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<document id="9C574F739F549D245C86669BD37C6192" ID-CLB-Dataset="20317" ID-DOI="10.11646/zootaxa.5105.4.5" ID-GBIF-Dataset="15ba8e38-a20d-4a8f-a9c3-95696d3f403b" ID-ISSN="1175-5326" ID-Zenodo-Dep="6333868" ID-ZooBank="ECE182B0-7053-450E-9CE7-29E3A1E81A89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646640578934" checkinUser="plazi" docAuthor="Kato, Taiga, Umemura, Shinya &amp; Nakano, Takafumi" docDate="2022" docId="03DB87B2E374FF84FF0418FCFABAFF3B" docLanguage="en" docName="zootaxa.5105.4.5.pdf" docOrigin="Zootaxa 5105 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Antrokoreana asuwaensis Kato &amp; Umemura &amp; Nakano 2022, sp. nov." docType="treatment" docUuid="CA391B59-C457-471B-8DEF-6BBF52DECC1E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="566" masterDocId="FFE2FFCAE377FF83FF93186BFFC1FFC1" masterDocTitle="A new species of the subterranean millipede genus Antrokoreana Verhoeff, 1938 from the Nanatsuoguchi Mine, central Honshu, Japan, and insights into the phylogenetic position of Antrokoreana (Diplopoda, Julida, Nemasomatidae)" masterLastPageNumber="570" masterPageNumber="559" pageNumber="562" updateTime="1699317307825" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CFF6FD56A728B5CB8539A92AB1653371">A new species of the subterranean millipede genus Antrokoreana Verhoeff, 1938 from the Nanatsuoguchi Mine, central Honshu, Japan, and insights into the phylogenetic position of Antrokoreana (Diplopoda, Julida, Nemasomatidae)</mods:title>
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<mods:namePart id="7F879AB0F7B9C23F44C49F6B2FD80E36">Kato, Taiga</mods:namePart>
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<mods:affiliation id="6D683ABECF93D0FE2D4E43EA1400507F">Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, Sakyo, Kyoto 606 - 8502, Japan. &amp; tkato @ zoo. zool. kyoto-u. ac. jp; https: // orcid. org / 0000 - 0001 - 7220 - 4502</mods:affiliation>
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<mods:namePart id="40FC245B234675C341F58E07DE37F83B">Umemura, Shinya</mods:namePart>
<mods:affiliation id="2604BDB34F697AD4C4A17DB3890097A1">Fukui City Museum of Natural History, 147 Asuwakami-cho, Fukui 918 - 8006, Japan.</mods:affiliation>
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<mods:namePart id="FD35FDA4A9920E9917CB277F1316069F">Nakano, Takafumi</mods:namePart>
<mods:affiliation id="1DDE5516758EFC3CA846F543A6954E0C">Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, Sakyo, Kyoto 606 - 8502, Japan.</mods:affiliation>
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<mods:date id="6350EABC43E3EF1C2ADA679A1C8D9D34">2022</mods:date>
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<taxonomicName id="4C724D27E374FF80FF0418FCFE10FF70" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[151,465,151,178]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E374FF80FF0418FCFE10FF70" bold="true" box="[151,465,151,178]" italics="true" pageId="3" pageNumber="562">Antrokoreana asuwaensis</emphasis>
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<figureCitation id="13492A21E374FF80FF0418D0FED5FF17" box="[151,276,187,214]" captionStart="FIGURE 1" captionStartId="3.[151,250,1720,1744]" captionTargetBox="[151,1436,1090,1696]" captionTargetId="figure-316@3.[151,1436,1090,1696]" captionTargetPageId="3" captionText="FIGURE 1. Antrokoreana asuwaensis sp. nov., in situ habitus, and its habitat at the type locality. A. Habitus of live male (not collected), dorsal view. B. Habitus of live male (not collected), lateral view. C. Collection site at the Nanatsuoguchi Mine in central Japan. Photographs were taken by Shinya Umemura (A, B), Taro Jonishi (C)." figureDoi="http://doi.org/10.5281/zenodo.6333870" httpUri="https://zenodo.org/record/6333870/files/figure.png" pageId="3" pageNumber="562">Figs 1A, B</figureCitation>
,
<figureCitation id="13492A21E374FF80FE8C18D0FE8BFF17" box="[287,330,187,214]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[151,250,1707,1731]" captionStartId-1="5.[151,250,1466,1490]" captionStartId-2="6.[151,250,964,988]" captionStartId-3="7.[151,250,1380,1404]" captionTargetBox-0="[151,1435,417,1684]" captionTargetBox-1="[151,1435,633,1443]" captionTargetBox-2="[157,1427,186,936]" captionTargetBox-3="[151,1436,273,1356]" captionTargetId-0="figure-143@4.[151,1435,417,1684]" captionTargetId-1="figure-210@5.[151,1435,633,1443]" captionTargetId-2="figure-19@6.[151,1436,181,941]" captionTargetId-3="figure-60@7.[151,1436,273,1356]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." captionText-1="FIGURE 3. Antrokoreana asuwaensis sp. nov., holotype ♂ (E, F: KUZ Z4004) and paratype ♂♂ (A, C, D: FCMNH-Ar213; B: FCMNH-Ar243). A. Body rings 5 and 6, lateral view. B. First leg pair, posterior view. C. Third leg pair, posterior view; white arrowheads indicate adhesive pads. D. Penes, posterior view. E. Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view. F. Telson, ventral view. Scale bars: 500 μm (A, E, F), 200 μm (B, C), 50 μm (D)." captionText-2="FIGURE 4. Antrokoreana asuwaensis sp. nov., holotype ♂ (KUZ Z4004). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. Scale bars: 250 μm." captionText-3="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi-0="http://doi.org/10.5281/zenodo.6333872" figureDoi-1="http://doi.org/10.5281/zenodo.6333874" figureDoi-2="http://doi.org/10.5281/zenodo.6333876" figureDoi-3="http://doi.org/10.5281/zenodo.6333878" httpUri-0="https://zenodo.org/record/6333872/files/figure.png" httpUri-1="https://zenodo.org/record/6333874/files/figure.png" httpUri-2="https://zenodo.org/record/6333876/files/figure.png" httpUri-3="https://zenodo.org/record/6333878/files/figure.png" pageId="3" pageNumber="562">25</figureCitation>
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urn:lsid:zoobank.org:act:
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<taxonomicName id="4C724D27E374FF80FF041975FE83FEF7" ID-CoL="7NQBY" box="[151,322,286,310]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B906EAB6E374FF80FF041975FEE3FEF7" box="[151,290,286,310]" italics="true" pageId="3" pageNumber="562">Antrokoreana</emphasis>
sp.
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<treatmentCitation id="0AD310B5E374FF80FEC81974FDB0FEF7" author="Umemura, S. &amp; Yoshizawa, Y. &amp; Ito, K." box="[347,625,286,310]" page="70" pageId="3" pageNumber="562" year="2019">
<bibRefCitation id="EFE34B55E374FF80FEC81974FDB0FEF7" author="Umemura, S. &amp; Yoshizawa, Y. &amp; Ito, K." box="[347,625,286,310]" pageId="3" pageNumber="562" pagination="63 - 74" refId="ref8607" refString="Umemura, S., Yoshizawa, Y. &amp; Ito, K. (2019) Arthropod fauna of Nanatsuoguchi mine in Mt. Asuwa, Fukui City, Fukui Prefecture. Bulletin of the Fukui City Museum of Natural History, 66, 63 - 74." type="journal article" year="2019">
Umemura
<emphasis id="B906EAB6E374FF80FE591974FE3CFEF7" box="[458,509,286,310]" italics="true" pageId="3" pageNumber="562">et al.</emphasis>
(2019: 70)
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.
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<emphasis id="B906EAB6E374FF80FF04190FFE41FEBF" bold="true" box="[151,384,356,382]" pageId="3" pageNumber="562">Material examined.</emphasis>
<materialsCitation id="3B1A3CF9E374FF80FE15190FFB37FE62" ID-GBIF-Occurrence="3698911303" collectingDate="2019-11-23" collectionCode="KUZ" collectorName="S. Umemura." country="Japan" county="Honshu" latitude="36.056007" location="Nanatsuoguchi Mine" longLatPrecision="1" longitude="136.19756" municipality="City" pageId="3" pageNumber="562" specimenCode="Z4004" specimenCount="1" specimenCount-male="1" stateProvince="Fukui" typeStatus="holotype">
<typeStatus id="54C98806E374FF80FE15190FFE32FEBF" box="[390,499,356,382]" pageId="3" pageNumber="562" type="holotype">Holotype</typeStatus>
:
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<specimenCode id="DBD49EDFE374FF80FDAE190EFD49FEBE" box="[573,648,357,383]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="562">Z4004</specimenCode>
,
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,
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(entrance:
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,
<geoCoordinate id="EE465063E374FF80FB53190FFAA4FEBE" box="[1216,1381,356,383]" degrees="136.197563" direction="east" orientation="longitude" pageId="3" pageNumber="562" precision="1" value="136.19756">136.197563°E</geoCoordinate>
;
<figureCitation id="13492A21E374FF80FAE3190FFF78FE62" captionStart="FIGURE 1" captionStartId="3.[151,250,1720,1744]" captionTargetBox="[151,1436,1090,1696]" captionTargetId="figure-316@3.[151,1436,1090,1696]" captionTargetPageId="3" captionText="FIGURE 1. Antrokoreana asuwaensis sp. nov., in situ habitus, and its habitat at the type locality. A. Habitus of live male (not collected), dorsal view. B. Habitus of live male (not collected), lateral view. C. Collection site at the Nanatsuoguchi Mine in central Japan. Photographs were taken by Shinya Umemura (A, B), Taro Jonishi (C)." figureDoi="http://doi.org/10.5281/zenodo.6333870" httpUri="https://zenodo.org/record/6333870/files/figure.png" pageId="3" pageNumber="562">Fig. 1C</figureCitation>
),
<location id="8EAD607FE374FF80FF5C19E2FE8CFE62" LSID="urn:lsid:plazi:treatment:03DB87B2E374FF84FF0418FCFABAFF3B:8EAD607FE374FF80FF5C19E2FE8CFE62" box="[207,333,393,419]" country="Japan" county="Honshu" latitude="36.056007" longLatPrecision="1" longitude="136.19756" municipality="City" name="Mt. Asuwa" pageId="3" pageNumber="562" stateProvince="Fukui">Mt. Asuwa</location>
,
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,
<collectingRegion id="49B6F846E374FF80FE4E19E3FD5AFE63" box="[477,667,392,418]" country="Japan" name="Fukui" pageId="3" pageNumber="562">Fukui Prefecture</collectingRegion>
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,
<collectingCountry id="F3657634E374FF80FC9819E2FC8FFE62" box="[779,846,393,419]" name="Japan" pageId="3" pageNumber="562">Japan</collectingCountry>
,
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<collectingDate id="EF88E98CE374FF80FCCA19E2FC1FFE62" box="[857,990,393,419]" pageId="3" pageNumber="562" value="2019-11-23">2019-11-23</collectingDate>
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/12-11, leg.
