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<mods:title id="07B82BD326C56639AF9F4807300CC1C2">Revision and phylogeny of Hypotelus Erichson: a Neotropical genus of minute rove beetles (Coleoptera, Staphylinidae, Piestinae)</mods:title>
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<heading id="D08581F7FF9CD531CEB5FACFD048FAC2" bold="true" box="[151,610,1357,1383]" fontSize="11" level="1" pageId="10" pageNumber="461" reason="1">
<taxonomicName id="4C724D18FF9CD531CEB5FACFD048FAC2" ID-CoL="3NZD3" authority="Bernhauer, 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,610,1357,1383]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B906EA89FF9CD531CEB5FACFD048FAC2" bold="true" box="[151,610,1357,1383]" pageId="10" pageNumber="461">
<emphasis id="B906EA89FF9CD531CEB5FACFD353FAC2" bold="true" box="[151,377,1357,1383]" italics="true" pageId="10" pageNumber="461">Hypotelus andinus</emphasis>
(
<bibRefCitation id="EFE34B6AFF9CD531CFABFACFD070FAC2" author="Bernhauer" box="[393,602,1357,1383]" pageId="10" pageNumber="461" refString="Bernhauer, M. (1917) Neue arten der gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. Neue Beitrage zur Systematischen Insektenkunde, 1, 45 - 53." type="journal article" year="1917">Bernhauer, 1917</bibRefCitation>
)
</emphasis>
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</paragraph>
<paragraph id="8BCD369BFF9CD531CEB5FAF2D311FA2C" blockId="10.[151,610,1357,1417]" box="[151,315,1392,1417]" pageId="10" pageNumber="461">
(
<figureCitation id="13492A1EFF9CD531CEBDFAF2D2D6FA2C" box="[159,252,1392,1417]" captionStart="FIGURES 9 19" captionStartId="9.[151,264,1913,1935]" captionTargetBox="[216,1355,230,1832]" captionTargetId="figure@9.[151,1436,157,1945]" captionTargetPageId="9" captionText="FIGURES 9 19. Habitus, dorsal view. 9, Hypotelus insulanus, syntype, male; 10, H. praecox, lectotype, female; 11, H. scheerpeltzi sp. nov., holotype, male; 12, H. testaceus, syntype, male; 13, H. melanodelta sp. nov., holotype, female; 14, H. castaneus sp. nov., holotype, male; 15, H. brevitarsus sp. nov., holotype, male; 16, H. corniculatus sp. nov., holotype, female; 17, H. marginatus, syntype, male; 18, H. laevis, male; 19, H. andinus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/803652/files/figure.png" pageId="10" pageNumber="461">Figs. 19</figureCitation>
,
<figureCitation id="13492A1EFF9CD531CF2AFAF3D318FA2C" box="[264,306,1393,1417]" captionStart="FIGURE 114" captionStartId="31.[151,250,1701,1723]" captionTargetBox="[190,1375,215,1655]" captionTargetId="figure@31.[150,1437,153,1679]" captionTargetPageId="31" captionText="FIGURE 114. Geographical records of Hypotelus. Hypotelus andinus, Colombia, Ecuador and Bolivia; H. insulanus, Caribbean islands; H. marginatus, Colombia, Venezuela and Guatemala; H. testaceus, Panama; H. brevitarsus sp. nov., Panama. Countries without dots but colored represent the occurrence but no locality given." httpUri="https://zenodo.org/record/803674/files/figure.png" pageId="10" pageNumber="461">114</figureCitation>
)
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<taxonomicName id="4C724D18FF9CD531CEB5FA3BD08BFA6B" ID-CoL="4HRWS" authority="Bernhauer, 1917: 45" authorityName="Bernhauer" authorityPageNumber="45" authorityYear="1917" box="[151,673,1464,1486]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andinus" subGenus="Antropiestus">
<emphasis id="B906EA89FF9CD531CEB5FA3BD2CAFA6B" box="[151,224,1465,1486]" italics="true" pageId="10" pageNumber="461">Piestus</emphasis>
(
<emphasis id="B906EA89FF9CD531CECCFA3BD347FA6B" box="[238,365,1465,1486]" italics="true" pageId="10" pageNumber="461">Antropiestus</emphasis>
)
<emphasis id="B906EA89FF9CD531CF5EFA3AD3E6FA6B" box="[380,460,1464,1486]" italics="true" pageId="10" pageNumber="461">andinus</emphasis>
