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<document id="21E07E5DB5EE8E280C1A1E850D171C0A" ID-CLB-Dataset="122177" ID-DOI="10.1093/zoolinnean/zlab091" ID-GBIF-Dataset="de4325f4-7b0a-41a6-8a42-22b9e766753a" ID-ISSN="0024-4082" ID-Zenodo-Dep="6993662" ID-ZooBank="73D99D73-1735-4F27-844E-796874E4A040" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660323608787" checkinUser="plazi" docAuthor="Bocakova, Milada, Campello-Gonçalves, Lucas &amp; Silveira, Luiz Felipe Lima Da" docDate="2022" docId="03DB87C0FFE08844F6306FEA13D97A9B" docLanguage="en" docName="zlab091.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Ledocas Olivier 1885" docType="treatment" docVersion="6" lastPageNumber="1189" masterDocId="FFE2FFB8FFE8884CF70B6A7410437C36" masterDocTitle="Phylogeny of the new subfamily Cladodinae: neotenic fireflies from the Neotropics (Coleoptera: Lampyridae)" masterLastPageNumber="1199" masterPageNumber="1181" pageNumber="1189" updateTime="1699366447678" updateUser="ExternalLinkService">
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<mods:title id="CDB4DCFA49917C4554C41BB8D61A6127">Phylogeny of the new subfamily Cladodinae: neotenic fireflies from the Neotropics (Coleoptera: Lampyridae)</mods:title>
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<mods:namePart id="FC20F0B4BC0DDB4E81910C76A6E27727">Bocakova, Milada</mods:namePart>
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<treatment id="03DB87C0FFE08844F6306FEA13D97A9B" ID-DOI="http://doi.org/10.5281/zenodo.6997824" ID-GBIF-Taxon="199422840" ID-Zenodo-Dep="6997824" LSID="urn:lsid:plazi:treatment:03DB87C0FFE08844F6306FEA13D97A9B" httpUri="http://treatment.plazi.org/id/03DB87C0FFE08844F6306FEA13D97A9B" lastPageNumber="1189" pageId="8" pageNumber="1189">
<subSubSection id="C368655DFFE08844F6306FEA120D7981" box="[315,590,1438,1463]" pageId="8" pageNumber="1189" type="nomenclature">
<paragraph id="8BCD36D6FFE08844F6306FEA120D7981" blockId="8.[315,590,1438,1463]" box="[315,590,1438,1463]" pageId="8" pageNumber="1189">
<heading id="D08581BAFFE08844F6306FEA120D7981" box="[315,590,1438,1463]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="1189" reason="2">
<taxonomicName id="4C724D55FFE08844F6306FEA120D7981" ID-CoL="5BRN" authority="OLIVIER, 1885" authorityName="Olivier" authorityYear="1885" box="[315,590,1438,1463]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAC4FFE08844F6306FEA11E67983" box="[315,421,1438,1461]" italics="true" pageId="8" pageNumber="1189">LEDOCAS</emphasis>
OLIVIER, 1885
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C368655DFFE08844F7AD6FB0126779CA" pageId="8" pageNumber="1189" type="description">
<paragraph id="8BCD36D6FFE08844F7AD6FB0126779CA" blockId="8.[166,739,1476,1532]" pageId="8" pageNumber="1189">
(
<figureCitation id="13492A53FFE08844F7A46FB0117779EB" box="[175,308,1476,1501]" captionStart="Figure 2" captionStartId="6.[145,225,1705,1727]" captionTargetBox="[227,1341,198,1663]" captionTargetId="figure-24@6.[225,1345,195,1665]" captionTargetPageId="6" captionText="Figure 2. Representatives of the Cladodinae; A, Cladodes flabellatus (photo: A. Reyes); B, Cladodes cincticollis (photo: V. Valdés); C, Cladodes ater (photo: M. Cid); D, Ledocas remixtus (photo: S. Quipildor-Sánchez); E, Nyctocrepis demoulini (photo: A. Madruga); F, Nyctocrepis stellata (photo:A.Katz); G, Dodacles elegans (photo: P. Smith); H, Brasilocladodes illigeri (photo: L. Silveira); I, Brasilocladodes delalandei (photo: A. Bouzan); J, Ledocas proteus (photo: M. Rojas-Quintero)." figureDoi="http://doi.org/10.5281/zenodo.6993668" httpUri="https://zenodo.org/record/6993668/files/figure.png" pageId="8" pageNumber="1189">FIGS 2D, 2J</figureCitation>
