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<mods:title id="A795BEBCE052E0FD2F891A364EF4C886">Centromedon zoe (Crustacea: Amphipoda: Lysianassoidea: Uristidae), a new deep-water scavenger species from the North Atlantic, with a key to the genus Centromedon</mods:title>
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<paragraph id="8BCD36F1FFA09D2062D9FD3F6C7723B7" blockId="1.[151,538,647,673]" box="[151,538,647,673]" pageId="1" pageNumber="55">
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<emphasis id="B906EAE3FFA09D2062D9FD3F6C7723B7" bold="true" box="[151,538,647,673]" pageId="1" pageNumber="55">
Genus
<taxonomicName id="4C724D72FFA09D2062A0FD3F6C7723B7" ID-CoL="7NS7J" authority="Sars, 1891" authorityName="Sars" authorityYear="1891" box="[238,538,647,673]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA09D2062A0FD3F6FFF23B7" bold="true" box="[238,402,647,673]" italics="true" pageId="1" pageNumber="55">Centromedon</emphasis>
<bibRefCitation id="EFE34B00FFA09D2063D4FD3F6C7723B7" author="Sars" box="[410,538,647,673]" pageId="1" pageNumber="55" refString="Sars G. O. (1891) An Account of the Crustacea of Norway, with short descriptions and figures of all the Species. Vol. I. Amphipoda. Parts 4 - 9. Alb. Cammermeyer, Christiana, pp. 69 - 212. Plates 25 - 72." type="book chapter" year="1891">Sars, 1891</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BCD36F1FFA09D2062D9FD776C552214" blockId="1.[151,1436,718,802]" pageId="1" pageNumber="55">
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<taxonomicName id="4C724D72FFA09D2062D9FD776FD023F2" ID-CoL="7NS7J" authority="Sars, 1891: 99" authorityName="Sars" authorityPageNumber="99" authorityYear="1891" box="[151,445,719,741]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA09D2062D9FD776F4D23F3" box="[151,288,719,741]" italics="true" pageId="1" pageNumber="55">Centromedon</emphasis>
<treatmentCitation id="0AD310E0FFA09D206369FD776FD023F2" author="Sars" box="[295,445,719,740]" page="99" pageId="1" pageNumber="55" year="1891">
<bibRefCitation id="EFE34B00FFA09D206369FD776FFA23F2" author="Sars" box="[295,407,719,740]" pageId="1" pageNumber="55" refString="Sars G. O. (1891) An Account of the Crustacea of Norway, with short descriptions and figures of all the Species. Vol. I. Amphipoda. Parts 4 - 9. Alb. Cammermeyer, Christiana, pp. 69 - 212. Plates 25 - 72." type="book chapter" year="1891">Sars, 1891</bibRefCitation>
: 99
</treatmentCitation>
</taxonomicName>
101, pl. 34, fig. 2. —
<treatmentCitation id="0AD310E0FFA09D2060EBFD766D0D23F2" author="Barnard" box="[677,864,718,740]" page="35" pageId="1" pageNumber="55" year="1962">
<bibRefCitation id="EFE34B00FFA09D2060EBFD766D5423F2" author="Barnard" box="[677,825,718,740]" pageId="1" pageNumber="55" refString="Barnard, J. L. (1962) South Atlantic abyssal amphipods collected by R. V. Vema. Abyssal Crustacea Research Series, 1, 1 - 79." type="journal article" year="1962">Barnard, 1962</bibRefCitation>
: 35
</treatmentCitation>
. —
<treatmentCitation id="0AD310E0FFA09D2061C8FD766A5A23F2" author="Olerod" box="[902,1079,718,740]" page="50" pageId="1" pageNumber="55" year="1980">
<bibRefCitation id="EFE34B00FFA09D2061C8FD766A7D23F2" author="Olerod" box="[902,1040,718,740]" pageId="1" pageNumber="55" refString="Olerod, R. (1980) A taxonomic study of the lysianassid genus Centromedon G. O. Sars (Crustacea, Amphipoda). Zoologica Scripta, 9, 35 - 52." type="journal article" year="1980">Olerod, 1980</bibRefCitation>
: 50
</treatmentCitation>
, 51. —
<treatmentCitation id="0AD310E0FFA09D2066CDFD766E9D2214" author="Barnard" page="474" pageId="1" pageNumber="55" year="1991">
<bibRefCitation id="EFE34B00FFA09D2066CDFD766BF523F2" author="Barnard" box="[1155,1432,718,740]" pageId="1" pageNumber="55" refString="Barnard J. L. &amp; Karaman, G. S. (1991) The families and genera of marine gammaridean Amphipoda (except marine gammaroids). Records of the Australian Museum, Supplement 13 (2), 419 - 866." type="journal article" year="1991">Barnard &amp; Karaman, 1991</bibRefCitation>
: 474
</treatmentCitation>
. —
<treatmentCitation id="0AD310E0FFA09D20635DFD566C582214" author="Lowry" box="[275,565,749,771]" page="13" pageId="1" pageNumber="55" year="1995">
<bibRefCitation id="EFE34B00FFA09D20635DFD566C622215" author="Lowry" box="[275,527,749,771]" pageId="1" pageNumber="55" refString="Lowry, J. K. &amp; Stoddart, H. E. (1995) New lysianassoid genera and species from south-eastern Australia (Crustacea: Amphipoda). Records of the Australian Museum, 47, 7 - 25." type="journal article" year="1995">Lowry &amp; Stoddart, 1995</bibRefCitation>
: 13
</treatmentCitation>
.