<treatmentCitation id="0AD3108AFF9CD531CFF1FA3AD08BFA6B" author="Bernhauer" box="[467,673,1464,1486]" page="45" pageId="10" pageNumber="461" year="1917">
<bibRefCitation id="EFE34B6AFF9CD531CFF1FA3AD054FA6B" author="Bernhauer" box="[467,638,1464,1486]" pageId="10" pageNumber="461" refString="Bernhauer, M. (1917) Neue arten der gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. Neue Beitrage zur Systematischen Insektenkunde, 1, 45 - 53." type="journal article" year="1917">Bernhauer, 1917</bibRefCitation>
: 45
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(original description,
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locality: West-Kolumbien: Umgebung von Cali am Rio Cauca);
<treatmentCitation id="0AD3108AFF9CD531CF55FA55D04BFA48" author="Scheerpeltz" box="[375,609,1495,1517]" page="993" pageId="10" pageNumber="461" year="1933">
<bibRefCitation id="EFE34B6AFF9CD531CF55FA55D01AFA48" author="Scheerpeltz" box="[375,560,1495,1517]" pageId="10" pageNumber="461" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 6 (129). Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 993
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(distribution);
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<bibRefCitation id="EFE34B6AFF9CD531CCD9FA55D19FFA48" author="Scheerpeltz" box="[763,949,1495,1517]" pageId="10" pageNumber="461" refString="Scheerpeltz, O. (1952) Revision der Gattung Piestus Gravh. (Coleoptera Staphylinidae). Revista Chilena de Entomologia, 2, 281 - 305." type="journal article" year="1952">Scheerpeltz, 1952</bibRefCitation>
: 295
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(characters, distribution);
<taxonomicName id="4C724D18FF9CD531CAD6FA5AD392F9AE" ID-CoL="4HRWS" authority="Blackwelder, 1944: 100" authorityName="Blackwelder" authorityPageNumber="100" authorityYear="1944" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B906EA89FF9CD531CAD6FA5AD7BFFA48" box="[1268,1429,1495,1517]" italics="true" pageId="10" pageNumber="461">Piestus andinus</emphasis>
:
<treatmentCitation id="0AD3108AFF9CD531CEE5FA77D392F9AE" author="Blackwelder" box="[199,440,1525,1547]" page="100" pageId="10" pageNumber="461" year="1944">
<bibRefCitation id="EFE34B6AFF9CD531CEE5FA77D3A3F9AE" author="Blackwelder" box="[199,393,1525,1547]" pageId="10" pageNumber="461" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. United States National Museum Bulletin, 185, xii + 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 100
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(distribution);
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<bibRefCitation id="EFE34B6AFF9CD531CC70FA74D0CCF9AE" author="Herman" box="[594,742,1525,1547]" pageId="10" pageNumber="461" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millennium, parts I - VII. Bulletin of the American Museum of Natural History, 167, 1 - 4218." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 1788
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(distribution);
<treatmentCitation id="0AD3108AFF9CD531CD99FA74D680F9AE" author="Newton" box="[955,1194,1525,1548]" page="37" pageId="10" pageNumber="461" year="2005">
<bibRefCitation id="EFE34B6AFF9CD531CD99FA74D6ADF9AE" author="Newton" box="[955,1159,1525,1548]" pageId="10" pageNumber="461" refString="Newton, A. F., Chacon, C. G. &amp; Chandler, D. S. (2005) Checklist of the Staphylinidae (Coleoptera) of Colombia. Biota Colombiana, 6, 1 - 72." type="journal article" year="2005">
Newton
<emphasis id="B906EA89FF9CD531CA30FA75D614F9AE" box="[1042,1086,1525,1548]" italics="true" pageId="10" pageNumber="461">et al</emphasis>
., 2005
</bibRefCitation>
: 37
</treatmentCitation>
(distribution).