<collectorName id="26875372E374FF80FBF219E2FB37FE62" box="[1121,1270,393,419]" pageId="3" pageNumber="562">S. Umemura.</collectorName>
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<typeStatus id="54C98806E374FF80FF5419C6FEF9FE06" box="[199,312,429,455]" pageId="3" pageNumber="562" type="paratype">Paratypes</typeStatus>
:
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collected from the same locality
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. FCMNH-Ar131,
<specimenCount id="9D74FD2DE374FF80FB8219C6FBFAFE06" box="[1041,1083,429,455]" pageId="3" pageNumber="562" type="female">1 ♀</specimenCount>
,
<date id="FFCC1064E374FF80FBDB19C6FB0CFE06" box="[1096,1229,429,455]" pageId="3" pageNumber="562" value="2017-02-12">2017-12-02</date>
, leg. S. Umemura; FCMNH-Ar132,
<specimenCount id="9D74FD2DE374FF80FECF19BAFE61FE2A" box="[348,416,465,491]" pageId="3" pageNumber="562" type="male">2 ♂♂</specimenCount>
,
<date id="FFCC1064E374FF80FE3D19BAFDF2FE2A" box="[430,563,465,491]" pageId="3" pageNumber="562" value="2018-02-02">2018-02-02</date>
/03-02, leg. S. Umemura; FCMNH-Ar206,
<specimenCount id="9D74FD2DE374FF80FBB719BAFBA9FE2A" box="[1060,1128,465,491]" pageId="3" pageNumber="562" type="male">2 ♂♂</specimenCount>
,
<date id="FFCC1064E374FF80FBE519BBFB3BFE2A" box="[1142,1274,464,491]" pageId="3" pageNumber="562" value="2018-03-28">2018-03-28</date>
/04-05, leg. S. Umemura; FCMNH-Ar207,
<specimenCount id="9D74FD2DE374FF80FE4A199EFDC4FDCE" box="[473,517,501,527]" pageId="3" pageNumber="562" type="male">1 ♂</specimenCount>
,
<date id="FFCC1064E374FF80FD81199FFD57FDCF" box="[530,662,500,526]" pageId="3" pageNumber="562" value="2018-02-02">2018-02-02</date>
/02-26, leg. S. Umemura; FCMNH-Ar208,
<specimenCount id="9D74FD2DE374FF80FBEF199EFB7CFDCE" box="[1148,1213,501,527]" pageId="3" pageNumber="562" type="male">2 ♂♂</specimenCount>
,
<date id="FFCC1064E374FF80FB5A199EFA8CFDCE" box="[1225,1357,501,527]" pageId="3" pageNumber="562" value="2017-12-14">2017-12-14</date>
/12-22, leg. S. Umemura; FCMNH-Ar212,
<specimenCount id="9D74FD2DE374FF80FDB61A72FDA2FDF2" box="[549,611,537,563]" pageId="3" pageNumber="562" type="female">2 ♀♀</specimenCount>
,
<date id="FFCC1064E374FF80FDE21A72FD34FDF2" box="[625,757,537,563]" pageId="3" pageNumber="562" value="2019-02-27">2019-02-27</date>
/03-22, leg. S. Umemura; FCMNH-Ar213,
<specimenCount id="9D74FD2DE374FF80FB4B1A72FAD9FDF2" box="[1240,1304,537,563]" pageId="3" pageNumber="562" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D74FD2DE374FF80FAB71A72FA8CFDF2" box="[1316,1357,537,563]" pageId="3" pageNumber="562" type="female">1 ♀</specimenCount>
,
<date id="FFCC1064E374FF80FAC81A72FF18FD96" pageId="3" pageNumber="562" value="2018-11-28">2018- 11-28</date>
/12-13, leg. S. Umemura; FCMNH-Ar228,
<materialsCitation id="3B1A3CF9E374FF80FD2D1A56FB49FD97" ID-GBIF-Occurrence="3698911302" box="[702,1160,572,599]" pageId="3" pageNumber="562" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D74FD2DE374FF80FD2D1A56FCC1FD96" box="[702,768,573,599]" pageId="3" pageNumber="562" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D74FD2DE374FF80FC9F1A56FC8AFD96" box="[780,843,573,599]" pageId="3" pageNumber="562" type="female">2 ♀♀</specimenCount>
, same data as for preceding
</materialsCitation>
; FCMNH-Ar232,
<specimenCount id="9D74FD2DE374FF80FAC51A56FA56FD96" box="[1366,1431,573,599]" pageId="3" pageNumber="562" type="male">2 ♂♂</specimenCount>
,
<date id="FFCC1064E374FF80FF041A0AFEDDFDBA" box="[151,284,609,635]" pageId="3" pageNumber="562" value="2018-09-11">2018-11-09</date>
/11-28, leg. S. Umemura; FCMNH-Ar240,
<specimenCount id="9D74FD2DE374FF80FD6C1A0AFCEAFDBA" box="[767,811,609,635]" pageId="3" pageNumber="562" type="male">1 ♂</specimenCount>
,
<date id="FFCC1064E374FF80FCA41A0AFC7AFDBA" box="[823,955,609,635]" pageId="3" pageNumber="562" value="2019-11-23">2019-11-23</date>
, leg. S. Umemura; FCMNH-Ar243,
<materialsCitation id="3B1A3CF9E374FF80FAC51A0AFDC2FD5F" ID-GBIF-Occurrence="3698911301" collectingDate="2019-11-23" collectionCode="KUZ" collectorName="S. Umemura." country="Japan" county="Honshu" latitude="36.056007" location="Nanatsuoguchi Mine" longLatPrecision="1" longitude="136.19756" municipality="City" pageId="3" pageNumber="562" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Fukui" typeStatus="paratype">
<specimenCount id="9D74FD2DE374FF80FAC51A0AFA56FDBA" box="[1366,1431,609,635]" pageId="3" pageNumber="562" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D74FD2DE374FF80FF041AEEFF17FD5E" box="[151,214,645,671]" pageId="3" pageNumber="562" type="female">3 ♀♀</specimenCount>
, same data as for holotype
</materialsCitation>
; FCMNH-Ar245,
<specimenCount id="9D74FD2DE374FF80FD431AEEFD38FD5E" box="[720,761,645,671]" pageId="3" pageNumber="562" type="female">1 ♀</specimenCount>
,
<date id="FFCC1064E374FF80FC941AEEFC4DFD5E" box="[775,908,645,671]" pageId="3" pageNumber="562" value="2018-11-28">2018-11-28</date>
/12-13, leg. S. Umemura; FCMNH-Ar246,
<specimenCount id="9D74FD2DE374FF80FAFF1AEEFA56FD5E" box="[1388,1431,645,671]" pageId="3" pageNumber="562" type="male">1 ♂</specimenCount>
,
<date id="FFCC1064E374FF80FF041AC2FED8FD02" box="[151,281,681,707]" pageId="3" pageNumber="562" value="2019-11-12">2019-12-11</date>
/12-24, leg. S. Umemura; FCMNH-Ar250,
<materialsCitation id="3B1A3CF9E374FF80FC931AC2FB9DFD03" ID-GBIF-Occurrence="3698911307" box="[768,1116,680,707]" collectingDate="2019-11-23" collectionCode="KUZ" collectorName="S. Umemura." country="Japan" county="Honshu" latitude="36.056007" location="Nanatsuoguchi Mine" longLatPrecision="1" longitude="136.19756" municipality="City" pageId="3" pageNumber="562" specimenCount="1" specimenCount-female="1" stateProvince="Fukui" typeStatus="paratype">
<specimenCount id="9D74FD2DE374FF80FC931AC2FCEAFD02" box="[768,811,681,707]" pageId="3" pageNumber="562" type="female">1 ♀</specimenCount>
, same data as for holotype
</materialsCitation>
; FCMNH-Ar251,
<materialsCitation id="3B1A3CF9E374FF80FAB91AC2FEB8FD27" ID-GBIF-Occurrence="3698911304" collectingDate="2019-11-23" collectionCode="KUZ" collectorName="S. Umemura." country="Japan" county="Honshu" latitude="36.056007" location="Nanatsuoguchi Mine" longLatPrecision="1" longitude="136.19756" municipality="City" pageId="3" pageNumber="562" specimenCount="1" specimenCount-male="1" stateProvince="Fukui" typeStatus="paratype">
<specimenCount id="9D74FD2DE374FF80FAB91AC2FA97FD02" box="[1322,1366,681,707]" pageId="3" pageNumber="562" type="male">1 ♂</specimenCount>
, same data as for holotype
</materialsCitation>
; FCMNH-Ar252,
<materialsCitation id="3B1A3CF9E374FF80FDD51AA6FC61FD27" ID-GBIF-Occurrence="3698911305" box="[582,928,716,743]" collectingDate="2019-11-23" collectionCode="KUZ" collectorName="S. Umemura." country="Japan" county="Honshu" latitude="36.056007" location="Nanatsuoguchi Mine" longLatPrecision="1" longitude="136.19756" municipality="City" pageId="3" pageNumber="562" specimenCount="1" specimenCount-female="1" stateProvince="Fukui" typeStatus="paratype">
<specimenCount id="9D74FD2DE374FF80FDD51AA6FDB1FD26" box="[582,624,717,743]" pageId="3" pageNumber="562" type="female">1 ♀</specimenCount>
, same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C368652FE374FF80FF541A9BFE68FC93" pageId="3" pageNumber="562" type="diagnosis">
<paragraph id="8BCD36A4E374FF80FF541A9BFE68FC93" blockId="3.[151,1437,356,1066]" pageId="3" pageNumber="562">
<emphasis id="B906EAB6E374FF80FF541A9BFE83FCCB" bold="true" box="[199,322,752,778]" pageId="3" pageNumber="562">Diagnosis.</emphasis>
The new species is clearly distinguishable from all congeners by each coxal process of the anterior gonopods being expanded distally in lateral view, its distal part concave, and its posterior part curved and directed mesad in posterior view.
</paragraph>
</subSubSection>
<subSubSection id="C368652FE374FF85FF541B37FD57FBAC" lastPageId="6" lastPageNumber="565" pageId="3" pageNumber="562" type="description">
<paragraph id="8BCD36A4E374FF80FF541B37FE98FCB7" blockId="3.[151,1437,356,1066]" box="[199,345,860,886]" pageId="3" pageNumber="562">
<emphasis id="B906EAB6E374FF80FF541B37FE98FCB7" bold="true" box="[199,345,860,886]" pageId="3" pageNumber="562">Description.</emphasis>
</paragraph>
<paragraph id="8BCD36A4E374FF80FF041BEAFF23FBEB" blockId="3.[151,1437,356,1066]" pageId="3" pageNumber="562">
<emphasis id="B906EAB6E374FF80FF041BEAFE2CFC5B" box="[151,493,896,923]" italics="true" pageId="3" pageNumber="562">Size and number of body rings</emphasis>
.