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</paragraph>
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<emphasis id="B906EAE3FFA09D2062D9FCB46F4F2234" bold="true" box="[151,290,779,802]" pageId="1" pageNumber="55">
<typeStatus id="54C98853FFA09D2062D9FCB46EA12237" box="[151,204,780,801]" pageId="1" pageNumber="55">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C724D72FFA09D206367FCB46C142237" authority="Lilljeborg, 1865" authorityName="Lilljeborg" authorityYear="1865" box="[297,633,779,802]" class="Malacostraca" family="Uristidae" genus="Anonyx" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B906EAE3FFA09D206367FCB46FA62234" box="[297,459,780,802]" italics="true" pageId="1" pageNumber="55">Anonyx pumilus</emphasis>
<bibRefCitation id="EFE34B00FFA09D20639CFCB36C142237" author="Lilljeborg" box="[466,633,779,801]" pageId="1" pageNumber="55" refString="Lilljeborg, W. (1865) On the Lysianassa magellanica H. Milne Edwards and on the Crustacea of the suborder Amphipoda and subfamily Lysianassina found an [sic] the coast of Sweden and Norway. Nova Acta Regiae Societatis Scientarum Upsaliensis, III Serie 6 (1), 1 - 38." type="journal article" year="1865">Lilljeborg, 1865</bibRefCitation>
</taxonomicName>
, original designation.
</paragraph>
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<subSubSection id="C368657AFFA09D2062D9FCF66FD62495" pageId="1" pageNumber="55" type="diagnosis">
<paragraph id="8BCD36F1FFA09D2062D9FCF66A9225E4" blockId="1.[151,1437,846,2023]" pageId="1" pageNumber="55">
<emphasis id="B906EAE3FFA09D2062D9FCF66F7C2271" bold="true" box="[151,273,846,871]" pageId="1" pageNumber="55">Diagnosis.</emphasis>
Lateral cephalic lobes triangular, subacute to acute. Eyes apparently absent. Callynophore well-developed in both males and females. Epistome flattened and separated from upper lip by a sinus or notch; upper lip rounded and projecting in front of epistome. Inner lobes of lower lip weak or absent. Mandibular molar conicolaminate with vestigial triturative area, palp attached level with molar. Maxilla 1, inner plate stout with two pappose setae on apex (one in
<taxonomicName id="4C724D72FFA09D2063C4FC676F8622E1" box="[394,491,991,1015]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="pavor">
<emphasis id="B906EAE3FFA09D2063C4FC676F8622E1" box="[394,491,991,1015]" italics="true" pageId="1" pageNumber="55">C. pavor</emphasis>
</taxonomicName>
); outer plate with 711 setal-teeth in 74 crown arrangement (reduced to 43 crown in
<taxonomicName id="4C724D72FFA09D2062FAFBBB6F7A250D" box="[180,279,1027,1051]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="pavor">
<emphasis id="B906EAE3FFA09D2062FAFBBB6F7A250D" box="[180,279,1027,1051]" italics="true" pageId="1" pageNumber="55">C. pavor</emphasis>
</taxonomicName>
and 63 crown in
<taxonomicName id="4C724D72FFA09D2063A4FBBB6C0E250D" box="[490,611,1027,1051]" class="Malacostraca" family="Uristidae" genus="Anonyx" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B906EAE3FFA09D2063A4FBBB6C0E250D" box="[490,611,1027,1051]" italics="true" pageId="1" pageNumber="55">C. pumilus</emphasis>
</taxonomicName>
); palp twoarticulate, reaching beyond outer plate. Maxilla 2, outer plate longer than inner plate. Maxilliped, outer plate not reaching article 3 of palp, suboval, with long robust setae apically, small and sparsely placed robust nodular setae on straight inner margin; article 4 of palp normal to vestigial. Gnathopod 1 variable from subchelate to simple; coxa 1 variable from strongly to weakly tapered, (not tapered in
<taxonomicName id="4C724D72FFA09D2062D9FB2B6F7425BD" box="[151,281,1171,1195]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="B906EAE3FFA09D2062D9FB2B6F7425BD" box="[151,281,1171,1195]" italics="true" pageId="1" pageNumber="55">C. typhlops</emphasis>
</taxonomicName>
); ischium short (1: &lt;1.13). Gnathopod 2 subchelate to chelate. Pereopod 7 with distal articles shortened. Epimeron 3 posteroventrally produced into an acute tooth. Urosomite 1 with dorsal carina. Outer ramus of uropod 3 two-articulate, article 2 long (&gt;50% article 1 length). Telson longer than wide, deeply cleft.