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<emphasis id="B906EA89FF9CD531CEB5F997D2CAF98F" box="[151,224,1557,1578]" italics="true" pageId="10" pageNumber="461">Piestus</emphasis>
(
<emphasis id="B906EA89FF9CD531CECDF997D344F98F" box="[239,366,1557,1578]" italics="true" pageId="10" pageNumber="461">Antropiestus</emphasis>
)
<emphasis id="B906EA89FF9CD531CF5EF997D3DEF98F" box="[380,500,1557,1578]" italics="true" pageId="10" pageNumber="461">strigipennis</emphasis>
<treatmentCitation id="0AD3108AFF9CD531CFD8F996D0E2F98F" author="Bernhauer" box="[506,712,1556,1578]" page="65" pageId="10" pageNumber="461" year="1921">
<bibRefCitation id="EFE34B6AFF9CD531CFD8F996D08FF98F" author="Bernhauer" box="[506,677,1556,1578]" pageId="10" pageNumber="461" refString="Bernhauer, M. (1921) Zur Staphyliniden fauna von Sudamerika. Deutsche Entomologische Zeitschrift, 1921, 65 - 77." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
: 65
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(original description,
<typeStatus id="54C98839FF9CD531CD88F997D1FCF98F" box="[938,982,1557,1578]" pageId="10" pageNumber="461">type</typeStatus>
locality;
<collectingCountry id="F365760BFF9CD531CA1BF996D6B3F98F" box="[1081,1177,1556,1578]" name="Bolivia" pageId="10" pageNumber="461">Bolivien</collectingCountry>
: Yuracaris);
<treatmentCitation id="0AD3108AFF9CD531CB3DF996D307F9EC" author="Scheerpeltz" page="993" pageId="10" pageNumber="461" year="1933">
<bibRefCitation id="EFE34B6AFF9CD531CB3DF996D2D7F9EC" author="Scheerpeltz" pageId="10" pageNumber="461" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 6 (129). Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 993
</treatmentCitation>
(distribution);
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<bibRefCitation id="EFE34B6AFF9CD531CFE4F9B1D0AAF9EC" author="Scheerpeltz" box="[454,640,1587,1609]" pageId="10" pageNumber="461" refString="Scheerpeltz, O. (1952) Revision der Gattung Piestus Gravh. (Coleoptera Staphylinidae). Revista Chilena de Entomologia, 2, 281 - 305." type="journal article" year="1952">Scheerpeltz, 1952</bibRefCitation>
: 295
</treatmentCitation>
(characters, distribution);
<treatmentCitation id="0AD3108AFF9CD531CD99F9B1D68EF9EC" author="Caron" box="[955,1188,1587,1609]" page="551" pageId="10" pageNumber="461" year="2012">
<bibRefCitation id="EFE34B6AFF9CD531CD99F9B1D65FF9EC" author="Caron" box="[955,1141,1587,1609]" pageId="10" pageNumber="461" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B906EA89FF9CD531CDDDF9B6D606F9EC" box="[1023,1068,1587,1609]" italics="true" pageId="10" pageNumber="461">et al</emphasis>
., 2012
</bibRefCitation>
: 551
</treatmentCitation>
(as junior synonym).
<taxonomicName id="4C724D18FF9CD531CEB5F9D0D077F9C2" authority="Blackwelder, 1944: 101" authorityName="Blackwelder" authorityPageNumber="101" authorityYear="1944" box="[151,605,1617,1639]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="strigipennis">
<emphasis id="B906EA89FF9CD531CEB5F9D0D374F9C2" box="[151,350,1618,1639]" italics="true" pageId="10" pageNumber="461">Piestus strigipennis</emphasis>
:
<treatmentCitation id="0AD3108AFF9CD531CF49F9D3D077F9C2" author="Blackwelder" box="[363,605,1617,1639]" page="101" pageId="10" pageNumber="461" year="1944">
<bibRefCitation id="EFE34B6AFF9CD531CF49F9D3D007F9C2" author="Blackwelder" box="[363,557,1617,1639]" pageId="10" pageNumber="461" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. United States National Museum Bulletin, 185, xii + 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 101
</treatmentCitation>
</taxonomicName>
(distribution);
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<bibRefCitation id="EFE34B6AFF9CD531CCD4F9D0D1A0F9C2" author="Herman" box="[758,906,1617,1639]" pageId="10" pageNumber="461" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millennium, parts I - VII. Bulletin of the American Museum of Natural History, 167, 1 - 4218." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 1795
</treatmentCitation>
(distribution).