<typeStatus id="54C98806E374FF80FE681BEBFDA2FC5B" box="[507,611,896,922]" pageId="3" pageNumber="562" type="holotype">Holotype</typeStatus>
male:
<quantity id="4C8A9B41E374FF80FD3C1BEAFCD2FC5A" box="[687,787,897,923]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="3" pageNumber="562" unit="mm" value="22.3">22.3 mm</quantity>
long, vertical diameter of largest body ring 1.0 mm; body with 33 podous rings + 1 apodous rings + telson.
<typeStatus id="54C98806E374FF80FD5B1BCEFCEBFC7E" box="[712,810,933,959]" pageId="3" pageNumber="562" type="paratype">Paratype</typeStatus>
males:
<quantity id="4C8A9B41E374FF80FC111BCFFBE7FC7E" box="[898,1062,932,959]" metricMagnitude="-2" metricUnit="m" metricValue="2.535" metricValueMax="2.92" metricValueMin="2.15" pageId="3" pageNumber="562" unit="mm" value="25.35" valueMax="29.2" valueMin="21.5">21.529.2 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4C8A9B41E374FF80FE9C1BA2FE5DFC22" box="[271,412,969,995]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="3" pageNumber="562" unit="mm" value="1.5" valueMax="1.8" valueMin="1.2">1.21.8 mm</quantity>
; body with 3236 podous rings + 1 apodous rings + telson.
<typeStatus id="54C98806E374FF80FBCB1BA2FB7BFC22" box="[1112,1210,969,995]" pageId="3" pageNumber="562" type="paratype">Paratype</typeStatus>
females:
<quantity id="4C8A9B41E374FF80FABF1BA3FF02FBC6" metricMagnitude="-2" metricUnit="m" metricValue="2.5149999999999997" metricValueMax="2.76" metricValueMin="2.27" pageId="3" pageNumber="562" unit="mm" value="25.15" valueMax="27.6" valueMin="22.7">22.727.6 mm</quantity>
long, vertical diameter of largest body ring
<quantity id="4C8A9B41E374FF80FD2F1B86FC86FBC6" box="[700,839,1005,1031]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.9" metricValueMin="1.2" pageId="3" pageNumber="562" unit="mm" value="1.5499999999999998" valueMax="1.9" valueMin="1.2">1.21.9 mm</quantity>
; body with 3133 podous rings + 1 apodous rings + telson.
</paragraph>
<caption id="DF0D662CE374FF80FF041ED3FC32F8D9" ID-DOI="http://doi.org/10.5281/zenodo.6333870" ID-Zenodo-Dep="6333870" httpUri="https://zenodo.org/record/6333870/files/figure.png" pageId="3" pageNumber="562" startId="3.[151,250,1720,1744]" targetBox="[151,1436,1090,1696]" targetPageId="3">
<paragraph id="8BCD36A4E374FF80FF041ED3FC32F8D9" blockId="3.[151,1437,1720,1816]" pageId="3" pageNumber="562">
<emphasis id="B906EAB6E374FF80FF041ED3FED5F911" bold="true" box="[151,276,1720,1744]" pageId="3" pageNumber="562">FIGURE 1.</emphasis>
<taxonomicName id="4C724D27E374FF80FE891ED3FDDEF911" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[282,543,1720,1744]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E374FF80FE891ED3FDDEF911" box="[282,543,1720,1744]" italics="true" pageId="3" pageNumber="562">Antrokoreana asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE374FF80FDB51ED3FDB9F90E" box="[550,632,1720,1743]" pageId="3" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="B906EAB6E374FF80FD111ED2FD03F911" box="[642,706,1721,1744]" italics="true" pageId="3" pageNumber="562">in situ</emphasis>
habitus, and its habitat at the type locality.
<emphasis id="B906EAB6E374FF80FBEE1ED3FB57F911" bold="true" box="[1149,1174,1720,1744]" pageId="3" pageNumber="562">A.</emphasis>
Habitus of live male (not collected), dorsal view.
<emphasis id="B906EAB6E374FF80FE1E1EB7FE65F935" bold="true" box="[397,420,1756,1780]" pageId="3" pageNumber="562">B.</emphasis>
Habitus of live male (not collected), lateral view.
<emphasis id="B906EAB6E374FF80FC3D1EB7FC07F935" bold="true" box="[942,966,1756,1780]" pageId="3" pageNumber="562">C.</emphasis>
Collection site at the Nanatsuoguchi Mine in central Japan. Photographs were taken by Shinya Umemura (
<emphasis id="B906EAB6E374FF80FC901F6BFCF2F8D9" bold="true" box="[771,819,1792,1816]" pageId="3" pageNumber="562">A, B</emphasis>
), Taro Jonishi (
<emphasis id="B906EAB6E374FF80FC411F6BFC25F8D9" bold="true" box="[978,996,1792,1816]" pageId="3" pageNumber="562">C</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BCD36A4E374FF87FF541F2DFA85FEA4" blockId="3.[151,1436,1862,2033]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="563" pageId="3" pageNumber="562">
<emphasis id="B906EAB6E374FF80FF541F2DFEC5F8A1" box="[199,260,1862,1888]" italics="true" pageId="3" pageNumber="562">Head</emphasis>
(
<figureCitation id="13492A21E374FF80FE801F2DFE99F8A1" box="[275,344,1862,1888]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">Fig. 2</figureCitation>
). Without ommatidia (
<figureCitation id="13492A21E374FF80FDC71F2DFD13F8A0" box="[596,722,1862,1889]" captionStart="FIGURE 1" captionStartId="3.[151,250,1720,1744]" captionTargetBox="[151,1436,1090,1696]" captionTargetId="figure-316@3.[151,1436,1090,1696]" captionTargetPageId="3" captionText="FIGURE 1. Antrokoreana asuwaensis sp. nov., in situ habitus, and its habitat at the type locality. A. Habitus of live male (not collected), dorsal view. B. Habitus of live male (not collected), lateral view. C. Collection site at the Nanatsuoguchi Mine in central Japan. Photographs were taken by Shinya Umemura (A, B), Taro Jonishi (C)." figureDoi="http://doi.org/10.5281/zenodo.6333870" httpUri="https://zenodo.org/record/6333870/files/figure.png" pageId="3" pageNumber="562">Figs 1A, B</figureCitation>
,
<figureCitation id="13492A21E374FF80FD4D1F2DFCC5F8A1" box="[734,772,1862,1888]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">2A</figureCitation>
); with 4 frontal setae. Labrum (
<figureCitation id="13492A21E374FF80FBF61F2DFB7AF8A1" box="[1125,1211,1862,1888]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">Fig. 2B</figureCitation>
) with 3 labral teeth, 4 supralabral setae and 15 labral setae (
<typeStatus id="54C98806E374FF80FDCF1F01FD04F844" box="[604,709,1898,1925]" pageId="3" pageNumber="562" type="holotype">holotype</typeStatus>
). Mandibular cardo with 1 seta; mandibular stipes with 2 setae, unexpanded in males. Gnathal lobe (
<figureCitation id="13492A21E374FF80FDA11FE5FD6DF869" box="[562,684,1934,1960]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">Fig. 2CE</figureCitation>
): external tooth with 3 cusps; internal tooth with 5 cusps (
<figureCitation id="13492A21E374FF80FAA51FE5FA4EF869" box="[1334,1423,1934,1960]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">Fig. 2C</figureCitation>
); 6 (4 complete + 2 incomplete) pectinate lamellae (
<figureCitation id="13492A21E374FF80FD401FD9FCECF80D" box="[723,813,1970,1996]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">Fig. 2D</figureCitation>
), posterior teeth of each lamella with 2 or 3 finger-like branches (
<figureCitation id="13492A21E374FF80FE991FBDFEA3F831" box="[266,354,2006,2032]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">Fig. 2E</figureCitation>
). Gnathochilarium (
<figureCitation id="13492A21E374FF80FDD01FBDFD56F831" box="[579,663,2006,2032]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="3" pageNumber="562">Fig. 2F</figureCitation>
) with triangular promentum. Lamellae linguales each with 4 setae in row, stipites each with 3 distolateral setae, without short medial setae. Antennae (
<figureCitation id="13492A21E373FF87FBA618FCFB64FF70" box="[1077,1189,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="4" pageNumber="563">Fig. 2GI</figureCitation>
) in
<typeStatus id="54C98806E373FF87FB4718FCFAF8FF70" box="[1236,1337,151,177]" pageId="4" pageNumber="563" type="holotype">holotype</typeStatus>
,
<quantity id="4C8A9B41E373FF87FAD618FCFA5CFF70" box="[1349,1437,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="563" unit="mm" value="2.8">2.8 mm</quantity>
long, ~2.8× longer than vertical diameter of largest body ring, with 4 apical cones. Lengths of antennomeres IVIII (VIII = apical cones) (in mm): 0.16 (I), 0.48 (II), 0.55 (III), 0.42 (IV), 0.41 (V), 0.39 (VI), 0.38 (VII) and 0.02 (VIII). Length/width ratio of antennomeres IVII: 1.2 (I), 3.1 (II), 4.3 (III), 3.4 (IV), 2.7 (V), 2.4 (VI) and 2.7 (VII). Antennomeres VVI (
<figureCitation id="13492A21E373FF87FE00194CFE2DFE80" box="[403,492,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="4" pageNumber="563">Fig. 2H</figureCitation>
) each with terminal corolla of large sensilla basiconica bacilliformia; antennomere VII (
<figureCitation id="13492A21E373FF87FF5E1920FEDBFEA4" box="[205,282,331,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1435,417,1684]" captionTargetId="figure-143@4.[151,1435,417,1684]" captionTargetPageId="4" captionText="FIGURE 2. Antrokoreana asuwaensis sp. nov., paratype ♂♂ (A, B, F: FCMNH-Ar213; GI: FCMNH-Ar243) and paratype ♀ (CE: FCMNH-Ar243). A. Head, frontal view. B. Labrum, frontal view. C. Right mandible, ventral view. D. Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae. E. Mandibular gnathal lobe of right mandible, posterior view. F. Gnathochilarium, ventral view. G. Left antenna, sub-mesal view. H. Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla. I. Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (A, B, F, H), 100 μm (C, I), 50 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="4" pageNumber="563">Fig. 2I</figureCitation>
) whole surface covered with numerous short sensilla chaetica among longer sensilla trichodea.