,
<figureCitation id="13492A53FFE08844F64A6FB0113C79EB" box="[321,383,1476,1501]" captionStart="Figure 3" captionStartId="7.[163,244,1665,1687]" captionTargetBox="[294,1311,215,1622]" captionTargetId="figure-24@7.[291,1315,195,1625]" captionTargetPageId="7" captionText="Figure 3. Habitus, dorsal view; A, Cladodes flabellatus; B, Ledocas parallelus; C, Ledocas pikillactanus sp. nov.; D, Nyctocrepis malleri; E, Andecladodes ovalis; F, Andecladodes cosangensis, lamella of antennomere III damaged; G, Brasilocladodes carinatus; H, Dryptelytra cayennensis; I, Dodacles elegans; J, Dodacles niger; K, Dodacles lateralis; L, Dryptelytra pampahermosae sp. nov.. Scales: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6993670" httpUri="https://zenodo.org/record/6993670/files/figure.png" pageId="8" pageNumber="1189">3B, C</figureCitation>
,
<figureCitation id="13492A53FFE08844F6876FB0118979EB" box="[396,458,1476,1501]" captionStart="Figure4" captionStartId="8.[145,221,1180,1202]" captionTargetBox="[150,1424,217,1139]" captionTargetId="figure-336@8.[145,1428,195,1140]" captionTargetPageId="8" captionText="Figure4. Pronotum:A,Cladodescincticollis;B,Ledocasparallelus;C,Ledocaspikillactanus;D,Nyctocrepismalleri;E,Andecladodes ovalis;F,Andecladodes cosangensis, lamellaof antennomere IIIdamaged;G, Brasilocladodescarinatus; H,Dryptelytra cayennensis; I, Dodacles elegans; J, Dodacles niger; K, Dodacles lateralis; L, Dryptelytra pampahermosae sp. nov.. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6993673" httpUri="https://zenodo.org/record/6993673/files/figure.png" pageId="8" pageNumber="1189">4B, C</figureCitation>
,
<figureCitation id="13492A53FFE08844F6DC6FB0125679EB" box="[471,533,1476,1501]" captionStart="Figure 5" captionStartId="9.[164,242,1236,1258]" captionTargetBox="[166,1441,196,1153]" captionTargetId="figure-307@9.[163,1443,195,1196]" captionTargetPageId="9" captionText="Figure 5. Antenna: A, Cladodes melanurus; B, Ledocas parallelus; C, Ledocas pikillactanus sp. nov.; D, Nyctocrepis malleri; E, Andecladodes ovalis; F, Andecladodes cosangensis; G, Dodacles elegans; H, Dryptelytra cayennensis; I, Dryptelytra pampahermosae sp. nov.; J, Vesta chevrolatii. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6993676" httpUri="https://zenodo.org/record/6993676/files/figure.png" pageId="8" pageNumber="1189">5B, C</figureCitation>
,
<figureCitation id="13492A53FFE08844F5296FB0120179EB" box="[546,578,1476,1501]" captionStart="Figure 6" captionStartId="10.[145,227,607,629]" captionTargetBox="[150,1420,204,563]" captionTargetId="figure-306@10.[145,1425,195,568]" captionTargetPageId="10" captionText="Figure 6. Head. A, ventral view; Ledocas pikillactanus sp. nov.; B, C, dorsal view: B, Andecladodes cosangensis; C, Andecladodes ovalis. Scales: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6993678" httpUri="https://zenodo.org/record/6993678/files/figure.png" pageId="8" pageNumber="1189">6A</figureCitation>