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</paragraph>
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<treatmentCitationGroup id="AB6211B5FF9CD531CEB5F9F2D650F923" box="[151,1146,1648,1670]" pageId="10" pageNumber="461">
<taxonomicName id="4C724D18FF9CD531CEB5F9F2D061F923" ID-CoL="3NZD3" authority="Caron et al., 2012: 551" authorityName="Caron et al." authorityPageNumber="551" authorityYear="2012" box="[151,587,1648,1670]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B906EA89FF9CD531CEB5F9F2D37EF923" box="[151,340,1648,1670]" italics="true" pageId="10" pageNumber="461">Hypotelus andinus</emphasis>
:
<treatmentCitation id="0AD3108AFF9CD531CF43F9F2D061F923" author="Caron" box="[353,587,1648,1670]" page="551" pageId="10" pageNumber="461" year="2012">
<bibRefCitation id="EFE34B6AFF9CD531CF43F9F2D036F923" author="Caron" box="[353,540,1648,1670]" pageId="10" pageNumber="461" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B906EA89FF9CD531CF87F9F3D3F8F923" box="[421,466,1648,1670]" italics="true" pageId="10" pageNumber="461">et al</emphasis>
., 2012
</bibRefCitation>
: 551
</treatmentCitation>
</taxonomicName>
(redescription,
<typeStatus id="54C98839FF9CD531CCC8F9F2D16DF923" box="[746,839,1648,1670]" pageId="10" pageNumber="461" type="lectotype">lectotype</typeStatus>
designed, distribution, notes).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9CD531CEB5F931D2D0F849" pageId="10" pageNumber="461" type="materials_examined">
<paragraph id="8BCD369BFF9CD531CEB5F931D730F801" blockId="10.[151,1436,1715,2028]" pageId="10" pageNumber="461">
<emphasis id="B906EA89FF9CD531CEB5F931D363F969" bold="true" box="[151,329,1715,1740]" pageId="10" pageNumber="461">Type material.</emphasis>
<taxonomicName id="4C724D18FF9CD531CF77F936D1A8F96E" authority="Bernhauer, 1917" authorityName="Bernhauer" authorityYear="1917" box="[341,898,1715,1740]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andinus" subGenus="Antropiestus">
<emphasis id="B906EA89FF9CD531CF77F936D38CF96E" box="[341,422,1716,1739]" italics="true" pageId="10" pageNumber="461">Piestus</emphasis>
(
<emphasis id="B906EA89FF9CD531CF99F936D06DF96E" box="[443,583,1716,1739]" italics="true" pageId="10" pageNumber="461">Antropiestus</emphasis>
)
<emphasis id="B906EA89FF9CD531CC7EF931D09FF969" box="[604,693,1715,1740]" italics="true" pageId="10" pageNumber="461">andinus</emphasis>
<bibRefCitation id="EFE34B6AFF9CD531CCE3F931D1A8F96E" author="Bernhauer" box="[705,898,1715,1740]" pageId="10" pageNumber="461" refString="Bernhauer, M. (1917) Neue arten der gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. Neue Beitrage zur Systematischen Insektenkunde, 1, 45 - 53." type="journal article" year="1917">Bernhauer, 1917</bibRefCitation>
</taxonomicName>
. Lectotype (designated by
<bibRefCitation id="EFE34B6AFF9CD531CAE0F931D7BFF96E" author="Caron" box="[1218,1429,1715,1740]" pageId="10" pageNumber="461" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B906EA89FF9CD531CB30F936D76DF969" box="[1298,1351,1715,1740]" italics="true" pageId="10" pageNumber="461">et al</emphasis>
. 2012
</bibRefCitation>
) deposited in FMNH, male [damaged specimen: without right anterior leg] with labels: (1) “♂” [white label, handwritten, together with the specimen]; (2) “Columbia occ/Cali. Fassl” [white label/printed in black]; (3) “
<taxonomicName id="4C724D18FF9CD531CE83F89DD31EF89D" box="[161,308,1823,1848]" class="Insecta" family="Staphylinidae" genus="Antropiestus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="462" phylum="Arthropoda" rank="genus">Antropiestus</taxonomicName>
/
<taxonomicName id="4C724D18FF9CD531CF1EF89DD3B9F89D" box="[316,403,1823,1848]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andinus">andinus</taxonomicName>
/Brnh. Typus” [light yellow label, handwritten]; (4) “♂” [white label, printed in black]; (5) “LECTOTYPE/
<taxonomicName id="4C724D18FF9CD531CF69F8C1D3D3F8F9" box="[331,505,1859,1884]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="andinus">Piestus andinus</taxonomicName>
/
<bibRefCitation id="EFE34B6AFF9CD531CC20F8C1D094F8F9" author="Bernhauer" box="[514,702,1859,1884]" pageId="10" pageNumber="461" refString="Bernhauer, M. (1917) Neue arten der gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. Neue Beitrage zur Systematischen Insektenkunde, 1, 45 - 53." type="journal article" year="1917">Bernhauer, 1917</bibRefCitation>
/det. E. Caron, 2009” [red label, printed in black]; (6) “Chicago NHMus/M.Bernhauer/Collection” [white label, printed in black]; (7) “FMNH-INS/0000 0 63 446” [white label, printed in black]; (8) &quot;Photographed/Kelsey Keaton 2014/Emub Catalog&quot; [blue label, printed in black].