</paragraph>
<paragraph id="8BCD36A4E373FF87FF541904FB28FE48" blockId="4.[151,1437,151,393]" box="[199,1257,367,393]" pageId="4" pageNumber="563">
<emphasis id="B906EAB6E373FF87FF541904FED8FE48" box="[199,281,367,393]" italics="true" pageId="4" pageNumber="563">Collum</emphasis>
. With whorl of 8 setae located about one-third of collum length from posterior margin.
</paragraph>
<caption id="DF0D662CE373FF87FF041EC0FE08F85D" ID-DOI="http://doi.org/10.5281/zenodo.6333872" ID-Zenodo-Dep="6333872" httpUri="https://zenodo.org/record/6333872/files/figure.png" pageId="4" pageNumber="563" startId="4.[151,250,1707,1731]" targetBox="[151,1435,417,1684]" targetPageId="4">
<paragraph id="8BCD36A4E373FF87FF041EC0FE08F85D" blockId="4.[151,1437,1707,1948]" pageId="4" pageNumber="563">
<emphasis id="B906EAB6E373FF87FF041EC0FED5F902" bold="true" box="[151,276,1707,1731]" pageId="4" pageNumber="563">FIGURE 2.</emphasis>
<taxonomicName id="4C724D27E373FF87FE8F1EC7FDE0F902" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[284,545,1708,1732]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="4" pageNumber="563" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E373FF87FE8F1EC7FDE0F902" box="[284,545,1708,1732]" italics="true" pageId="4" pageNumber="563">Antrokoreana asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE373FF87FDBA1EC7FDBAF902" box="[553,635,1708,1731]" pageId="4" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♂♂ (
<emphasis id="B906EAB6E373FF87FC881EC0FCB3F905" bold="true" box="[795,882,1707,1732]" pageId="4" pageNumber="563">A, B, F:</emphasis>
FCMNH-Ar213;
<emphasis id="B906EAB6E373FF87FBB81EC0FB9FF902" bold="true" box="[1067,1118,1707,1731]" pageId="4" pageNumber="563">GI:</emphasis>
FCMNH-Ar243) and paratype ♀ (
<emphasis id="B906EAB6E373FF87FF281EA4FF32F926" bold="true" box="[187,243,1743,1767]" pageId="4" pageNumber="563">CE:</emphasis>
FCMNH-Ar243).
<emphasis id="B906EAB6E373FF87FE261EA4FE0FF926" bold="true" box="[437,462,1743,1767]" pageId="4" pageNumber="563">A.</emphasis>
Head, frontal view.
<emphasis id="B906EAB6E373FF87FD301EBBFD7AF929" bold="true" box="[675,699,1744,1768]" pageId="4" pageNumber="563">B.</emphasis>
Labrum, frontal view.
<emphasis id="B906EAB6E373FF87FC381EA4FC05F926" bold="true" box="[939,964,1743,1767]" pageId="4" pageNumber="563">C.</emphasis>
Right mandible, ventral view.
<emphasis id="B906EAB6E373FF87FA961EBBFADFF929" bold="true" box="[1285,1310,1744,1768]" pageId="4" pageNumber="563">D.</emphasis>
Mandibular gnathal lobe of left mandible, anterior view; white arrowheads indicate complete pectinate lamellae, black arrowheads denote incomplete lamellae.
<emphasis id="B906EAB6E373FF87FEE01F73FE4BF8F1" bold="true" box="[371,394,1816,1840]" pageId="4" pageNumber="563">E.</emphasis>
Mandibular gnathal lobe of right mandible, posterior view.
<emphasis id="B906EAB6E373FF87FC611F73FBC4F8F1" bold="true" box="[1010,1029,1816,1840]" pageId="4" pageNumber="563">F.</emphasis>
Gnathochilarium, ventral view.
<emphasis id="B906EAB6E373FF87FAC21F7CFAAAF8EE" bold="true" box="[1361,1387,1815,1839]" pageId="4" pageNumber="563">G.</emphasis>
Left antenna, sub-mesal view.
<emphasis id="B906EAB6E373FF87FE331F57FE7BF895" bold="true" box="[416,442,1852,1876]" pageId="4" pageNumber="563">H.</emphasis>
Antennomeres VVI, showing large sensilla basiconica bacilliformia at each terminal corolla.
<emphasis id="B906EAB6E373FF87FA1F1F57FA5DF895" bold="true" box="[1420,1436,1852,1876]" pageId="4" pageNumber="563">I.</emphasis>
Antennomere VII, showing short sensilla chaetica and longer sensilla trichodea. Scale bars: 200 μm (
<emphasis id="B906EAB6E373FF87FB301F34FAD1F8B9" bold="true" box="[1187,1296,1887,1912]" pageId="4" pageNumber="563">A, B, F, H</emphasis>
), 100 μm (
<emphasis id="B906EAB6E373FF87FA171F34FF60F85D" bold="true" pageId="4" pageNumber="563">C, I</emphasis>
), 50 μm (
<emphasis id="B906EAB6E373FF87FE951FEFFEF7F85D" bold="true" box="[262,310,1924,1948]" pageId="4" pageNumber="563">D, E</emphasis>
), 500 μm (
<emphasis id="B906EAB6E373FF87FE341FE8FE7AF85A" bold="true" box="[423,443,1923,1947]" pageId="4" pageNumber="563">G</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BCD36A4E372FF86FF5418FCFD1CFE80" blockId="5.[151,1436,151,609]" pageId="5" pageNumber="564">
<emphasis id="B906EAB6E372FF86FF5418FCFE82FF71" box="[199,323,151,177]" italics="true" pageId="5" pageNumber="564">Body rings</emphasis>
(
<figureCitation id="13492A21E372FF86FEC118FCFE6CFF70" box="[338,429,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1466,1490]" captionTargetBox="[151,1435,633,1443]" captionTargetId="figure-210@5.[151,1435,633,1443]" captionTargetPageId="5" captionText="FIGURE 3. Antrokoreana asuwaensis sp. nov., holotype ♂ (E, F: KUZ Z4004) and paratype ♂♂ (A, C, D: FCMNH-Ar213; B: FCMNH-Ar243). A. Body rings 5 and 6, lateral view. B. First leg pair, posterior view. C. Third leg pair, posterior view; white arrowheads indicate adhesive pads. D. Penes, posterior view. E. Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view. F. Telson, ventral view. Scale bars: 500 μm (A, E, F), 200 μm (B, C), 50 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.6333874" httpUri="https://zenodo.org/record/6333874/files/figure.png" pageId="5" pageNumber="564">Fig. 3A</figureCitation>
). Prozona to metazona of each ring with 48 transverse furrows on ventral half of lateral surfaces. Metazona dorsal surface smooth, ventral surface with 48 longitudinal striae; with posterior marginal whorl of setae, length of mid-body setae ~1.2% of vertical diameter of rings; one pair of ozopores present at center of lateral surface of each metazona of 6th ring to ultimate podous ring. (
<figureCitation id="13492A21E372FF86FC211968FBEBFEDC" box="[946,1066,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1720,1744]" captionTargetBox="[151,1436,1090,1696]" captionTargetId="figure-316@3.[151,1436,1090,1696]" captionTargetPageId="3" captionText="FIGURE 1. Antrokoreana asuwaensis sp. nov., in situ habitus, and its habitat at the type locality. A. Habitus of live male (not collected), dorsal view. B. Habitus of live male (not collected), lateral view. C. Collection site at the Nanatsuoguchi Mine in central Japan. Photographs were taken by Shinya Umemura (A, B), Taro Jonishi (C)." figureDoi="http://doi.org/10.5281/zenodo.6333870" httpUri="https://zenodo.org/record/6333870/files/figure.png" pageId="5" pageNumber="564">Fig. 1A, B</figureCitation>
). Sternum of each ring, free from pleurotergum, with wing-like expansions anteriorly.
</paragraph>
<paragraph id="8BCD36A4E372FF86FF541920FE66FE34" blockId="5.[151,1436,151,609]" pageId="5" pageNumber="564">
<emphasis id="B906EAB6E372FF86FF541920FF3DFEA5" box="[199,252,331,356]" italics="true" pageId="5" pageNumber="564">Legs</emphasis>
(
<figureCitation id="13492A21E372FF86FE9E1920FE4BFEA4" box="[269,394,331,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1466,1490]" captionTargetBox="[151,1435,633,1443]" captionTargetId="figure-210@5.[151,1435,633,1443]" captionTargetPageId="5" captionText="FIGURE 3. Antrokoreana asuwaensis sp. nov., holotype ♂ (E, F: KUZ Z4004) and paratype ♂♂ (A, C, D: FCMNH-Ar213; B: FCMNH-Ar243). A. Body rings 5 and 6, lateral view. B. First leg pair, posterior view. C. Third leg pair, posterior view; white arrowheads indicate adhesive pads. D. Penes, posterior view. E. Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view. F. Telson, ventral view. Scale bars: 500 μm (A, E, F), 200 μm (B, C), 50 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.6333874" httpUri="https://zenodo.org/record/6333874/files/figure.png" pageId="5" pageNumber="564">Fig. 3BD</figureCitation>
). In
<typeStatus id="54C98806E372FF86FE531920FDE4FEA4" box="[448,549,331,357]" pageId="5" pageNumber="564" type="holotype">holotype</typeStatus>
, mid-body legs ~1.7× longer than vertical diameter of ring; prefemur to claw in length ratio of 1.0:1.2:1.2:1.0:1.4:0.3 (leg-pair 10). Claw slender, ~7.0× longer than height (
<typeStatus id="54C98806E372FF86FB781904FA92FE4B" box="[1259,1363,367,394]" pageId="5" pageNumber="564" type="holotype">holotype</typeStatus>
); with accessory claw. In males, tibia of leg-pair 1 with 1 long spine directed proximad (
<figureCitation id="13492A21E372FF86FBBA19F8FB40FE6C" box="[1065,1153,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1466,1490]" captionTargetBox="[151,1435,633,1443]" captionTargetId="figure-210@5.[151,1435,633,1443]" captionTargetPageId="5" captionText="FIGURE 3. Antrokoreana asuwaensis sp. nov., holotype ♂ (E, F: KUZ Z4004) and paratype ♂♂ (A, C, D: FCMNH-Ar213; B: FCMNH-Ar243). A. Body rings 5 and 6, lateral view. B. First leg pair, posterior view. C. Third leg pair, posterior view; white arrowheads indicate adhesive pads. D. Penes, posterior view. E. Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view. F. Telson, ventral view. Scale bars: 500 μm (A, E, F), 200 μm (B, C), 50 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.6333874" httpUri="https://zenodo.org/record/6333874/files/figure.png" pageId="5" pageNumber="564">Fig. 3B</figureCitation>
). Femur, post-femur, and tibia of following legs each with adhesive pad (
<figureCitation id="13492A21E372FF86FD3519DCFD3FFE10" box="[678,766,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1466,1490]" captionTargetBox="[151,1435,633,1443]" captionTargetId="figure-210@5.[151,1435,633,1443]" captionTargetPageId="5" captionText="FIGURE 3. Antrokoreana asuwaensis sp. nov., holotype ♂ (E, F: KUZ Z4004) and paratype ♂♂ (A, C, D: FCMNH-Ar213; B: FCMNH-Ar243). A. Body rings 5 and 6, lateral view. B. First leg pair, posterior view. C. Third leg pair, posterior view; white arrowheads indicate adhesive pads. D. Penes, posterior view. E. Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view. F. Telson, ventral view. Scale bars: 500 μm (A, E, F), 200 μm (B, C), 50 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.6333874" httpUri="https://zenodo.org/record/6333874/files/figure.png" pageId="5" pageNumber="564">Fig. 3C</figureCitation>
). Penes (
<figureCitation id="13492A21E372FF86FCF219DCFC79FE10" box="[865,952,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1466,1490]" captionTargetBox="[151,1435,633,1443]" captionTargetId="figure-210@5.[151,1435,633,1443]" captionTargetPageId="5" captionText="FIGURE 3. Antrokoreana asuwaensis sp. nov., holotype ♂ (E, F: KUZ Z4004) and paratype ♂♂ (A, C, D: FCMNH-Ar213; B: FCMNH-Ar243). A. Body rings 5 and 6, lateral view. B. First leg pair, posterior view. C. Third leg pair, posterior view; white arrowheads indicate adhesive pads. D. Penes, posterior view. E. Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view. F. Telson, ventral view. Scale bars: 500 μm (A, E, F), 200 μm (B, C), 50 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.6333874" httpUri="https://zenodo.org/record/6333874/files/figure.png" pageId="5" pageNumber="564">Fig. 3D</figureCitation>
) shorter than coxae of leg-pair 2, each with 13 microsetae apically.