,
<figureCitation id="13492A53FFE08844F55B6FB012CD79EB" box="[592,654,1476,1501]" captionStart="Figure 7" captionStartId="10.[145,226,1157,1179]" captionTargetBox="[154,1418,750,1111]" captionTargetId="figure-332@10.[145,1425,740,1117]" captionTargetPageId="10" captionText="Figure 7. A, CH, Abdomen, ventral view: A, Ledocas pikillactanus sp. nov.; C, Andecladodes cosangensis sp.nov.; D, Andecladodes ovalis; E, Nyctocrepis malleri; F, Dodacles niger; G, Dodacles lateralis; H, Dryptelytra pampahermosae sp.nov.; B, distal portion of hind tibia and tarsus, Ledocas pikillactanus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6993680" httpUri="https://zenodo.org/record/6993680/files/figure.png" pageId="8" pageNumber="1189">7A, B</figureCitation>
,
<figureCitation id="13492A53FFE08844F5976FB0129E79EB" box="[668,733,1476,1501]" captionStart="Figure 8" captionStartId="11.[163,241,1019,1041]" captionTargetBox="[165,1443,195,979]" captionTargetId="figure-317@11.[163,1443,195,979]" captionText="Figure 8. A, C, E, G, I, K, M, O, Q, S, U, W, Sternum VIII; B, D, F, H, J, L, N, P, R, T, V, X, Pygidium; A, B, Ledocas parallelus; C, D, Ledocas pikillactanus sp. nov.; E, F, Andecladodes cosangensis; G, H, Andecladodes ovalis; I, J, Dryptelytra cayennensis; K, L, Dryptelytra pampahermosae sp. nov.; M, N, Dodacles elegans; O, P, Dodacles niger; Q, R, Dodacles lateralis; S, T, Nyctocrepis malleri; U, V, Brasilocladodes illigeri; W, X, Brasilocladodes carinatus. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6993682" httpUri="https://zenodo.org/record/6993682/files/figure.png" pageId="8" pageNumber="1189">8AD</figureCitation>
, 9A, B, 10B1C2)
</paragraph>
</subSubSection>
<subSubSection id="C368655DFFE08844F79A6C7F11837A17" box="[145,448,1547,1569]" pageId="8" pageNumber="1189" type="reference_group">
<paragraph id="8BCD36D6FFE08844F79A6C7F11837A17" blockId="8.[145,448,1547,1569]" box="[145,448,1547,1569]" pageId="8" pageNumber="1189">
<treatmentCitationGroup id="AB6211F8FFE08844F79A6C7F11837A17" box="[145,448,1547,1569]" pageId="8" pageNumber="1189">
<taxonomicName id="4C724D55FFE08844F79A6C7F11F87A17" ID-CoL="5BRN" authority="Olivier, 1885: 136" authorityName="Olivier" authorityPageNumber="136" authorityYear="1885" box="[145,443,1547,1569]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAC4FFE08844F79A6C7F10AF7A16" box="[145,236,1547,1568]" italics="true" pageId="8" pageNumber="1189">Ledocas</emphasis>
<treatmentCitation id="0AD310C7FFE08844F7F96C7F11F87A17" author="Olivier" box="[242,443,1547,1569]" page="136" pageId="8" pageNumber="1189" year="1885">Olivier, 1885: 136</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C368655DFFE08844F79A6C3F11537A48" pageId="8" pageNumber="1189" type="type_taxon">
<paragraph id="8BCD36D6FFE08844F79A6C3F11537A48" blockId="8.[145,760,1610,1662]" pageId="8" pageNumber="1189">
<emphasis id="B906EAC4FFE08844F79A6C3F116C7A56" box="[145,303,1611,1632]" italics="true" pageId="8" pageNumber="1189">