</paragraph>
<paragraph id="8BCD36F1FFA09D20628BFB466FD62495" blockId="1.[151,1437,846,2023]" pageId="1" pageNumber="55">
<emphasis id="B906EAE3FFA09D20628BFB466FD02401" bold="true" box="[197,445,1278,1303]" pageId="1" pageNumber="55">Species composition.</emphasis>
<taxonomicName id="4C724D72FFA09D206389FB466DFA2400" authority="Lilljeborg, 1865" authorityName="Lilljeborg" authorityYear="1865" box="[455,919,1278,1302]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B906EAE3FFA09D206389FB466CAC2400" box="[455,705,1278,1302]" italics="true" pageId="1" pageNumber="55">Centromedon pumilus</emphasis>
(
<bibRefCitation id="EFE34B00FFA09D20609FFB466DE22400" author="Lilljeborg" box="[721,911,1278,1302]" pageId="1" pageNumber="55" refString="Lilljeborg, W. (1865) On the Lysianassa magellanica H. Milne Edwards and on the Crustacea of the suborder Amphipoda and subfamily Lysianassina found an [sic] the coast of Sweden and Norway. Nova Acta Regiae Societatis Scientarum Upsaliensis, III Serie 6 (1), 1 - 38." type="journal article" year="1865">Lilljeborg, 1865</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C724D72FFA09D2061EBFB466B3A2400" authority="Goes, 1866" authorityName="Goes" authorityYear="1866" box="[933,1367,1278,1302]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="productus">
<emphasis id="B906EAE3FFA09D2061EBFB466AD82400" box="[933,1205,1278,1302]" italics="true" pageId="1" pageNumber="55">Centromedon productus</emphasis>
(
<bibRefCitation id="EFE34B00FFA09D206689FB466B222400" author="Goes" box="[1223,1359,1278,1302]" pageId="1" pageNumber="55" refString="Goes, A. (1866) Crustacea Amphipoda maris Spetsbergiam alluentis, cum specibus aliis arcticis enumerat. Ofversigt af Kongelige Vetenskaps-Akademiens Forhandligar, 1865, 517 - 536." type="journal article" year="1866">Goës, 1866</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C724D72FFA09D20672BFB476F90242C" authority="Sars, 1879" authorityName="Sars" authorityYear="1879" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="B906EAE3FFA09D20672BFB476F05242D" italics="true" pageId="1" pageNumber="55">Centromedon typhlops</emphasis>
(
<bibRefCitation id="EFE34B00FFA09D206336FA9A6F98242C" author="Sars" box="[376,501,1314,1338]" pageId="1" pageNumber="55" refString="Sars G. O. (1879) Crustacea et Pycnogonida nova in itinere 2 do &amp; 3 tio expeditionis norvegicae anno 1877 &amp; 78 collecta. (Prodromus descriptionis.). Archiv for Mathmatik og Naturvidenskab, 4, 427 - 476." type="journal article" year="1879">Sars, 1879</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C724D72FFA09D206047FA9B6DD9242C" authority="Sars, 1879" authorityName="Sars" authorityYear="1879" box="[521,948,1314,1339]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="B906EAE3FFA09D206047FA9B6D72242D" box="[521,799,1315,1339]" italics="true" pageId="1" pageNumber="55">Centromedon calcaratus</emphasis>
(
<bibRefCitation id="EFE34B00FFA09D206161FA9A6DC6242C" author="Sars" box="[815,939,1314,1338]" pageId="1" pageNumber="55" refString="Sars G. O. (1879) Crustacea et Pycnogonida nova in itinere 2 do &amp; 3 tio expeditionis norvegicae anno 1877 &amp; 78 collecta. (Prodromus descriptionis.). Archiv for Mathmatik og Naturvidenskab, 4, 427 - 476." type="journal article" year="1879">Sars, 1879</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C724D72FFA09D20618FFA9B6B31242C" authority="Bonnier, 1896" authorityName="Bonnier" authorityYear="1896" box="[961,1372,1314,1339]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B906EAE3FFA09D20618FFA9B6ACC242D" box="[961,1185,1315,1339]" italics="true" pageId="1" pageNumber="55">Centromedon laevis</emphasis>
(
<bibRefCitation id="EFE34B00FFA09D2066FFFA9A6B39242C" author="Bonnier" box="[1201,1364,1314,1338]" pageId="1" pageNumber="55" refString="Bonnier, J. (1896) Edriopthalmes. Resultats scientifiques de la campagne du &quot; Caudan &quot; dans le Golfe de Gascogne. Annals de l'Universite de Lyon, 26, 527 - 689." type="journal article" year="1896">Bonnier, 1896</bibRefCitation>
)
</taxonomicName>
(new combination);
<taxonomicName id="4C724D72FFA09D206375FAFE6C942448" authority="Barnard, 1962" authorityName="Barnard" authorityYear="1962" box="[315,761,1350,1374]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="mediator">
<emphasis id="B906EAE3FFA09D206375FAFE6C522448" box="[315,575,1350,1374]" italics="true" pageId="1" pageNumber="55">Centromedon mediator</emphasis>
(
<bibRefCitation id="EFE34B00FFA09D206000FAFE6C9C2448" author="Barnard" box="[590,753,1350,1374]" pageId="1" pageNumber="55" refString="Barnard, J. L. (1962) South Atlantic abyssal amphipods collected by R. V. Vema. Abyssal Crustacea Research Series, 1, 1 - 79." type="journal article" year="1962">Barnard, 1962</bibRefCitation>
)
</taxonomicName>
(new combination);