</paragraph>
<paragraph id="8BCD369BFF9CD531CEE5F832D2D0F849" blockId="10.[151,1436,1715,2028]" pageId="10" pageNumber="461">
<materialsCitation id="3B1A3CC6FF9CD531CEE5F832D2D0F849" ID-GBIF-Occurrence="1563407728" collectionCode="NMW" country="Austria" location="Wien" pageId="10" pageNumber="461" specimenCount="1" stateProvince="Wien" typeStatus="paralectotype">
<emphasis id="B906EA89FF9CD531CEE5F832D2D6F862" box="[199,252,1968,1991]" italics="true" pageId="10" pageNumber="461">Note</emphasis>
:
<bibRefCitation id="EFE34B6AFF9CD531CF2EF82DD3CEF862" box="[268,484,1967,1992]" pageId="10" pageNumber="461" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article">
Caron
<emphasis id="B906EA89FF9CD531CF7AF832D3A0F86D" box="[344,394,1967,1992]" italics="true" pageId="10" pageNumber="461">et al</emphasis>
. (2012)
</bibRefCitation>
cited one
<typeStatus id="54C98839FF9CD531CC79F82DD0D8F86D" box="[603,754,1967,1992]" pageId="10" pageNumber="461" type="paralectotype">paralectotype</typeStatus>
deposited in
<collectionCode id="ED63AE5EFF9CD531CDA9F832D1F8F862" box="[907,978,1968,1991]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="461">NMW</collectionCode>
(Naturhistorische Museum
<collectingRegion id="49B6F879FF9CD531CB2DF82DD764F86D" box="[1295,1358,1967,1992]" country="Austria" name="Wien" pageId="10" pageNumber="461">Wien</collectingRegion>
,
<collectingRegion id="49B6F879FF9CD531CB7BF82DD7B2F86D" box="[1369,1432,1967,1992]" country="Austria" name="Wien" pageId="10" pageNumber="461">Wien</collectingRegion>
,
<collectingCountry id="F365760BFF9CD531CEB5F851D2C5F849" box="[151,239,2003,2028]" name="Austria" pageId="10" pageNumber="461">Austria</collectingCountry>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9DD530CEE5FF1BD33FFF70" pageId="11" pageNumber="462" type="reference_group">
<paragraph id="8BCD369BFF9DD530CEE5FF1BD33FFF70" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<taxonomicName id="4C724D18FF9DD530CEE5FF1BD136FF15" authority="Bernhauer, 1921" authorityName="Bernhauer" authorityYear="1921" box="[199,796,151,176]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="strigipennis" subGenus="Antropiestus">
<emphasis id="B906EA89FF9DD530CEE5FF1BD332FF15" box="[199,280,153,176]" italics="true" pageId="11" pageNumber="462">Piestus</emphasis>
(
<emphasis id="B906EA89FF9DD530CF0EFF1AD392FF0A" box="[300,440,152,175]" italics="true" pageId="11" pageNumber="462">Antropiestus</emphasis>
)
<emphasis id="B906EA89FF9DD530CFEEFF1BD07AFF15" box="[460,592,153,176]" italics="true" pageId="11" pageNumber="462">strigipennis</emphasis>
<bibRefCitation id="EFE34B6AFF9DD530CC7EFF15D136FF15" author="Bernhauer" box="[604,796,151,176]" pageId="11" pageNumber="462" refString="Bernhauer, M. (1921) Zur Staphyliniden fauna von Sudamerika. Deutsche Entomologische Zeitschrift, 1921, 65 - 77." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
</taxonomicName>
. Syntype deposited in FMNH (
<bibRefCitation id="EFE34B6AFF9DD530CAA9FF1AD776FF15" author="Caron" box="[1163,1372,151,176]" pageId="11" pageNumber="462" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B906EA89FF9DD530CAFEFF1BD732FF15" box="[1244,1304,151,176]" italics="true" pageId="11" pageNumber="462">et al.</emphasis>
2012
</bibRefCitation>
) (not examined).