<typeStatus id="54C98874FFE08844F79A6C3F108A7A56" box="[145,201,1611,1632]" pageId="8" pageNumber="1189">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C724D55FFE08844F6326C3E128A7A56" ID-CoL="3SWR7" authority="Olivier, 1885" authorityName="Olivier" authorityYear="1885" box="[313,713,1610,1632]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B906EAC4FFE08844F6326C3E12637A69" box="[313,544,1610,1631]" italics="true" pageId="8" pageNumber="1189">Ledocas parallelus</emphasis>
Olivier, 1885
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C368655DFFE08844F79A6CD114E979FE" pageId="8" pageNumber="1189" type="diagnosis">
<paragraph id="8BCD36D6FFE08844F79A6CD114E979FE" blockId="8.[145,761,1701,1907]" lastBlockId="8.[809,1425,1305,1480]" pageId="8" pageNumber="1189">
<emphasis id="B906EAC4FFE08844F79A6CD1114F7A8C" box="[145,268,1701,1722]" italics="true" pageId="8" pageNumber="1189">Diagnosis:</emphasis>
Pronotum and elytra subparallel-sided. Eyes small to medium-sized, mandibles semicircular, never right-angled, antennae uniflabellate, fan-like, branches of median antennomeres about 4× longer than the antennomere body, flabella of antennomere III markedly shorter than the following branches, translucent spots on pronotum absent. Abdomen strongly laterally lobed, sternum VIII bisinuate, mucronate, pygidium usually as long as wide, widest medially, with median extension about half as long as the lateral corners, lateral margins rounded. Male genitalia small, parameres as long as phallus, parameral apices simply straight.
</paragraph>
</subSubSection>
<subSubSection id="C368655DFFE08844F4226F9215D17A61" pageId="8" pageNumber="1189" type="discussion">
<paragraph id="8BCD36D6FFE08844F4226F9215D17A61" blockId="8.[809,1426,1510,1624]" pageId="8" pageNumber="1189">
<emphasis id="B906EAC4FFE08844F4226F9213BE79CD" box="[809,1021,1510,1531]" italics="true" pageId="8" pageNumber="1189">Species included:</emphasis>
<taxonomicName id="4C724D55FFE08844F31E6F9215C879CD" authority="Olivier, 1908" authorityName="Olivier" authorityYear="1908" box="[1045,1419,1510,1532]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="species" species="detectus">
<emphasis id="B906EAC4FFE08844F31E6F9214A179CD" box="[1045,1250,1510,1531]" italics="true" pageId="8" pageNumber="1189">Ledocas detectus</emphasis>
Olivier, 1908
</taxonomicName>
,
<taxonomicName id="4C724D55FFE08844F4226C71140F7A2C" authority="(Olivier, 1908)" authorityName="Bocakova &amp; Campello-Gonçalves &amp; Silveira" authorityYear="2022" baseAuthorityName="Olivier" baseAuthorityYear="1908" box="[809,1100,1541,1563]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="species" species="emissus" status="comb. nov.">
<emphasis id="B906EAC4FFE08844F4226C7113EB7A2C" bold="true" box="[809,936,1541,1562]" italics="true" pageId="8" pageNumber="1189">L. emissus</emphasis>
(Olivier, 1908)
</taxonomicName>
<emphasis id="B906EAC4FFE08844F35A6C7114977A2C" bold="true" box="[1105,1236,1541,1562]" pageId="8" pageNumber="1189">
<taxonomicNameLabel id="A23557BFFFE08844F35A6C7114977A2C" box="[1105,1236,1541,1562]" pageId="8" pageNumber="1189" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C724D55FFE08844F3EB6C7115D27A2C" authority="Bocakova &amp; Campello-Gonçalves &amp; Silveira, 2022" authorityName="Bocakova &amp; Campello-Gonçalves &amp; Silveira" authorityYear="2022" box="[1248,1425,1541,1562]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="species" species="pikillactanus" status="sp. nov.">