</paragraph>
<paragraph id="8BCD36A4E372FF86FF541994FF16FDA0" blockId="5.[151,1436,151,609]" pageId="5" pageNumber="564">
<emphasis id="B906EAB6E372FF86FF541994FECEFDD8" box="[199,271,511,537]" italics="true" pageId="5" pageNumber="564">Telson</emphasis>
(
<figureCitation id="13492A21E372FF86FE8C1994FE55FDD8" box="[287,404,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1466,1490]" captionTargetBox="[151,1435,633,1443]" captionTargetId="figure-210@5.[151,1435,633,1443]" captionTargetPageId="5" captionText="FIGURE 3. Antrokoreana asuwaensis sp. nov., holotype ♂ (E, F: KUZ Z4004) and paratype ♂♂ (A, C, D: FCMNH-Ar213; B: FCMNH-Ar243). A. Body rings 5 and 6, lateral view. B. First leg pair, posterior view. C. Third leg pair, posterior view; white arrowheads indicate adhesive pads. D. Penes, posterior view. E. Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view. F. Telson, ventral view. Scale bars: 500 μm (A, E, F), 200 μm (B, C), 50 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.6333874" httpUri="https://zenodo.org/record/6333874/files/figure.png" pageId="5" pageNumber="564">Fig. 3E, F</figureCitation>
). Preanal ring without projection, with marginal whorl of setae and 5 additional dorsal setae in front of marginal whorl. Anal valves rounded, each with 7 setae; subanal scale without any modifications, with 2 setae.
</paragraph>
<caption id="DF0D662CE372FF86FF041DD1FB8EF9FF" ID-DOI="http://doi.org/10.5281/zenodo.6333874" ID-Zenodo-Dep="6333874" httpUri="https://zenodo.org/record/6333874/files/figure.png" pageId="5" pageNumber="564" startId="5.[151,250,1466,1490]" targetBox="[151,1435,633,1443]" targetPageId="5">
<paragraph id="8BCD36A4E372FF86FF041DD1FB8EF9FF" blockId="5.[151,1437,1466,1599]" pageId="5" pageNumber="564">
<emphasis id="B906EAB6E372FF86FF041DD1FED3FA13" bold="true" box="[151,274,1466,1490]" pageId="5" pageNumber="564">FIGURE 3.</emphasis>
<taxonomicName id="4C724D27E372FF86FE8B1DD1FDDAFA13" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[280,539,1466,1490]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E372FF86FE8B1DD1FDDAFA13" box="[280,539,1466,1490]" italics="true" pageId="5" pageNumber="564">Antrokoreana asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE372FF86FDB21DD0FDB0FA13" box="[545,625,1467,1490]" pageId="5" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (
<emphasis id="B906EAB6E372FF86FD6A1DD1FCEFFA13" bold="true" box="[761,814,1466,1490]" pageId="5" pageNumber="564">E, F:</emphasis>
KUZ Z4004) and paratype ♂♂ (
<emphasis id="B906EAB6E372FF86FBE41DD1FB0CFA13" bold="true" box="[1143,1229,1466,1490]" pageId="5" pageNumber="564">A, C, D:</emphasis>
FCMNH-Ar213;
<emphasis id="B906EAB6E372FF86FA101DD1FA5CFA13" bold="true" box="[1411,1437,1466,1490]" pageId="5" pageNumber="564">B:</emphasis>
FCMNH-Ar243).
<emphasis id="B906EAB6E372FF86FEC31DB5FEA8FA37" bold="true" box="[336,361,1502,1526]" pageId="5" pageNumber="564">A.</emphasis>
Body rings 5 and 6, lateral view.
<emphasis id="B906EAB6E372FF86FD511DB5FD18FA37" bold="true" box="[706,729,1502,1526]" pageId="5" pageNumber="564">B.</emphasis>
First leg pair, posterior view.
<emphasis id="B906EAB6E372FF86FB9E1DB5FBE4FA37" bold="true" box="[1037,1061,1502,1526]" pageId="5" pageNumber="564">C.</emphasis>
Third leg pair, posterior view; white arrowheads indicate adhesive pads.
<emphasis id="B906EAB6E372FF86FD981E69FDE2F9DB" bold="true" box="[523,547,1538,1562]" pageId="5" pageNumber="564">D.</emphasis>
Penes, posterior view.
<emphasis id="B906EAB6E372FF86FC851E69FCECF9DB" bold="true" box="[790,813,1538,1562]" pageId="5" pageNumber="564">E.</emphasis>
Caudal end with 4 podous ring + 1 apodous rings + telson, lateral view.
<emphasis id="B906EAB6E372FF86FE8A1E4DFEEDF9FF" bold="true" box="[281,300,1574,1598]" pageId="5" pageNumber="564">F.</emphasis>
Telson, ventral view. Scale bars: 500 μm (
<emphasis id="B906EAB6E372FF86FD4F1E4DFCE9F9FF" bold="true" box="[732,808,1574,1598]" pageId="5" pageNumber="564">A, E, F</emphasis>
), 200 μm (
<emphasis id="B906EAB6E372FF86FC091E4DFC08F9FF" bold="true" box="[922,969,1574,1598]" pageId="5" pageNumber="564">B, C</emphasis>
), 50 μm (
<emphasis id="B906EAB6E372FF86FBBC1E4DFB80F9FF" bold="true" box="[1071,1089,1574,1598]" pageId="5" pageNumber="564">D</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BCD36A4E372FF86FF541E07FD38F843" blockId="5.[151,1437,1644,2031]" pageId="5" pageNumber="564">
<emphasis id="B906EAB6E372FF86FF541E07FEFBF947" box="[199,314,1644,1670]" italics="true" pageId="5" pageNumber="564">Gonopods</emphasis>
(
<figureCitation id="13492A21E372FF86FED91E07FE54F947" box="[330,405,1644,1670]" captionStart="FIGURE 4" captionStartId="6.[151,250,964,988]" captionTargetBox="[157,1427,186,936]" captionTargetId="figure-19@6.[151,1436,181,941]" captionTargetPageId="6" captionText="FIGURE 4. Antrokoreana asuwaensis sp. nov., holotype ♂ (KUZ Z4004). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. Scale bars: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.6333876" httpUri="https://zenodo.org/record/6333876/files/figure.png" pageId="5" pageNumber="564">Figs 4</figureCitation>
,
<figureCitation id="13492A21E372FF86FE311E07FE27F946" box="[418,486,1644,1671]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">5AF</figureCitation>
). Both anterior and posterior gonopods projecting ventrad. Anterior gonopods (
<figureCitation id="13492A21E372FF86FAF91E07FF66F96B" captionStart="FIGURE 4" captionStartId="6.[151,250,964,988]" captionTargetBox="[157,1427,186,936]" captionTargetId="figure-19@6.[151,1436,181,941]" captionTargetPageId="6" captionText="FIGURE 4. Antrokoreana asuwaensis sp. nov., holotype ♂ (KUZ Z4004). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. Scale bars: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.6333876" httpUri="https://zenodo.org/record/6333876/files/figure.png" pageId="5" pageNumber="564">Figs 4</figureCitation>
,
<figureCitation id="13492A21E372FF86FF261EFBFF3EF96A" box="[181,255,1680,1707]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">5AD</figureCitation>
): each coxal process (
<emphasis id="B906EAB6E372FF86FE6E1EFBFDD8F96B" bold="true" box="[509,537,1680,1706]" pageId="5" pageNumber="564">cp</emphasis>
) expanded distally, distal part concave (
<figureCitation id="13492A21E372FF86FC7B1EFBFB80F96A" box="[1000,1089,1680,1707]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">Fig. 5C</figureCitation>
), anterior part with 6 setae on lateral surface (
<figureCitation id="13492A21E372FF86FED81EDFFE68F90F" box="[331,425,1716,1742]" captionStart="FIGURE 4" captionStartId="6.[151,250,964,988]" captionTargetBox="[157,1427,186,936]" captionTargetId="figure-19@6.[151,1436,181,941]" captionTargetPageId="6" captionText="FIGURE 4. Antrokoreana asuwaensis sp. nov., holotype ♂ (KUZ Z4004). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. Scale bars: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.6333876" httpUri="https://zenodo.org/record/6333876/files/figure.png" pageId="5" pageNumber="564">Fig. 4A</figureCitation>
), posterior part with several setae, curved and directed mesad (
<figureCitation id="13492A21E372FF86FB191EDFFB27F90F" box="[1162,1254,1716,1742]" captionStart="FIGURE 4" captionStartId="6.[151,250,964,988]" captionTargetBox="[157,1427,186,936]" captionTargetId="figure-19@6.[151,1436,181,941]" captionTargetPageId="6" captionText="FIGURE 4. Antrokoreana asuwaensis sp. nov., holotype ♂ (KUZ Z4004). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. Scale bars: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.6333876" httpUri="https://zenodo.org/record/6333876/files/figure.png" pageId="5" pageNumber="564">Fig. 4B</figureCitation>
); telopodite (
<emphasis id="B906EAB6E372FF86FA111EDFFA52F90F" bold="true" box="[1410,1427,1716,1742]" pageId="5" pageNumber="564">tl</emphasis>
) slender, club-shaped, composed of single article, with several long setae on distal margin, field of spine-like setae absent; each flagellum (
<emphasis id="B906EAB6E372FF86FE3B1E97FE78F8D7" bold="true" box="[424,441,1788,1814]" pageId="5" pageNumber="564">fl</emphasis>
) with minute spinules (
<emphasis id="B906EAB6E372FF86FD5A1E97FD2FF8D7" bold="true" box="[713,750,1788,1814]" pageId="5" pageNumber="564">sps</emphasis>
) distally (
<figureCitation id="13492A21E372FF86FCF01E97FC7FF8D6" box="[867,958,1788,1815]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">Fig. 5D</figureCitation>
). Posterior gonopods (
<figureCitation id="13492A21E372FF86FB531E97FAF4F8D6" box="[1216,1333,1788,1815]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">Fig. 5E, F</figureCitation>
) slender, straight, with 1 longitudinal furrow, field of spine-like setae absent; anterior surface covered with minute scalelike processes (
<emphasis id="B906EAB6E372FF86FED51F2FFEB2F89F" bold="true" box="[326,371,1860,1886]" pageId="5" pageNumber="564">slps</emphasis>
); posterior surface smooth, with 3 or 4 long bristles (
<emphasis id="B906EAB6E372FF86FC471F2FFC37F89F" bold="true" box="[980,1014,1860,1886]" pageId="5" pageNumber="564">lbs</emphasis>
) on distal part reaching apex of each posterior gonopod; apex forming 3 flattened branches.
