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<mods:title id="224265D18993AE5BBB6A4F45115658F7">The flat-footed fly genus Seri Kessel &amp; Kessel (Diptera: Platypezidae)</mods:title>
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<mods:namePart id="06F0CB298EDF73D3CB604EEBAFDCAD06">Cumming, Jeffrey M.</mods:namePart>
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<treatment id="03DB1D4DFFEDFFEDFF4AFE0A6D04F8EB" ID-DOI="http://doi.org/10.5281/zenodo.6187565" ID-GBIF-Taxon="119427279" ID-Zenodo-Dep="6187565" LSID="urn:lsid:plazi:treatment:03DB1D4DFFEDFFEDFF4AFE0A6D04F8EB" httpUri="http://treatment.plazi.org/id/03DB1D4DFFEDFFEDFF4AFE0A6D04F8EB" lastPageNumber="62" pageId="1" pageNumber="62">
<subSubSection id="C368FFD0FFEDFFEDFF4AFE0A6EA5FE01" box="[151,493,437,463]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="8BCDAC5BFFEDFFEDFF4AFE0A6EA5FE01" blockId="1.[151,493,437,463]" box="[151,493,437,463]" pageId="1" pageNumber="62">
<heading id="D0851B37FFEDFFEDFF4AFE0A6EA5FE01" bold="true" box="[151,493,437,463]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="B9067049FFEDFFEDFF4AFE0A6EA5FE01" bold="true" box="[151,493,437,463]" pageId="1" pageNumber="62">
Genus
<taxonomicName id="4C72D7D8FFEDFFEDFF33FE0A6EA5FE01" ID-CoL="7FRV" ID-ENA="1445834" authority="Kessel &amp; Kessel" authorityName="Kessel &amp; Kessel" box="[238,493,437,463]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFF33FE0A6E57FE01" bold="true" box="[238,287,437,463]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
Kessel &amp; Kessel
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368FFD0FFEDFFEDFF4AFE446BF4FD9B" pageId="1" pageNumber="62" type="reference_group">
<paragraph id="8BCDAC5BFFEDFFEDFF4AFE446BA7FDDE" blockId="1.[151,1263,506,528]" box="[151,1263,506,528]" pageId="1" pageNumber="62">
<treatmentCitationGroup id="AB628B75FFEDFFEDFF4AFE446BA7FDDE" box="[151,1263,506,528]" pageId="1" pageNumber="62">
<taxonomicName id="4C72D7D8FFEDFFEDFF4AFE446E9EFDDE" ID-CoL="7FRV" authority="Kessel &amp; Kessel, 1966: 98" authorityName="Kessel &amp; Kessel" authorityPageNumber="98" authorityYear="1966" box="[151,470,506,528]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFF4AFE446F88FDDE" box="[151,192,507,528]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
<treatmentCitation id="0AD38A4AFFEDFFEDFF18FE456E9EFDDE" author="Kessel" box="[197,470,506,528]" page="98" pageId="1" pageNumber="62" year="1966">
<bibRefCitation id="EFE3D1AAFFEDFFEDFF18FE456EF8FDDE" author="Kessel" box="[197,432,506,528]" pageId="1" pageNumber="62" refString="Kessel, E. L. &amp; Kessel, B. B. (1966) Seri, a new genus of Platypezidae from North America (Diptera). The Wasmann Journal of Biology, 24, 97 - 100." type="journal article" year="1966">Kessel &amp; Kessel, 1966</bibRefCitation>
: 98
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54C912F9FFEDFFEDFE02FE446D59FDDE" box="[479,529,507,528]" pageId="1" pageNumber="62">Type</typeStatus>
species:
<taxonomicName id="4C72D7D8FFEDFFEDFDB2FE456CCDFDDE" authority="Kessel 1961" authorityName="Kessel" authorityYear="1961" box="[623,901,506,528]" class="Insecta" family="Platypezidae" genus="Clythia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="dymka">
<emphasis id="B9067049FFEDFFEDFDB2FE456C4AFDDE" box="[623,770,506,528]" italics="true" pageId="1" pageNumber="62">Clythia dymka</emphasis>
<bibRefCitation id="EFE3D1AAFFEDFFEDFCD5FE456CCDFDDE" author="Kessel" box="[776,901,506,528]" pageId="1" pageNumber="62" refString="Kessel, E. L. (1961) New species of flat-footed flies from North America (Diptera: Platypezidae). The Wasmann Journal of Biology, 19, 191 - 227." type="journal article" year="1961">Kessel 1961</bibRefCitation>
</taxonomicName>
[Nearctic], by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCDAC5BFFEDFFEDFF4AFD826BF4FD9B" blockId="1.[151,1436,573,1829]" box="[151,1212,573,598]" pageId="1" pageNumber="62">
<emphasis id="B9067049FFEDFFEDFF4AFD826E29FD98" bold="true" box="[151,353,573,598]" pageId="1" pageNumber="62">Included species.</emphasis>