<taxonomicName id="4C724D72FFA09D2061AAFAFE6B0A2448" authority="Barnard 1966" authorityName="Barnard" authorityYear="1966" box="[996,1383,1350,1375]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="pavor">
<emphasis id="B906EAE3FFA09D2061AAFAFE6AA92449" box="[996,1220,1350,1375]" italics="true" pageId="1" pageNumber="55">Centromedon pavor</emphasis>
<bibRefCitation id="EFE34B00FFA09D206685FAFE6B0A2448" author="Barnard" box="[1227,1383,1350,1374]" pageId="1" pageNumber="55" refString="Barnard, J. L. (1966) Benthic Amphipoda of Monterey Bay, California. Proceedings of the United States National Museum, 119 (3541), 1 - 41." type="journal article" year="1966">Barnard 1966</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C724D72FFA09D2062D9FAD36F372495" box="[151,346,1387,1411]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="zoe" status="sp. nov.">
<emphasis id="B906EAE3FFA09D2062D9FAD36F372495" box="[151,346,1387,1411]" italics="true" pageId="1" pageNumber="55">Centromedon zoe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2355798FFA09D20632FFAD36FD62495" box="[353,443,1387,1411]" pageId="1" pageNumber="55" rank="species">
<emphasis id="B906EAE3FFA09D20632FFAD36FD62495" bold="true" box="[353,443,1387,1411]" pageId="1" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C368657AFFA09D20628BFA376FB8274C" pageId="1" pageNumber="55" type="discussion">
<paragraph id="8BCD36F1FFA09D20628BFA376FB8274C" blockId="1.[151,1437,846,2023]" pageId="1" pageNumber="55">
<emphasis id="B906EAE3FFA09D20628BFA376F5424B1" bold="true" box="[197,313,1423,1447]" pageId="1" pageNumber="55">Remarks.</emphasis>
<taxonomicName id="4C724D72FFA09D20630FFA376D6324B0" authority="Barnard, 1962" authorityName="Barnard" authorityYear="1962" box="[321,782,1422,1446]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="typhlops" subSpecies="mediator">
<emphasis id="B906EAE3FFA09D20630FFA376C0824B0" box="[321,613,1422,1446]" italics="true" pageId="1" pageNumber="55">Uristes typhlops mediator</emphasis>
<bibRefCitation id="EFE34B00FFA09D206023FA366D6324B0" author="Barnard" box="[621,782,1422,1446]" pageId="1" pageNumber="55" refString="Barnard, J. L. (1962) South Atlantic abyssal amphipods collected by R. V. Vema. Abyssal Crustacea Research Series, 1, 1 - 79." type="journal article" year="1962">Barnard, 1962</bibRefCitation>
</taxonomicName>
was originally described as a subspecies of
<taxonomicName id="4C724D72FFA09D20674AFA366FE024DC" authority="Sars, 1879" authorityName="Sars" authorityYear="1879" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="B906EAE3FFA09D20674AFA366E9B24DD" italics="true" pageId="1" pageNumber="55">Centromedon typhlops</emphasis>
(
<bibRefCitation id="EFE34B00FFA09D206349FA0A6FE824DC" author="Sars" box="[263,389,1458,1482]" pageId="1" pageNumber="55" refString="Sars G. O. (1879) Crustacea et Pycnogonida nova in itinere 2 do &amp; 3 tio expeditionis norvegicae anno 1877 &amp; 78 collecta. (Prodromus descriptionis.). Archiv for Mathmatik og Naturvidenskab, 4, 427 - 476." type="journal article" year="1879">Sars, 1879</bibRefCitation>
)
</taxonomicName>
. An examination of the
<typeStatus id="54C98853FFA09D2060E9FA0B6CBA24DD" box="[679,727,1459,1483]" pageId="1" pageNumber="55">type</typeStatus>
material of both
<taxonomicName id="4C724D72FFA09D2061ECFA0C6AA424DD" box="[930,1225,1459,1483]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="typhlops" subSpecies="mediator">
<emphasis id="B906EAE3FFA09D2061ECFA0C6AA424DD" box="[930,1225,1459,1483]" italics="true" pageId="1" pageNumber="55">Uristes typhlops mediator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C724D72FFA09D20674AFA0B6E9B24F8" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="B906EAE3FFA09D20674AFA0B6E9B24F8" italics="true" pageId="1" pageNumber="55">Centromedon typhlops</emphasis>
</taxonomicName>
shows that they differ by characters of the gnathopod 2, epimeron 3 and telson. There is also a clear size difference between the two species, with mature females of
<taxonomicName id="4C724D72FFA09D20617DFA436DDF2705" box="[819,946,1531,1555]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis id="B906EAE3FFA09D20617DFA436DDF2705" box="[819,946,1531,1555]" italics="true" pageId="1" pageNumber="55">C. typhlops</emphasis>
</taxonomicName>
reaching
<quantity id="4C8A9B14FFA09D20666EFA426A032704" box="[1056,1134,1530,1554]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="55" unit="mm" value="17.0">17 mm</quantity>
in length while those of
<taxonomicName id="4C724D72FFA09D2067CCFA436E912720" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="mediator">
<emphasis id="B906EAE3FFA09D2067CCFA436E912720" italics="true" pageId="1" pageNumber="55">C. mediator</emphasis>
</taxonomicName>
are
<quantity id="4C8A9B14FFA09D206362F9A66F1D2720" box="[300,368,1566,1590]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="55" unit="mm" value="6.0">6 mm</quantity>
.