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9DD530CEE5FF5DD63CFEB9" pageId="11" pageNumber="462" type="diagnosis">
<paragraph id="8BCD369BFF9DD530CEE5FF5DD63CFEB9" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<emphasis id="B906EA89FF9DD530CEE5FF5DD36BFF5D" bold="true" box="[199,321,223,248]" pageId="11" pageNumber="462">Diagnosis.</emphasis>
<taxonomicName id="4C724D18FF9DD530CF71FF5DD005FF5D" box="[339,559,223,248]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B906EA89FF9DD530CF71FF5DD005FF5D" box="[339,559,223,248]" italics="true" pageId="11" pageNumber="462">Hypotelus andinus</emphasis>
</taxonomicName>
may be easily distinguished from other species of
<taxonomicName id="4C724D18FF9DD530CAEAFF5DD713FF5D" box="[1224,1337,223,248]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA89FF9DD530CAEAFF5DD713FF5D" box="[1224,1337,223,248]" italics="true" pageId="11" pageNumber="462">Hypotelus</emphasis>
</taxonomicName>
by the conspicuous, deep and large depression at the middle of the pronotum (
<figureCitation id="13492A1EFF9DD530CD97FE86D622FEB9" box="[949,1032,260,285]" captionStart="FIGURES 9 19" captionStartId="9.[151,264,1913,1935]" captionTargetBox="[216,1355,230,1832]" captionTargetId="figure@9.[151,1436,157,1945]" captionTargetPageId="9" captionText="FIGURES 9 19. Habitus, dorsal view. 9, Hypotelus insulanus, syntype, male; 10, H. praecox, lectotype, female; 11, H. scheerpeltzi sp. nov., holotype, male; 12, H. testaceus, syntype, male; 13, H. melanodelta sp. nov., holotype, female; 14, H. castaneus sp. nov., holotype, male; 15, H. brevitarsus sp. nov., holotype, male; 16, H. corniculatus sp. nov., holotype, female; 17, H. marginatus, syntype, male; 18, H. laevis, male; 19, H. andinus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/803652/files/figure.png" pageId="11" pageNumber="462">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9DD530CEE5FEA5D62AFAA9" pageId="11" pageNumber="462" type="description">
<paragraph id="8BCD369BFF9DD530CEE5FEA5D645FE08" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<emphasis id="B906EA89FF9DD530CEE5FEA5D35EFEE5" bold="true" box="[199,372,295,320]" pageId="11" pageNumber="462">Redescription.</emphasis>
BL:
<quantity id="4C8A9B7EFF9DD530CF8DFEA5D022FE9A" box="[431,520,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="11" pageNumber="462" unit="mm" value="6.1">6.1 mm</quantity>
, BW:
<quantity id="4C8A9B7EFF9DD530CC72FEAAD083FE9A" box="[592,681,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="462" unit="mm" value="1.3">1.3 mm</quantity>
. Body slightly convex; dorsal surface matte, except elytra; entirely black with legs a little lighter (
<figureCitation id="13492A1EFF9DD530CFCEFECED015FEC1" box="[492,575,332,357]" captionStart="FIGURES 9 19" captionStartId="9.[151,264,1913,1935]" captionTargetBox="[216,1355,230,1832]" captionTargetId="figure@9.[151,1436,157,1945]" captionTargetPageId="9" captionText="FIGURES 9 19. Habitus, dorsal view. 9, Hypotelus insulanus, syntype, male; 10, H. praecox, lectotype, female; 11, H. scheerpeltzi sp. nov., holotype, male; 12, H. testaceus, syntype, male; 13, H. melanodelta sp. nov., holotype, female; 14, H. castaneus sp. nov., holotype, male; 15, H. brevitarsus sp. nov., holotype, male; 16, H. corniculatus sp. nov., holotype, female; 17, H. marginatus, syntype, male; 18, H. laevis, male; 19, H. andinus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/803652/files/figure.png" pageId="11" pageNumber="462">Fig. 19</figureCitation>
). Dorsal integument of head and pronotum with fine punctures and granulate microsculpture; elytra each with one longitudinal finely punctate stria close to sutural margin, conspicuous only on basal half; metaventrite and abdomen with fine punctures and granulate microsculpture.