<taxonomicName id="4C72D7D8FFEDFFEDFEB5FD816EDDFD9B" ID-CoL="7FRV" box="[360,405,574,597]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFEB5FD816EDDFD9B" box="[360,405,574,597]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
</taxonomicName>
currently includes
<taxonomicName id="4C72D7D8FFEDFFEDFDB1FD816C7EFD9B" authority="Kessel" authorityName="Kessel" box="[620,822,573,597]" class="Insecta" family="Platypezidae" genus="Clythia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="dymka">
<emphasis id="B9067049FFEDFFEDFDB1FD816D99FD9B" box="[620,721,573,597]" italics="true" pageId="1" pageNumber="62">S. dymka</emphasis>
(Kessel)
</taxonomicName>
and
<taxonomicName id="4C72D7D8FFEDFFEDFCB1FD816BF0FD9B" ID-CoL="6YHGH" authority="Oldenberg" authorityName="Oldenberg" box="[876,1208,573,597]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B9067049FFEDFFEDFCB1FD816B6DFD9B" box="[876,1061,573,597]" italics="true" pageId="1" pageNumber="62">S. obscuripennis</emphasis>
(Oldenberg)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C368FFD0FFEDFFEDFF1AFDDE6B68FB9E" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="8BCDAC5BFFEDFFEDFF1AFDDE6B68FB9E" blockId="1.[151,1436,573,1829]" pageId="1" pageNumber="62">
<emphasis id="B9067049FFEDFFEDFF1AFDDE6E09FDB4" bold="true" box="[199,321,609,634]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
<taxonomicName id="4C72D7D8FFEDFFEDFE95FDDD6E3DFDB7" box="[328,373,610,633]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFE95FDDD6E3DFDB7" box="[328,373,610,633]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
</taxonomicName>
is distinguished from other platypezid genera by the following combination of characters: relatively small flies
<quantity id="4C8A01BEFFEDFFEDFEB2FD3A6E87FD53" box="[367,463,645,669]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="62" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
in length with body (Figs. 1, 2) uniformly colored grey to black (male darker than female), without abdominal markings in female; head with parafacial bare, frons bearing long frontal setae in male (short setae in female); antenna with scape bare, pedicel with circle of preapical setae extended less than half length of postpedicel, postpedicel small and ovate, arista apical; thorax lacking acrostichal setae, with 3 notopleural setae; wing (Fig. 3) tinted brown in male and lighter greyish brown in female, covered with conspicuous microtrichia, crossvein dm-m less than or subequal to its own length from wing margin, vein M1+2 branched with furcation nearer to crossvein dm-m than to wing margin, cell cua about equal in length to apical section of anal vein (CuA+CuP); legs with mid tibia bearing one apical spur (lacking anteroventral spur); female hind tarsus with soles on anteroventral surface of tarsomeres 3 and 4, tarsomeres 13 broad, tarsomere 2 distinctly narrowed ventrally, tarsomere 3 longest; female abdomen with terminalia short, mostly retracted into segment 7, sternite 7 with one or two pairs of strong erect setae; hypopygium of male (
<figureCitation id="1349B0DEFFEDFFEDFDB4FC4F6DE5FBC6" box="[617,685,1008,1032]" captionStart="FIGURE 4" captionStartId="3.[151,250,1468,1491]" captionTargetBox="[263,1321,387,1435]" captionTargetId="figure@3.[250,1336,381,1448]" captionTargetPageId="3" captionText="FIGURE 4. Male genitalia of Seri dymka (Kessel), left lateral view. Abbreviations: cerc—cercus, epand—epandrium, ej apod—ejaculatory apodeme, hypd—hypandrium, hypd lb—hypandrial lobe, hyprct—hypoproct, phapod—phallapodeme, ph—phallus, spm sac—sperm sac, sur—surstylus. Scale in mm." httpUri="https://zenodo.org/record/279514/files/figure.png" pageId="1" pageNumber="62">Fig. 4</figureCitation>
) with epandrium terminating in short blunt surstylus, hypandrium with paired elongate ventrally curved apical processes, phallapodeme keel-like, postgonite absent, phallus slender and elongate, sperm pump large and sclerotized with small ejaculatory apodeme.