</paragraph>
<paragraph id="8BCD36A4E372FF86FF541FE7FCBAF82E" blockId="5.[151,1437,1644,2031]" pageId="5" pageNumber="564">
<emphasis id="B906EAB6E372FF86FF541FE7FED2F867" box="[199,275,1932,1958]" italics="true" pageId="5" pageNumber="564">Vulvae</emphasis>
(
<figureCitation id="13492A21E372FF86FEB21FE7FE52F866" box="[289,403,1932,1959]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">Fig. 5GI</figureCitation>
). Each vulva pyriform, located inside body ring 3. Operculum (
<emphasis id="B906EAB6E372FF86FBC71FE7FBB3F867" bold="true" box="[1108,1138,1932,1958]" pageId="5" pageNumber="564">op</emphasis>
) slightly longer than bursa (
<emphasis id="B906EAB6E372FF86FF321FDBFF01F80B" bold="true" box="[161,192,1968,1994]" pageId="5" pageNumber="564">bu</emphasis>
), anterior surface of distal part with 4 setae located in 2 longitudinal rows (
<figureCitation id="13492A21E372FF86FB9A1FDBFB44F80A" box="[1033,1157,1968,1995]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">Fig. 5G, H</figureCitation>
). Each valve with pair of setae. Receptaculum seminis (
<emphasis id="B906EAB6E372FF86FE791FBFFDC0F82F" bold="true" box="[490,513,2004,2030]" pageId="5" pageNumber="564">rs</emphasis>
) single, long, claviform (
<figureCitation id="13492A21E372FF86FC8C1FBFFCACF82E" box="[799,877,2004,2031]" captionStart="FIGURE 5" captionStartId="7.[151,250,1380,1404]" captionTargetBox="[151,1436,273,1356]" captionTargetId="figure-60@7.[151,1436,273,1356]" captionTargetPageId="7" captionText="FIGURE 5. Antrokoreana asuwaensis sp. nov., holotype ♂ (A, B: KUZ Z4004), paratype ♂♂ (C: FCMNH-Ar243; D: FCMNH- Ar213; E, F: FCMNH-Ar132) and paratype ♀♀ (G, H: FCMNH-Ar243; I: FCMNH-Ar250). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, right lateral view. D. Tip of right flagellum of anterior gonopods. E. Right posterior gonopod, lateral view. F. Apical part of right posterior gonopod, lateral view. G. Right vulva, posterior view. H. Left vulva, mesal view. I. Left vulva, lateral view; broken line outlines the receptaculum seminis. Scale bars: 250 μm (AC), 25 μm (D), 100 μm (E, G, H), 50 μm (F), 200 μm (I)." figureDoi="http://doi.org/10.5281/zenodo.6333878" httpUri="https://zenodo.org/record/6333878/files/figure.png" pageId="5" pageNumber="564">Fig. 5I</figureCitation>
).
</paragraph>
<caption id="DF0D662CE371FF85FF041BAFFDACFBC0" ID-DOI="http://doi.org/10.5281/zenodo.6333876" ID-Zenodo-Dep="6333876" httpUri="https://zenodo.org/record/6333876/files/figure.png" pageId="6" pageNumber="565" startId="6.[151,250,964,988]" targetBox="[157,1427,186,936]" targetPageId="6">
<paragraph id="8BCD36A4E371FF85FF041BAFFDACFBC0" blockId="6.[151,1436,964,1025]" pageId="6" pageNumber="565">
<emphasis id="B906EAB6E371FF85FF041BAFFED4FC1C" bold="true" box="[151,277,964,989]" pageId="6" pageNumber="565">FIGURE 4.</emphasis>
<taxonomicName id="4C724D27E371FF85FE8E1BAEFDE2FC1D" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[285,547,965,989]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E371FF85FE8E1BAEFDE2FC1D" box="[285,547,965,989]" italics="true" pageId="6" pageNumber="565">Antrokoreana asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE371FF85FDBF1BAEFDBEFC1D" box="[556,639,965,988]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (KUZ Z4004).
<emphasis id="B906EAB6E371FF85FC371BAFFC7CFC1D" bold="true" box="[932,957,964,988]" pageId="6" pageNumber="565">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B906EAB6E371FF85FABB1BAEFAFEFC1C" bold="true" box="[1320,1343,965,989]" pageId="6" pageNumber="565">B.</emphasis>
Anterior gonopods, posterior view. Scale bars: 250 μm.
</paragraph>
</caption>
<paragraph id="8BCD36A4E371FF85FF541C44FD57FBAC" blockId="6.[151,1437,1071,2033]" pageId="6" pageNumber="565">
<emphasis id="B906EAB6E371FF85FF541C44FE8BFB88" box="[199,330,1071,1097]" italics="true" pageId="6" pageNumber="565">Coloration.</emphasis>
In life, yellowish brown with the defense glands visible by transparency as reddish spots (
<figureCitation id="13492A21E371FF85FAA41C44FA4FFB88" box="[1335,1422,1071,1097]" captionStart="FIGURE 1" captionStartId="3.[151,250,1720,1744]" captionTargetBox="[151,1436,1090,1696]" captionTargetId="figure-316@3.[151,1436,1090,1696]" captionTargetPageId="3" captionText="FIGURE 1. Antrokoreana asuwaensis sp. nov., in situ habitus, and its habitat at the type locality. A. Habitus of live male (not collected), dorsal view. B. Habitus of live male (not collected), lateral view. C. Collection site at the Nanatsuoguchi Mine in central Japan. Photographs were taken by Shinya Umemura (A, B), Taro Jonishi (C)." figureDoi="http://doi.org/10.5281/zenodo.6333870" httpUri="https://zenodo.org/record/6333870/files/figure.png" pageId="6" pageNumber="565">Fig. 1B</figureCitation>
). In ethanol, yellowish white or reddish brown.
</paragraph>
</subSubSection>
<subSubSection id="C368652FE371FF85FF541C1CFB1CFB74" pageId="6" pageNumber="565" type="distribution">
<paragraph id="8BCD36A4E371FF85FF541C1CFB1CFB74" blockId="6.[151,1437,1071,2033]" pageId="6" pageNumber="565">
<emphasis id="B906EAB6E371FF85FF541C1CFD8AFB50" bold="true" box="[199,587,1143,1169]" pageId="6" pageNumber="565">Distribution and natural history.</emphasis>
Known only from the
<typeStatus id="54C98806E371FF85FCD41C1CFCB6FB50" box="[839,887,1143,1169]" pageId="6" pageNumber="565">type</typeStatus>
locality in
<collectingCountry id="F3657634E371FF85FC601C1CFBF7FB50" box="[1011,1078,1143,1169]" name="Japan" pageId="6" pageNumber="565">Japan</collectingCountry>
. Observed at the Nanatsuoguchi Mine (
<figureCitation id="13492A21E371FF85FF711CF0FEF7FB74" box="[226,310,1179,1205]" captionStart="FIGURE 1" captionStartId="3.[151,250,1720,1744]" captionTargetBox="[151,1436,1090,1696]" captionTargetId="figure-316@3.[151,1436,1090,1696]" captionTargetPageId="3" captionText="FIGURE 1. Antrokoreana asuwaensis sp. nov., in situ habitus, and its habitat at the type locality. A. Habitus of live male (not collected), dorsal view. B. Habitus of live male (not collected), lateral view. C. Collection site at the Nanatsuoguchi Mine in central Japan. Photographs were taken by Shinya Umemura (A, B), Taro Jonishi (C)." figureDoi="http://doi.org/10.5281/zenodo.6333870" httpUri="https://zenodo.org/record/6333870/files/figure.png" pageId="6" pageNumber="565">Fig. 1C</figureCitation>
) from late autumn (November) to early spring (March) (see
<bibRefCitation id="EFE34B55E371FF85FC4D1CF0FB10FB74" author="Umemura, S. &amp; Yoshizawa, Y. &amp; Ito, K." box="[990,1233,1179,1205]" pageId="6" pageNumber="565" pagination="63 - 74" refId="ref8607" refString="Umemura, S., Yoshizawa, Y. &amp; Ito, K. (2019) Arthropod fauna of Nanatsuoguchi mine in Mt. Asuwa, Fukui City, Fukui Prefecture. Bulletin of the Fukui City Museum of Natural History, 66, 63 - 74." type="journal article" year="2019">
Umemura
<emphasis id="B906EAB6E371FF85FBC61CF7FB4CFB74" box="[1109,1165,1179,1205]" italics="true" pageId="6" pageNumber="565">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C368652FE371FF85FF541CD4FC84FA87" pageId="6" pageNumber="565" type="etymology">
<paragraph id="8BCD36A4E371FF85FF541CD4FB52FB18" blockId="6.[151,1437,1071,2033]" box="[199,1171,1215,1241]" pageId="6" pageNumber="565">
<emphasis id="B906EAB6E371FF85FF541CD4FE65FB18" bold="true" box="[199,420,1215,1241]" pageId="6" pageNumber="565">Derivatio nominis.</emphasis>
The specific name is derived from the
<typeStatus id="54C98806E371FF85FCC81CD4FC4AFB18" box="[859,907,1215,1241]" pageId="6" pageNumber="565">type</typeStatus>
locality of this species.