</paragraph>
<paragraph id="8BCD369BFF9DD530CEE5FE35D0CAFD20" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<emphasis id="B906EA89FF9DD530CEE5FE35D364FE75" box="[199,334,439,464]" italics="true" pageId="11" pageNumber="462">Male. Head</emphasis>
. With five moderately sized punctures with long setae on basal half of dorsal margin of eyes; supra-antennal area visibly prominent. Antennae almost reaching apex of elytra; every antennomeres with some dispersed long setae, setae on inner face are longest; antennomere 3 longer than 2; 511 gradually increasing in length toward antennal apex. Labrum with median third deeply emarginate, three pairs of long setae medially; lateral thirds with two pairs of long setae, two apical and two subapical. Mandibles symmetrical, one acute tooth on each inner margin. Mentum 2 times as wide as long.
</paragraph>
<paragraph id="8BCD369BFF9DD530CEE5FD0DD644FCB1" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<emphasis id="B906EA89FF9DD530CEE5FD0DD30AFD0D" box="[199,288,655,680]" italics="true" pageId="11" pageNumber="462">Thorax.</emphasis>
Pronotum wider than long (PW/PL=1.25); anterior angles rounded and slightly prominent; apical twothirds with somewhat curved sides and basal one-third gradually narrowing toward the base; with complete internal mid-longitudinal ridge and conspicuous longitudinal median sulcus; conspicuous and deep large depression at middle. Elytra somewhat longer than wide (EL/BW=1.2), covering abdominal tergite 3.
</paragraph>
<paragraph id="8BCD369BFF9DD530CEE5FC9DD067FC48" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<emphasis id="B906EA89FF9DD530CEE5FC9DD312FC9D" box="[199,312,799,824]" italics="true" pageId="11" pageNumber="462">Abdomen.</emphasis>
Apical abdominal segments 8 to 10 and genitalia were not studied, but according to
<bibRefCitation id="EFE34B6AFF9DD530CB31FCA2D2CBFCF9" pageId="11" pageNumber="462" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article">
Caron
<emphasis id="B906EA89FF9DD530CB43FCA3D7BFFC9D" box="[1377,1429,799,824]" italics="true" pageId="11" pageNumber="462">et al</emphasis>
. (2012)
</bibRefCitation>
the shape of the structures are: sternite 8 with posterior margin truncate; tergite 9 with short ventral struts; tergite 10 with some long setae on apex and posterior margin with short fringes, basal half not divided longitudinally; sternite 9 with two pairs of long setae and truncate apex. Median lobe of aedeagus somewhat globose apex in lateral view, tube closed in dorsal view; length of parameres exceeds apex of median lobe by a little (see
<bibRefCitation id="EFE34B6AFF9DD530CEE9FC56D3BAFC49" author="Caron" box="[203,400,980,1005]" pageId="11" pageNumber="462" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B906EA89FF9DD530CF34FC57D36CFC48" box="[278,326,980,1005]" italics="true" pageId="11" pageNumber="462">et al</emphasis>
. 2012
</bibRefCitation>
, Figs. 248250).
</paragraph>
<paragraph id="8BCD369BFF9DD530CEE5FC75D15DFB05" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<emphasis id="B906EA89FF9DD530CEE5FC75D309FBB5" box="[199,291,1015,1040]" italics="true" pageId="11" pageNumber="462">Female.</emphasis>
Not examined. According to
<bibRefCitation id="EFE34B6AFF9DD530CC4DFC7AD169FBB5" box="[623,835,1015,1040]" pageId="11" pageNumber="462" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article">
Caron
<emphasis id="B906EA89FF9DD530CC98FC7BD0C1FBB5" box="[698,747,1015,1040]" italics="true" pageId="11" pageNumber="462">et al</emphasis>
. (2012)
</bibRefCitation>
the female is similar to male except for: elytra a little shorter, covering only basal half of abdominal tergite 3; abdominal sternite 8 with pointed apex and short setae on posterior margin; tergite 10 divided longitudinally in basal half (see
<bibRefCitation id="EFE34B6AFF9DD530CDE4FBC2D6B1FBFD" author="Caron" box="[966,1179,1087,1112]" pageId="11" pageNumber="462" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B906EA89FF9DD530CA34FBC3D667FBFD" box="[1046,1101,1087,1112]" italics="true" pageId="11" pageNumber="462">et al</emphasis>
. 2012
</bibRefCitation>
, Fig. 151); ovipositor consisting of a pair of weakly pigmented hemisternites and a pair of more apical coxites, and with many long setae on apex (see
<bibRefCitation id="EFE34B6AFF9DD530CF08FB0AD3C5FB05" author="Caron" box="[298,495,1159,1184]" pageId="11" pageNumber="462" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B906EA89FF9DD530CF57FB0BD38CFB05" box="[373,422,1159,1184]" italics="true" pageId="11" pageNumber="462">et al</emphasis>
. 2012
</bibRefCitation>
, Fig. 155). Spermatheca unknown.