</paragraph>
</subSubSection>
<subSubSection id="C368FFD0FFEDFFEDFF1AFBE46AC9FBBA" box="[199,1409,1115,1141]" pageId="1" pageNumber="62" type="distribution">
<paragraph id="8BCDAC5BFFEDFFEDFF1AFBE46AC9FBBA" blockId="1.[151,1436,573,1829]" box="[199,1409,1115,1141]" pageId="1" pageNumber="62">
<emphasis id="B9067049FFEDFFEDFF1AFBE46E28FBBA" bold="true" box="[199,352,1115,1140]" pageId="1" pageNumber="62">Distribution.</emphasis>
The genus is known from the Palaearctic (
<taxonomicName id="4C72D7D8FFEDFFEDFC9DFBE26CB1FBBA" box="[832,1017,1116,1140]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B9067049FFEDFFEDFC9DFBE26CB1FBBA" box="[832,1017,1116,1140]" italics="true" pageId="1" pageNumber="62">S. obscuripennis</emphasis>
</taxonomicName>
) and Nearctic (
<taxonomicName id="4C72D7D8FFEDFFEDFB75FBE26A45FBBA" box="[1192,1293,1116,1140]" class="Insecta" family="Platypezidae" genus="Clythia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="dymka">
<emphasis id="B9067049FFEDFFEDFB75FBE26A45FBBA" box="[1192,1293,1116,1140]" italics="true" pageId="1" pageNumber="62">S. dymka</emphasis>
</taxonomicName>
) Regions.
</paragraph>
</subSubSection>
<subSubSection id="C368FFD0FFEDFFEDFF1AFB3F6D04F8EB" pageId="1" pageNumber="62" type="discussion">
<paragraph id="8BCDAC5BFFEDFFEDFF1AFB3F6C38FAC9" blockId="1.[151,1436,573,1829]" pageId="1" pageNumber="62">
<emphasis id="B9067049FFEDFFEDFF1AFB3F6E73FB56" bold="true" box="[199,315,1152,1176]" pageId="1" pageNumber="62">Remarks.</emphasis>
Adult specimens of
<taxonomicName id="4C72D7D8FFEDFFEDFDF8FB3E6D1AFB56" box="[549,594,1153,1176]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFDF8FB3E6D1AFB56" box="[549,594,1153,1176]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
</taxonomicName>
are most easily recognized by their tinted wings, which are conspicuously covered with microtrichia unlike many other platypezids that have clear glassier wings. The venation is also distinctive in that the furcation of M1+2 is nearer to crossvein dm-m than to the wing margin, rather than the more common condition of having the furcation nearer to the wing margin.