<emphasis id="B906EAC4FFE08844F3EB6C7115D27A2C" box="[1248,1425,1541,1562]" italics="true" pageId="8" pageNumber="1189">L. pikillactanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557BFFFE08844F4226C5013C47A0F" box="[809,903,1572,1593]" pageId="8" pageNumber="1189" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="B906EAC4FFE08844F49D6C5013F67A0F" box="[918,949,1572,1593]" italics="true" pageId="8" pageNumber="1189">L..</emphasis>
<taxonomicName id="4C724D55FFE08844F4B46C5714A47A0F" authority="Olivier, 1885" authorityName="Olivier" authorityYear="1885" box="[959,1255,1571,1593]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B906EAC4FFE08844F4B46C5714787A0E" box="[959,1083,1571,1592]" italics="true" pageId="8" pageNumber="1189">parallelus</emphasis>
Olivier, 1885
</taxonomicName>
,
<taxonomicName id="4C724D55FFE08844F3FD6C50138B7A61" authority="(Olivier, 1911)" authorityName="Bocakova &amp; Campello-Gonçalves &amp; Silveira" authorityYear="2022" baseAuthorityName="Olivier" baseAuthorityYear="1911" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="species" species="remixtus" status="comb. nov.">
<emphasis id="B906EAC4FFE08844F3FD6C5015D27A0E" bold="true" box="[1270,1425,1571,1593]" italics="true" pageId="8" pageNumber="1189">L. remixtus</emphasis>
(Olivier, 1911)
</taxonomicName>
<emphasis id="B906EAC4FFE08844F4C56C3614127A61" bold="true" box="[974,1105,1602,1623]" pageId="8" pageNumber="1189">
<taxonomicNameLabel id="A23557BFFFE08844F4C56C3614127A61" box="[974,1105,1602,1623]" pageId="8" pageNumber="1189" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C724D55FFE08844F3566C3715CD7A61" authority="Olivier, 1894" authorityName="Olivier" authorityYear="1894" box="[1117,1422,1602,1624]" class="Insecta" family="Lampyridae" genus="Ledocas" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1189" phylum="Arthropoda" rank="species" species="xanthomus">
<emphasis id="B906EAC4FFE08844F3566C3714B47A61" box="[1117,1271,1602,1624]" italics="true" pageId="8" pageNumber="1189">L. xanthomus</emphasis>
Olivier, 1894
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C368655DFFE08844F4226C0D13D97A9B" pageId="8" pageNumber="1189" type="distribution">
<paragraph id="8BCD36D6FFE08844F4226C0D13D97A9B" blockId="8.[809,1424,1657,1709]" pageId="8" pageNumber="1189">
<emphasis id="B906EAC4FFE08844F4226C0D13857AB8" box="[809,966,1657,1678]" italics="true" pageId="8" pageNumber="1189">Distribution:</emphasis>
<collectingCountry id="F3657646FFE08844F4D66C0D14187AB9" box="[989,1115,1657,1679]" name="Argentina" pageId="8" pageNumber="1189">Argentina</collectingCountry>
,
<collectingCountry id="F3657646FFE08844F3626C0D14F57AB9" box="[1129,1206,1657,1679]" name="Brazil" pageId="8" pageNumber="1189">Brazil</collectingCountry>
,
<collectingCountry id="F3657646FFE08844F3CE6C0D15437AB8" box="[1221,1280,1657,1678]" name="Peru" pageId="8" pageNumber="1189">Peru</collectingCountry>
,
<collectingCountry id="F3657646FFE08844F2056C0D15CF7AB9" box="[1294,1420,1657,1679]" name="Venezuela" pageId="8" pageNumber="1189">Venezuela</collectingCountry>
,
<collectingCountry id="F3657646FFE08844F4226CEC13D57A9B" box="[809,918,1688,1709]" name="Paraguay" pageId="8" pageNumber="1189">Paraguay</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>