<taxonomicName id="4C724D72FFA09D206334F9986CF12720" box="[378,668,1566,1591]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="typhlops" subSpecies="mediator">
<emphasis id="B906EAE3FFA09D206334F9986CF12720" box="[378,668,1566,1591]" italics="true" pageId="1" pageNumber="55">Uristes typhlops mediator</emphasis>
</taxonomicName>
conforms to the diagnosis of
<taxonomicName id="4C724D72FFA09D2061A7F9A66AED2720" box="[1001,1152,1566,1590]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA09D2061A7F9A66AED2720" box="[1001,1152,1566,1590]" italics="true" pageId="1" pageNumber="55">Centromedon</emphasis>
</taxonomicName>
and is hereby transferred to this genus as
<taxonomicName id="4C724D72FFA09D206304F9FB6FA3274D" box="[330,462,1603,1627]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="mediator">
<emphasis id="B906EAE3FFA09D206304F9FB6FA3274D" box="[330,462,1603,1627]" italics="true" pageId="1" pageNumber="55">C. mediator</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C368657AFFA09D23628BF9DE6BF1222E" lastPageId="2" lastPageNumber="56" pageId="1" pageNumber="55" type="reference_group">
<paragraph id="8BCD36F1FFA09D20628BF9DE6F6D2618" blockId="1.[151,1437,846,2023]" pageId="1" pageNumber="55">
<taxonomicName id="4C724D72FFA09D20628BF9DE6C3E2768" authority="Bonnier, 1896" authorityName="Bonnier" authorityYear="1896" box="[197,595,1638,1662]" class="Malacostraca" family="Lysianassidae" genus="Orchomenella" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B906EAE3FFA09D20628BF9DE6FC12768" box="[197,428,1638,1662]" italics="true" pageId="1" pageNumber="55">Orchomenella laevis</emphasis>
<bibRefCitation id="EFE34B00FFA09D2063FAF9DE6C3E2768" author="Bonnier" box="[436,595,1638,1662]" pageId="1" pageNumber="55" refString="Bonnier, J. (1896) Edriopthalmes. Resultats scientifiques de la campagne du &quot; Caudan &quot; dans le Golfe de Gascogne. Annals de l'Universite de Lyon, 26, 527 - 689." type="journal article" year="1896">Bonnier, 1896</bibRefCitation>
</taxonomicName>
was described from a single adult male (
<quantity id="4C8A9B14FFA09D20666EF9DE6A0E2768" box="[1056,1123,1638,1662]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="55" unit="mm" value="5.0">5 mm</quantity>
) taken in the Bay of Biscay (
<geoCoordinate id="EE465036FFA09D2062D1F9326F6B27B4" box="[159,262,1674,1698]" direction="north" orientation="latitude" pageId="1" pageNumber="55" precision="925" value="44.283333">44°17' N</geoCoordinate>
,
<geoCoordinate id="EE465036FFA09D20635CF9326F1027B4" box="[274,381,1674,1698]" direction="west" orientation="longitude" pageId="1" pageNumber="55" precision="925" value="-2.3">02°18' W</geoCoordinate>
) at
<specimenCount id="9D74FD78FFA09D2063E9F9326F9B27B4" box="[423,502,1674,1698]" pageId="1" pageNumber="55" type="generic">
<quantity id="4C8A9B14FFA09D2063E9F9326F9C27B4" box="[423,497,1674,1698]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="55" unit="m" value="950.0">950 m</quantity>
.