</paragraph>
<paragraph id="8BCD369BFF9DD530CEE5FB2ED110FB4D" blockId="11.[151,1436,151,1293]" pageId="11" pageNumber="462">
<materialsCitation id="3B1A3CC6FF9DD530CEE5FB2ED110FB4D" ID-GBIF-Occurrence="1563407727" country="Colombia" location="Beni" pageId="11" pageNumber="462" specimenCount="1" stateProvince="Beni">
<emphasis id="B906EA89FF9DD530CEE5FB2ED3F9FB60" bold="true" box="[199,467,1196,1221]" pageId="11" pageNumber="462">Geographical records.</emphasis>
In the current study, the species was examined from
<collectingCountry id="F365760BFF9DD530CA62FB2ED698FB60" box="[1088,1202,1196,1221]" name="Colombia" pageId="11" pageNumber="462">Colombia</collectingCountry>
.
<bibRefCitation id="EFE34B6AFF9DD530CA9DFB2ED7B6FB61" box="[1215,1436,1196,1221]" pageId="11" pageNumber="462" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25, 490 - 585. https: // doi. org / 10.1071 / IS 10016" type="journal article">
Caron
<emphasis id="B906EA89FF9DD530CB2FFB2FD76AFB60" box="[1293,1344,1196,1221]" italics="true" pageId="11" pageNumber="462">et al</emphasis>
. (2012)
</bibRefCitation>
also recorded it from
<collectingCountry id="F365760BFF9DD530CFAAFB4DD3CFFB4D" box="[392,485,1231,1256]" name="Ecuador" pageId="11" pageNumber="462">Ecuador</collectingCountry>
and
<collectingCountry id="F365760BFF9DD530CC3EFB4DD045FB4D" box="[540,623,1231,1256]" name="Bolivia" pageId="11" pageNumber="462">Bolivia</collectingCountry>
(
<collectingRegion id="49B6F879FF9DD530CC5FFB4DD09FFB4D" box="[637,693,1231,1256]" country="Bolivia" name="Beni" pageId="11" pageNumber="462">Beni</collectingRegion>
) (
<figureCitation id="13492A1EFF9DD530CCEEFB4DD107FB4D" box="[716,813,1231,1256]" captionStart="FIGURE 114" captionStartId="31.[151,250,1701,1723]" captionTargetBox="[190,1375,215,1655]" captionTargetId="figure@31.[150,1437,153,1679]" captionTargetPageId="31" captionText="FIGURE 114. Geographical records of Hypotelus. Hypotelus andinus, Colombia, Ecuador and Bolivia; H. insulanus, Caribbean islands; H. marginatus, Colombia, Venezuela and Guatemala; H. testaceus, Panama; H. brevitarsus sp. nov., Panama. Countries without dots but colored represent the occurrence but no locality given." httpUri="https://zenodo.org/record/803674/files/figure.png" pageId="11" pageNumber="462">Fig. 114</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCD369BFF9DD530CEE5FB76D62AFAA9" blockId="11.[151,1436,151,1293]" box="[199,1024,1268,1293]" pageId="11" pageNumber="462">
<emphasis id="B906EA89FF9DD530CEE5FB76D3A3FAA8" bold="true" box="[199,393,1268,1293]" pageId="11" pageNumber="462">Biological notes.</emphasis>
<taxonomicName id="4C724D18FF9DD530CFB2FB76D048FAA8" box="[400,610,1268,1293]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B906EA89FF9DD530CFB2FB76D048FAA8" box="[400,610,1268,1293]" italics="true" pageId="11" pageNumber="462">Hypotelus andinus</emphasis>
</taxonomicName>
has been collected in a Malaise trap.
</paragraph>
</subSubSection>
</treatment>
</document>