</paragraph>
<paragraph id="8BCD36A4E371FF85FF541C88FC84FA87" blockId="6.[151,1437,1071,2033]" pageId="6" pageNumber="565">
<emphasis id="B906EAB6E371FF85FF541C88FE45FB3C" bold="true" box="[199,388,1251,1277]" pageId="6" pageNumber="565">DNA sequences.</emphasis>
Five sequences were obtained: two of a male (FCMNH-Ar251)—28S (INSD accession number,
<accessionNumber id="9421AB47E371FF85FF041D6CFED2FAE3" box="[151,275,1287,1314]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC654434" pageId="6" pageNumber="565">LC654434</accessionNumber>
; 517 bp) and 16S (
<accessionNumber id="9421AB47E371FF85FE741D6CFDA5FAE3" box="[487,612,1287,1314]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC654436" pageId="6" pageNumber="565">LC654436</accessionNumber>
; 425 bp); and three of a female (FCMNH-Ar252)—28S (
<accessionNumber id="9421AB47E371FF85FB791D6CFAA6FAE3" box="[1258,1383,1287,1314]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC654435" pageId="6" pageNumber="565">LC654435</accessionNumber>
; 517 bp), COI (
<accessionNumber id="9421AB47E371FF85FE9A1D40FE46FA87" box="[265,391,1323,1350]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC654438" pageId="6" pageNumber="565">LC654438</accessionNumber>
; 658 bp), and 16S (
<accessionNumber id="9421AB47E371FF85FDF31D40FD1CFA87" box="[608,733,1323,1350]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC654437" pageId="6" pageNumber="565">LC654437</accessionNumber>
; 425 bp).
</paragraph>
</subSubSection>
<subSubSection id="C368652FE371FF84FF541D24FABAFF3B" lastPageId="7" lastPageNumber="566" pageId="6" pageNumber="565" type="discussion">
<paragraph id="8BCD36A4E371FF85FF541D24FAC9F9DC" blockId="6.[151,1437,1071,2033]" pageId="6" pageNumber="565">
<emphasis id="B906EAB6E371FF85FF541D24FEFAFAA8" bold="true" box="[199,315,1359,1385]" pageId="6" pageNumber="565">Remarks.</emphasis>
The new species is clearly judged to be a member of the genus
<taxonomicName id="4C724D27E371FF85FBBB1D24FB00FAA8" authorityName="Verhoeff" authorityYear="1938" box="[1064,1217,1359,1385]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E371FF85FBBB1D24FB00FAA8" box="[1064,1217,1359,1385]" italics="true" pageId="6" pageNumber="565">Antrokoreana</emphasis>
</taxonomicName>
as it possesses the following diagnostic characteristics of the genus (
<bibRefCitation id="EFE34B55E371FF85FD2E1D18FC9AFA4F" author="Enghoff, H." box="[701,859,1395,1422]" pageId="6" pageNumber="565" pagination="27 - 67" refId="ref6947" refString="Enghoff, H. (1985) The millipede family Nemasomatidae. With the description of a new genus, and a revision of Orinisobates (Diplopoda: Julida). Entomologica Scandinavica, 16, 27 - 67. https: // doi. org / 10.1163 / 187631285 X 00045" type="journal article" year="1985">Enghoff 1985</bibRefCitation>
;
<bibRefCitation id="EFE34B55E371FF85FCF51D18FB96FA4F" author="Shinohara, K. &amp; Tanabe, T. &amp; Korsos, Z." box="[870,1111,1395,1422]" pageId="6" pageNumber="565" pagination="941 - 984" refId="ref8228" refString="Shinohara, K., Tanabe, T. &amp; Korsos, Z. (2015) Arthropoda. Myriapoda. Diplopoda. In: Aoki, J. - I. (Ed.), Pictorial Keys to Soil Animals of Japan. 2 nd Edition. Tokai University Press, Hadano, pp. 941 - 984." type="book chapter" year="2015">
Shinohara
<emphasis id="B906EAB6E371FF85FC4E1D1FFBD4FA4C" box="[989,1045,1395,1421]" italics="true" pageId="6" pageNumber="565">et al.</emphasis>
2015
</bibRefCitation>
): 1) head without ommatidia; 2) antennomere VII almost as long as antennomere VI, and its whole surface covered with short sensilla chaetica among longer sensilla trichodea; 3) posterior margin of each ring with one row of setae; 4) tibia of leg-pair 1 with 1 long spine; 5) femur, post-femur, and tibia of other leg pairs with adhesive pads; 6) coxa of leg-pair 2 with pair of penes; 7) both gonopods without fields of spine-like setae; and 8) anterior gonopods with telopodites.
</paragraph>
<paragraph id="8BCD36A4E371FF85FF541E4CFBE4F8FC" blockId="6.[151,1437,1071,2033]" pageId="6" pageNumber="565">
<taxonomicName id="4C724D27E371FF85FF541E4CFE28F980" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[199,489,1575,1601]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E371FF85FF541E4CFE28F980" box="[199,489,1575,1601]" italics="true" pageId="6" pageNumber="565">Antrokoreana asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE371FF85FE611E4CFD8BF980" box="[498,586,1575,1601]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
is distinguishable from all seven congeners, including
<taxonomicName id="4C724D27E371FF85FB531E4CFA8AF980" authorityName="Masuda" authorityYear="2010" box="[1216,1355,1575,1601]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="gujoensis">
<emphasis id="B906EAB6E371FF85FB531E4CFA8AF980" box="[1216,1355,1575,1601]" italics="true" pageId="6" pageNumber="565">A. gujoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C724D27E371FF85FA171E4CFF14F9A4" authorityName="Takashima &amp; Haga" authorityYear="1956" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B906EAB6E371FF85FA171E4CFF14F9A4" italics="true" pageId="6" pageNumber="565">A. uenoi</emphasis>
</taxonomicName>
which are endemic to central Honshu, by the distally expanded coxal processes of the anterior gonopods, in accordance with the descriptions of the six species known thus far (
<bibRefCitation id="EFE34B55E371FF85FC251E04FB17F94B" author="Takashima, H. &amp; Haga, A." box="[950,1238,1647,1674]" pageId="6" pageNumber="565" pagination="329 - 343" refId="ref8346" refString="Takashima, H. &amp; Haga, A. (1956) A contribution towards the Japanese cave-dwelling species of the class Diplopoda. Journal of the Yamashina Institute for Ornithology, 1, 329 - 343. https: // doi. org / 10.3312 / jyio 1952.1.329" type="journal article" year="1956">Takashima &amp; Haga 1956</bibRefCitation>
;
<bibRefCitation id="EFE34B55E371FF85FB701E04FA59F948" author="Shinohara, K." box="[1251,1432,1647,1673]" pageId="6" pageNumber="565" pagination="23 - 30" refId="ref8156" refString="Shinohara, K. (1960) Three new species of Juloidea (Diplopoda) from Chichibu. Bulletin of the Chichibu Museum of Natural History, 10, 23 - 30." type="journal article" year="1960">Shinohara 1960</bibRefCitation>
,
<bibRefCitation id="EFE34B55E371FF85FF041EF8FF13F96C" author="Shinohara, K." box="[151,210,1683,1709]" pageId="6" pageNumber="565" pagination="217 - 251" refId="ref8189" refString="Shinohara, K. (1973) The fauna of the lava caves around Mt. Fuji-san. XIII. Diplopoda and Chilopoda. Bulletin of the National Science Museum, Tokyo, 16, 217 - 251." type="journal article" year="1973">1973</bibRefCitation>
;
<bibRefCitation id="EFE34B55E371FF85FF4E1EF8FE26F96C" author="Murakami, Y. &amp; Paik, K. Y." box="[221,487,1683,1709]" pageId="6" pageNumber="565" pagination="363 - 384" refId="ref7814" refString="Murakami, Y. &amp; Paik, K. Y. (1968) Results of the Speleological Survey in South Korea 1966. XI. Cave-dwelling myriapods from the southern part of Korea. Bulletin of the National Science Museum (Tokyo), 11, 363 - 384." type="journal article" year="1968">Murakami &amp; Paik 1968</bibRefCitation>
;
<bibRefCitation id="EFE34B55E371FF85FE611EF8FD69F96C" author="Murakami, Y." box="[498,680,1683,1709]" pageId="6" pageNumber="565" pagination="557 - 582" refId="ref7746" refString="Murakami, Y. (1969) Myriapods found in limestone caves of northern Honshu, Japan. Bulletin of the National Science Museum, Tokyo, 12, 557 - 582." type="journal article" year="1969">Murakami 1969</bibRefCitation>
;
<bibRefCitation id="EFE34B55E371FF85FD201EF8FC8EF96C" author="Masuda, K." box="[691,847,1683,1709]" pageId="6" pageNumber="565" pagination="29 - 32" refId="ref7588" refString="Masuda, K. (2010) A new species of the genus Antrokoreana (Diplopoda) from Japanese caves. Journal of the Speleological Society of Japan, 35, 29 - 32." type="journal article" year="2010">Masuda 2010</bibRefCitation>
). In common with
<taxonomicName id="4C724D27E371FF85FB8D1EF8FB66F96C" authorityName="Masuda" authorityYear="2010" box="[1054,1191,1683,1709]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="gujoensis">
<emphasis id="B906EAB6E371FF85FB8D1EF8FB66F96C" box="[1054,1191,1683,1709]" italics="true" pageId="6" pageNumber="565">A. gujoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C724D27E371FF85FB481EF8FAACF96C" baseAuthorityName="Verhoeff" baseAuthorityYear="1941" box="[1243,1389,1683,1709]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="takakuwai">
<emphasis id="B906EAB6E371FF85FB481EF8FAACF96C" box="[1243,1389,1683,1709]" italics="true" pageId="6" pageNumber="565">A. takakuwai</emphasis>
</taxonomicName>
, the posterior gonopods of
<taxonomicName id="4C724D27E371FF85FE001EDCFDF0F910" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[403,561,1719,1745]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E371FF85FE001EDCFDF0F910" box="[403,561,1719,1745]" italics="true" pageId="6" pageNumber="565">A. asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE371FF85FDA41EDCFD4CF910" box="[567,653,1719,1745]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
each bears 3 or 4 long bristles reaching the distal edge of the gonopod (
<bibRefCitation id="EFE34B55E371FF85FF0C1EB0FE97F934" author="Shinohara, K." box="[159,342,1755,1781]" pageId="6" pageNumber="565" pagination="23 - 30" refId="ref8156" refString="Shinohara, K. (1960) Three new species of Juloidea (Diplopoda) from Chichibu. Bulletin of the Chichibu Museum of Natural History, 10, 23 - 30." type="journal article" year="1960">Shinohara 1960</bibRefCitation>
,
<bibRefCitation id="EFE34B55E371FF85FEF71EB0FE5EF934" author="Shinohara, K." box="[356,415,1755,1781]" pageId="6" pageNumber="565" pagination="217 - 251" refId="ref8189" refString="Shinohara, K. (1973) The fauna of the lava caves around Mt. Fuji-san. XIII. Diplopoda and Chilopoda. Bulletin of the National Science Museum, Tokyo, 16, 217 - 251." type="journal article" year="1973">1973</bibRefCitation>
;
<bibRefCitation id="EFE34B55E371FF85FE3E1EB0FD8DF934" author="Masuda, K." box="[429,588,1755,1781]" pageId="6" pageNumber="565" pagination="29 - 32" refId="ref7588" refString="Masuda, K. (2010) A new species of the genus Antrokoreana (Diplopoda) from Japanese caves. Journal of the Speleological Society of Japan, 35, 29 - 32." type="journal article" year="2010">Masuda 2010</bibRefCitation>
); this shared feature suggests these species may be closely related. Future studies are needed to elucidate phylogenetic relationships among
<taxonomicName id="4C724D27E371FF85FC011E94FB4BF8D8" box="[914,1162,1791,1817]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B906EAB6E371FF85FC011E94FBEAF8D8" box="[914,1067,1791,1817]" italics="true" pageId="6" pageNumber="565">Antrokoreana</emphasis>
species
</taxonomicName>
, and clarify whether
<taxonomicName id="4C724D27E371FF85FA171E94FED6F8FC" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E371FF85FA171E94FED6F8FC" italics="true" pageId="6" pageNumber="565">A. asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE371FF85FE8D1F48FEB6F8FC" box="[286,375,1827,1853]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C724D27E371FF85FE111F48FDCAF8FC" authorityName="Masuda" authorityYear="2010" box="[386,523,1827,1853]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="gujoensis">
<emphasis id="B906EAB6E371FF85FE111F48FDCAF8FC" box="[386,523,1827,1853]" italics="true" pageId="6" pageNumber="565">A. gujoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C724D27E371FF85FDD21F48FD12F8FC" baseAuthorityName="Verhoeff" baseAuthorityYear="1941" box="[577,723,1827,1853]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="takakuwai">
<emphasis id="B906EAB6E371FF85FDD21F48FD12F8FC" box="[577,723,1827,1853]" italics="true" pageId="6" pageNumber="565">A. takakuwai</emphasis>
</taxonomicName>
form a monophyletic lineage.