</specimenCount>
It was transferred to
<taxonomicName id="4C724D72FFA09D2060ABF9336D0527B5" box="[741,872,1675,1699]" class="Malacostraca" family="Lysianassidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA09D2060ABF9336D0527B5" box="[741,872,1675,1699]" italics="true" pageId="1" pageNumber="55">Tryphosella</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE34B00FFA09D2061DAF9326A5927B4" author="Barnard" box="[916,1076,1674,1698]" pageId="1" pageNumber="55" refString="Barnard, J. L. (1964) Marine Amphipoda of Bahia de San Quintin, Baja California. Pacific Naturalist, 4, 55 - 139." type="journal article" year="1964">Barnard, 1964</bibRefCitation>
but it has a nondominant epistome, reduced distal articles of pereopod 7 and otherwise conforms to the new diagnosis of
<emphasis id="B906EAE3FFA09D2066E2F9166B2727D0" box="[1196,1354,1710,1734]" italics="true" pageId="1" pageNumber="55">
<taxonomicName id="4C724D72FFA09D2066E2F9166B2B27D0" ID-CoL="7NS7J" box="[1196,1350,1710,1734]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">Centromedon</taxonomicName>
,
</emphasis>
except for the outer plate of the maxilliped, which does reach article 3 of the palp. It is hereby transferred to this genus as
<emphasis id="B906EAE3FFA09D2062D9F94F6F6D2618" box="[151,256,1782,1806]" italics="true" pageId="1" pageNumber="55">
<taxonomicName id="4C724D72FFA09D2062D9F94F6E912618" box="[151,252,1782,1806]" class="Malacostraca" family="Lysianassidae" genus="Orchomenella" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="laevis">C. laevis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BCD36F1FFA09D23628BF8A26E942072" blockId="1.[151,1437,846,2023]" lastBlockId="2.[151,1436,151,824]" lastPageId="2" lastPageNumber="56" pageId="1" pageNumber="55">
The distinctions between the genus
<taxonomicName id="4C724D72FFA09D20602EF8A46CDC2625" ID-CoL="7Q5VZ" box="[608,689,1820,1843]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA09D20602EF8A46CDC2625" box="[608,689,1820,1843]" italics="true" pageId="1" pageNumber="55">Uristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C724D72FFA09D2060A2F8A36DEE2625" ID-CoL="7NS7J" box="[748,899,1819,1843]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA09D2060A2F8A36DEE2625" box="[748,899,1819,1843]" italics="true" pageId="1" pageNumber="55">Centromedon</emphasis>
</taxonomicName>
are unclear but clarification awaits a thorough redescription of
<taxonomicName id="4C724D72FFA09D206300F8F86C002640" ID-CoL="7DRTY" authority="Dana, 1849" authorityName="Dana" authorityYear="1849" box="[334,621,1854,1879]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B906EAE3FFA09D206300F8F86F8E2641" box="[334,483,1856,1879]" italics="true" pageId="1" pageNumber="55">Uristes gigas</emphasis>
<bibRefCitation id="EFE34B00FFA09D2063A7F8876C002640" author="Dana" box="[489,621,1854,1878]" pageId="1" pageNumber="55" refString="Dana, J. D. (1849) Synopsis of the genera of Gammaracea. American Journal of Sciences and the Arts, Series 2, 8, 135 - 140." type="journal article" year="1849">Dana, 1849</bibRefCitation>
</taxonomicName>
, the
<typeStatus id="54C98853FFA09D2060EFF8876CBC2641" box="[673,721,1855,1879]" pageId="1" pageNumber="55">type</typeStatus>
species of
<taxonomicName id="4C724D72FFA09D206103F8F86DF32641" ID-CoL="7Q5VZ" box="[845,926,1856,1879]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA09D206103F8F86DF32641" box="[845,926,1856,1879]" italics="true" pageId="1" pageNumber="55">Uristes</emphasis>
</taxonomicName>
and a re-examination of the species currently included in that genus, an analysis which is beyond the scope of this paper. The location of Danas material is unknown. It is very likely that the
<typeStatus id="54C98853FFA09D206058F83F6C2B2689" box="[534,582,1927,1951]" pageId="1" pageNumber="55">type</typeStatus>
material was part of a collection on loan from the Smithsonian Institution to William Stimpson, at the Chicago Academy of Science, at the time of the Chicago fire in 1871. All of Stimpsons material was lost in the fire and it is known that he had on loan 10,000 jars of Crustacea, many of which contained
<typeStatus id="54C98853FFA39D2362D9FF206EAA21A6" box="[151,199,152,176]" pageId="2" pageNumber="56">type</typeStatus>
material described by James Dana (
<bibRefCitation id="EFE34B00FFA39D236026FF2F6C9E21B9" author="Evans" box="[616,755,151,175]" pageId="2" pageNumber="56" refString="Evans, A. C. (1967) Syntypes of Decapoda described by William Stimpson &amp; James Dana in the collections of the British Museum (Natural History). Journal of Natural History, 1, 399 - 411." type="journal article" year="1967">Evans 1967</bibRefCitation>
).
<typeStatus id="54C98853FFA39D236147FF206D2D21A6" box="[777,832,152,176]" pageId="2" pageNumber="56">Type</typeStatus>
material is not held at the Natural History Museum, London (
<bibRefCitation id="EFE34B00FFA39D2362B0FF046C6021C2" author="Thurston" box="[254,525,188,212]" pageId="2" pageNumber="56" refString="Thurston, M. H. &amp; Allen, E. A. (1969) Type material of the families Lysianassidae, Stegocephalidae, Ampeliscidae and Haustoriidae (Crustacea: Amphipoda) in the collections of the British Museum (Natural History). Bulletin of the British Museum (Natural History) Zoology, 17, 347 - 388." type="journal article" year="1969">Thurston &amp; Allen 1969</bibRefCitation>
) and is presumed to be no longer extant. It is generally accepted, however, that
<taxonomicName id="4C724D72FFA39D2362D9FF586C3C21E1" authority="Stebbing, 1888" authorityName="Stebbing" authorityYear="1888" box="[151,593,223,248]" class="Malacostraca" family="Lysianassidae" genus="Tryphosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="antennipotens">
<emphasis id="B906EAE3FFA39D2362D9FF586FCE21EE" box="[151,419,224,248]" italics="true" pageId="2" pageNumber="56">Tryphosa antennipotens</emphasis>
<bibRefCitation id="EFE34B00FFA39D2363E6FF676C3C21E1" author="Stebbing" box="[424,593,223,247]" pageId="2" pageNumber="56" refString="Stebbing, T. R. R. (1888) Report on the Amphipoda collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76, Zoology, 29, Eyre &amp; Spottiswoodie, London. 1737 pp, 210 plates." type="book" year="1888">Stebbing, 1888</bibRefCitation>
</taxonomicName>
is synonymous with
<taxonomicName id="4C724D72FFA39D236172FF596DBC21EE" ID-CoL="7DRTY" box="[828,977,225,248]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B906EAE3FFA39D236172FF596DBC21EE" box="[828,977,225,248]" italics="true" pageId="2" pageNumber="56">Uristes gigas</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE34B00FFA39D236644FF676AC221E1" author="Stebbing" box="[1034,1199,223,247]" pageId="2" pageNumber="56" refString="Stebbing, T. R. R. (1899) Revision of Amphipoda (continued). Annals and Magazine of Natural History, Series 7, 4, 205 - 211." type="journal article" year="1899">Stebbing 1899</bibRefCitation>
). A full redescription of this species and designation of a
<typeStatus id="54C98853FFA39D236061FEBD6CEA200B" box="[559,647,261,285]" pageId="2" pageNumber="56" type="neotype">neotype</typeStatus>
is needed to clarify the status of the genus. It is probable that a large number of the species currently included in the genus (22+) will be transferred to other genera following a critical analysis.
</paragraph>
<paragraph id="8BCD36F1FFA39D23628BFEC86A6120D9" blockId="2.[151,1436,151,824]" pageId="2" pageNumber="56">
<taxonomicName id="4C724D72FFA39D23628BFEC86F31209E" ID-CoL="7NS7J" box="[197,348,368,392]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA39D23628BFEC86F31209E" box="[197,348,368,392]" italics="true" pageId="2" pageNumber="56">Centromedon</emphasis>
</taxonomicName>
should be retained for those species characterised by: sub-acute to acute head lobes; tapered coxa 1; laminar, unridged molar with vestigial triturative area; maxilla 1 outer plate with 7-4 crown setal-tooth arrangement; shortened distal articles of pereopod 7 and acutely produced epimeron 3.
</paragraph>
<paragraph id="8BCD36F1FFA39D23628BFE646BF1222E" blockId="2.[151,1436,151,824]" pageId="2" pageNumber="56">
The similarity of
<taxonomicName id="4C724D72FFA39D2363DBFE646C4020E2" ID-CoL="7NS7J" box="[405,557,476,500]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA39D2363DBFE646C4020E2" box="[405,557,476,500]" italics="true" pageId="2" pageNumber="56">Centromedon</emphasis>
</taxonomicName>
to
<taxonomicName id="4C724D72FFA39D236017FE646A7E20E2" ID-CoL="7PGF8" authority="Barnard &amp; Karaman, 1987" authorityName="Barnard &amp; Karaman" authorityYear="1987" box="[601,1043,476,500]" class="Malacostraca" family="Uristidae" genus="Galathella" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA39D236017FE646CBC20E2" box="[601,721,476,500]" italics="true" pageId="2" pageNumber="56">Galathella</emphasis>
<bibRefCitation id="EFE34B00FFA39D236092FE646A7E20E2" author="Barnard" box="[732,1043,476,500]" pageId="2" pageNumber="56" refString="Barnard, J. L. &amp; Karaman, G. S. (1987) Revisions in classification of gammaridean Amphipoda (Crustacea), Part 3. Proceedings of the Biological Society of Washington, 100, 856 - 875." type="journal article" year="1987">Barnard &amp; Karaman, 1987</bibRefCitation>
</taxonomicName>
was noted by