</paragraph>
<paragraph id="8BCD36A4E371FF84FF541F2CFBE1FF14" blockId="6.[151,1437,1071,2033]" lastBlockId="7.[151,1436,151,250]" lastPageId="7" lastPageNumber="566" pageId="6" pageNumber="565">
<emphasis id="B906EAB6E371FF85FF541F2CFE75F8A0" bold="true" box="[199,436,1863,1889]" pageId="6" pageNumber="565">Molecular analyses.</emphasis>
The topology of the BI tree (
<figureCitation id="13492A21E371FF85FD6D1F2CFC82F8A3" box="[766,835,1863,1890]" captionStart="FIGURE 6" captionStartId="8.[151,250,1156,1180]" captionTargetBox="[155,1432,413,1124]" captionTargetId="figure-451@8.[151,1436,403,1132]" captionTargetPageId="8" captionText="FIGURE 6. Bayesian inference tree for the the combineid 965 bp of nuclear 28S rRNA and mitochondrial 16S rRNA marker sequences. Numbers on nodes are the bootstrap values for maximum likelihood ≥ 60% and Bayesian posterior probabilities ≥ 0.95." figureDoi="http://doi.org/10.5281/zenodo.6333880" httpUri="https://zenodo.org/record/6333880/files/figure.png" pageId="6" pageNumber="565">Fig. 6</figureCitation>
), which was constructed to estimate the phylogenetic position of
<taxonomicName id="4C724D27E371FF85FE881F00FDFFF844" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[283,574,1899,1925]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E371FF85FE881F00FDFFF844" box="[283,574,1899,1925]" italics="true" pageId="6" pageNumber="565">Antrokoreana asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE371FF85FDD51F00FD60F844" box="[582,673,1899,1925]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
, was identical to that of the ML tree (not shown). The monophyly of
<taxonomicName id="4C724D27E371FF85FF251FE4FE97F868" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[182,342,1935,1961]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E371FF85FF251FE4FE97F868" box="[182,342,1935,1961]" italics="true" pageId="6" pageNumber="565">A. asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE371FF85FECE1FE4FE75F868" box="[349,436,1935,1961]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C724D27E371FF85FE781FE4FDBAF868" authorityName="Verhoeff" authorityYear="1938" box="[491,635,1935,1961]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B906EAB6E371FF85FE781FE4FDBAF868" box="[491,635,1935,1961]" italics="true" pageId="6" pageNumber="565">A. gracilipes</emphasis>
</taxonomicName>
was fully supported (BS = 96%, PP = 0.99), and the monophyly of the nemasomatid
<taxonomicName id="4C724D27E371FF85FEA41FDFFDBEF80C" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[311,639,1971,1997]" class="Diplopoda" family="Nemasomatidae" genus="Nemasoma" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E371FF85FEA41FDFFE72F80C" box="[311,435,1972,1997]" italics="true" pageId="6" pageNumber="565">Nemasoma</emphasis>
C.L.
<bibRefCitation id="EFE34B55E371FF85FE6A1FD8FDBEF80C" author="Koch, C. L." box="[505,639,1971,1997]" pageId="6" pageNumber="565" refId="ref7461" refString="Koch, C. L. (1847) System der Myriapoden, mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden, und Arachniden. Heft 1 - 40. Friedrich Pustet, Regensburg, 196 pp. https: // doi. org / 10.5962 / bhl. title. 49866" type="book" year="1847">Koch, 1847</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C724D27E371FF85FD281FD8FBE2F80C" authority="Verhoeff, 1908" authorityName="Verhoeff" authorityYear="1908" box="[699,1059,1971,1997]" class="Diplopoda" family="Nemasomatidae" genus="Thalassisobates" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E371FF85FD281FD8FCAFF80C" box="[699,878,1971,1997]" italics="true" pageId="6" pageNumber="565">Thalassisobates</emphasis>
<bibRefCitation id="EFE34B55E371FF85FCEB1FD8FBE2F80C" author="Verhoeff, K. W." box="[888,1059,1971,1997]" pageId="6" pageNumber="565" pagination="486 - 495" refId="ref8656" refString="Verhoeff, K. W. (1908) Uber Diplopoden. 8. (28.) Aufsatz: Ein neuer Strand-Iulide und seine biologisch-morphologische Bedeutung. Zoologischer Anzeiger, 32, 486 - 495." type="journal article" year="1908">Verhoeff, 1908</bibRefCitation>
</taxonomicName>
was also recovered (BS = 94%, PP = 1.0). The monophyly of the nemasomatid genera
<taxonomicName id="4C724D27E371FF85FC941FBCFC61F830" authorityName="Verhoeff" authorityYear="1938" box="[775,928,2007,2033]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E371FF85FC941FBCFC61F830" box="[775,928,2007,2033]" italics="true" pageId="6" pageNumber="565">Antrokoreana</emphasis>
</taxonomicName>
and (
<taxonomicName id="4C724D27E371FF85FC701FB3FB9EF830" authorityName="C. L. Koch" authorityYear="1847" box="[995,1119,2008,2033]" class="Diplopoda" family="Nemasomatidae" genus="Nemasoma" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E371FF85FC701FB3FB9EF830" box="[995,1119,2008,2033]" italics="true" pageId="6" pageNumber="565">Nemasoma</emphasis>
</taxonomicName>
+
<taxonomicName id="4C724D27E371FF85FB131FBCFAF2F830" authorityName="Verhoeff" authorityYear="1908" box="[1152,1331,2007,2033]" class="Diplopoda" family="Nemasomatidae" genus="Thalassisobates" kingdom="Animalia" order="Julida" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E371FF85FB131FBCFAF2F830" box="[1152,1331,2007,2033]" italics="true" pageId="6" pageNumber="565">Thalassisobates</emphasis>
</taxonomicName>
) was not elucidated, but nonetheless, the obtained phylogenies failed to resolve the basal nodes and phylogenetic relationships among
<taxonomicName id="4C724D27E370FF84FF7918D0FE42FF14" authorityName="Verhoeff" authorityYear="1938" box="[234,387,187,213]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="7" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E370FF84FF7918D0FE42FF14" box="[234,387,187,213]" italics="true" pageId="7" pageNumber="566">Antrokoreana</emphasis>
</taxonomicName>
, (
<taxonomicName id="4C724D27E370FF84FE0818D0FDD6FF15" authorityName="C. L. Koch" authorityYear="1847" box="[411,535,187,212]" class="Diplopoda" family="Nemasomatidae" genus="Nemasoma" kingdom="Animalia" order="Julida" pageId="7" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E370FF84FE0818D0FDD6FF15" box="[411,535,187,212]" italics="true" pageId="7" pageNumber="566">Nemasoma</emphasis>
</taxonomicName>
+
<taxonomicName id="4C724D27E370FF84FDA618D0FD29FF14" authorityName="Verhoeff" authorityYear="1908" box="[565,744,187,213]" class="Diplopoda" family="Nemasomatidae" genus="Thalassisobates" kingdom="Animalia" order="Julida" pageId="7" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="B906EAB6E370FF84FDA618D0FD29FF14" box="[565,744,187,213]" italics="true" pageId="7" pageNumber="566">Thalassisobates</emphasis>
</taxonomicName>
) and parajuloidean species.
</paragraph>
<paragraph id="8BCD36A4E370FF84FF5418B4FABAFF3B" blockId="7.[151,1436,151,250]" box="[199,1403,222,250]" pageId="7" pageNumber="566">
The COI uncorrected
<emphasis id="B906EAB6E370FF84FE2F18B4FE0BFF39" box="[444,458,223,248]" italics="true" pageId="7" pageNumber="566">p</emphasis>
-distance between
<taxonomicName id="4C724D27E370FF84FD0918B4FCFBFF39" authority="Kato &amp; Umemura &amp; Nakano, 2022" authorityName="Kato &amp; Umemura &amp; Nakano" authorityYear="2022" box="[666,826,223,249]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="asuwaensis" status="sp. nov.">
<emphasis id="B906EAB6E370FF84FD0918B4FCFBFF39" box="[666,826,223,249]" italics="true" pageId="7" pageNumber="566">A. asuwaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557CDE370FF84FCD318B4FC56FF38" box="[832,919,223,249]" pageId="7" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C724D27E370FF84FC5E18B4FB9DFF38" authorityName="Verhoeff" authorityYear="1938" box="[973,1116,223,249]" class="Diplopoda" family="Nemasomatidae" genus="Antrokoreana" kingdom="Animalia" order="Julida" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B906EAB6E370FF84FC5E18B4FB9DFF38" box="[973,1116,223,249]" italics="true" pageId="7" pageNumber="566">A. gracilipes</emphasis>
</taxonomicName>
was 16.9% (111/658 bp).
</paragraph>
</subSubSection>
</treatment>
</document>