<bibRefCitation id="EFE34B00FFA39D236688FE656E8E2301" author="Lowry" pageId="2" pageNumber="56" refString="Lowry, J. K. &amp; Stoddart, H. E. (1995) New lysianassoid genera and species from south-eastern Australia (Crustacea: Amphipoda). Records of the Australian Museum, 47, 7 - 25." type="journal article" year="1995">Lowry &amp; Stoddart (1995)</bibRefCitation>
, who pointed out that these are the only two genera in the
<taxonomicName id="4C724D72FFA39D236121FE476DBB2301" ID-CoL="7NLH7" box="[879,982,511,535]" class="Malacostraca" family="Uristidae" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="family">Uristidae</taxonomicName>
with a tapering first coxa and 7-4 crown setal-tooth arrangement. It was also noted that both genera show a range of gnathopod 1 morphologies with species characterised by a simple, weak structure through to others with a strongly subchelate gnathopod 1. The only differences between the two genera appear to be the sub-acute lateral cephalic lobe (rounded in
<taxonomicName id="4C724D72FFA39D2366FDFDD46B472392" ID-CoL="7PGF8" box="[1203,1322,620,644]" class="Malacostraca" family="Uristidae" genus="Galathella" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA39D2366FDFDD46B472392" box="[1203,1322,620,644]" italics="true" pageId="2" pageNumber="56">Galathella</emphasis>
</taxonomicName>
), the lack of proximal A3 setae on the mandibular palp, the less well-developed molar, and the acutely produced posteroventral corner of epimeron
<quantity id="4C8A9B14FFA39D2363EEFD0C6FA723DA" box="[416,458,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="56" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C724D72FFA39D23639FFD0C6C0523DA" ID-CoL="7NS7J" box="[465,616,692,716]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA39D23639FFD0C6C0523DA" box="[465,616,692,716]" italics="true" pageId="2" pageNumber="56">Centromedon</emphasis>
</taxonomicName>
. Contrary to
<bibRefCitation id="EFE34B00FFA39D2360B2FD0D6A7123DA" author="Lowry" box="[764,1052,692,717]" pageId="2" pageNumber="56" refString="Lowry, J. K. &amp; Stoddart, H. E. (1995) New lysianassoid genera and species from south-eastern Australia (Crustacea: Amphipoda). Records of the Australian Museum, 47, 7 - 25." type="journal article" year="1995">Lowry &amp; Stoddart (1995)</bibRefCitation>
, most species of
<taxonomicName id="4C724D72FFA39D236695FD0C6B1F23DA" ID-CoL="7NS7J" box="[1243,1394,692,716]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAE3FFA39D236695FD0C6B1F23DA" box="[1243,1394,692,716]" italics="true" pageId="2" pageNumber="56">Centromedon</emphasis>
</taxonomicName>
(
<emphasis id="B906EAE3FFA39D2367CCFD0D6CC623E6" italics="true" pageId="2" pageNumber="56">
<taxonomicName id="4C724D72FFA39D2367CCFD0D6F6423E6" ID-CoL="5XHXQ" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="productus">C. productus</taxonomicName>
,
<taxonomicName id="4C724D72FFA39D23635BFD606FFA23E6" ID-CoL="7DS6B" box="[277,407,728,752]" class="Malacostraca" family="Uristidae" genus="Uristes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="typhlops">C. typhlops</taxonomicName>
,
<taxonomicName id="4C724D72FFA39D2363EAFD606F8723E6" ID-CoL="69HY2" box="[420,490,728,752]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="zoe">C. zoe</taxonomicName>
and
<taxonomicName id="4C724D72FFA39D23606AFD606CC623E6" ID-CoL="69HXR" box="[548,683,728,752]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="mediator">C. mediator</taxonomicName>
</emphasis>
) have callynophores in the female. There is a degree of variability within the genus in that
<taxonomicName id="4C724D72FFA39D2363EAFD456C732202" ID-CoL="5XHY2" box="[420,542,764,788]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B906EAE3FFA39D2363EAFD456C732202" box="[420,542,764,788]" italics="true" pageId="2" pageNumber="56">C. pumilus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C724D72FFA39D23601AFD456CDB2202" ID-CoL="69HY3" box="[596,694,765,788]" class="Malacostraca" family="Uristidae" genus="Centromedon" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="pavor">
<emphasis id="B906EAE3FFA39D23601AFD456CDB2202" box="[596,694,765,788]" italics="true" pageId="2" pageNumber="56">C. pavor</emphasis>
</taxonomicName>
have a reduced number of setal-teeth on the outer plate of maxilla 1, a reduced article 4 of the maxilliped palp and less well-developed callynophore and gnathopod 1 (
<bibRefCitation id="EFE34B00FFA39D2366B9FC986BFD222E" author="Olerod" box="[1271,1424,800,824]" pageId="2" pageNumber="56" refString="Olerod, R. (1980) A taxonomic study of the lysianassid genus Centromedon G. O. Sars (Crustacea, Amphipoda). Zoologica Scripta, 9, 35 - 52." type="journal article" year="1980">Olerod, 1980</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>