</paragraph>
<paragraph id="8BCDAC5BFFEDFFEDFF1AFAAC6B70F9BF" blockId="1.[151,1436,573,1829]" pageId="1" pageNumber="62">
According to
<bibRefCitation id="EFE3D1AAFFEDFFEDFEBEFAAC6D54FAE5" author="Chandler" box="[355,540,1299,1323]" pageId="1" pageNumber="62" refString="Chandler, P. J. (2001) The flat-footed flies (Diptera: Opetiidae and Platypezidae) of Europe. Fauna Entomologica Scandinavica, 36, 1 - 276." type="journal article" year="2001">Chandler (2001)</bibRefCitation>
<taxonomicName id="4C72D7D8FFEDFFEDFDF9FAAB6D19FAE5" box="[548,593,1300,1323]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFDF9FAAB6D19FAE5" box="[548,593,1300,1323]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
</taxonomicName>
is most similar to the monotypic Palaearctic genus
<taxonomicName id="4C72D7D8FFEDFFEDFB46FAAC6A24FAE5" authority="Enderlein" authorityName="Enderlein" box="[1179,1388,1299,1323]" class="Insecta" family="Platypezidae" genus="Bolopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFB46FAAC6BBFFAE5" box="[1179,1271,1299,1323]" italics="true" pageId="1" pageNumber="62">Bolopus</emphasis>
Enderlein
</taxonomicName>
and is also similar to the small Holarctic/Oriental genus
<taxonomicName id="4C72D7D8FFEDFFEDFD02FA886BADFA81" authority="Kessel &amp; Maggioncalda. Unlike" authorityName="Kessel &amp; Maggioncalda. Unlike" box="[735,1253,1335,1359]" class="Insecta" family="Platypezidae" genus="Polyporivora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFD02FA886C3CFA81" box="[735,884,1335,1359]" italics="true" pageId="1" pageNumber="62">Polyporivora</emphasis>
Kessel &amp; Maggioncalda. Unlike
</taxonomicName>
most other genera in the
<taxonomicName id="4C72D7D8FFEDFFEDFEDAFAE46EDEFABD" box="[263,406,1371,1395]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="subFamily" subFamily="Platypezinae">Platypezinae</taxonomicName>
the mid tibia of these three genera ends in a single posteroventral spur, with the anteroventral spur being extremely reduced or absent. The antennal scape and parafacial are also bare in these three genera, and the male genitalia includes a large sclerotized sperm pump. Unlike
<taxonomicName id="4C72D7D8FFEDFFEDFC16FA1C6B28FA75" box="[971,1120,1443,1467]" class="Insecta" family="Platypezidae" genus="Polyporivora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFC16FA1C6B28FA75" box="[971,1120,1443,1467]" italics="true" pageId="1" pageNumber="62">Polyporivora</emphasis>
</taxonomicName>
however,
<taxonomicName id="4C72D7D8FFEDFFEDFB04FA1C6A7DFA75" box="[1241,1333,1443,1467]" class="Insecta" family="Platypezidae" genus="Bolopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFB04FA1C6A7DFA75" box="[1241,1333,1443,1467]" italics="true" pageId="1" pageNumber="62">Bolopus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C72D7D8FFEDFFEDFAB2FA1B6AD4FA75" box="[1391,1436,1444,1467]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFAB2FA1B6AD4FA75" box="[1391,1436,1444,1467]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
</taxonomicName>
both have the wing covered with conspicuous microtrichia, the furcation of M1+2 nearer to crossvein dm-m than to the wing margin, and females that lack a pattern of abdominal markings.
<taxonomicName id="4C72D7D8FFEDFFEDFC19FA526B68F9CB" box="[964,1056,1517,1541]" class="Insecta" family="Platypezidae" genus="Bolopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFC19FA526B68F9CB" box="[964,1056,1517,1541]" italics="true" pageId="1" pageNumber="62">Bolopus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C72D7D8FFEDFFEDFB6FFA516B97F9CB" box="[1202,1247,1518,1541]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFB6FFA516B97F9CB" box="[1202,1247,1518,1541]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
</taxonomicName>
in that crossvein dm-m is more than 1.5X its length from the wing margin and the antenna has a circle of much longer preapical setae on the pedicel and a subdorsal rather than apical arista. In addition, the male genitalia of
<taxonomicName id="4C72D7D8FFEDFFEDFB36F9896A0FF980" box="[1259,1351,1590,1614]" class="Insecta" family="Platypezidae" genus="Bolopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFB36F9896A0FF980" box="[1259,1351,1590,1614]" italics="true" pageId="1" pageNumber="62">Bolopus</emphasis>
</taxonomicName>
have a larger pointed surstylus, a shorter hypandrial process and a short straighter phallus.
</paragraph>
<paragraph id="8BCDAC5BFFEDFFEDFF1AF9C26D04F8EB" blockId="1.[151,1436,573,1829]" pageId="1" pageNumber="62">
<bibRefCitation id="EFE3D1AAFFEDFFEDFF1AF9C26EA0F95B" author="Kessel" box="[199,488,1661,1685]" pageId="1" pageNumber="62" refString="Kessel, E. L. &amp; Kessel, B. B. (1966) Seri, a new genus of Platypezidae from North America (Diptera). The Wasmann Journal of Biology, 24, 97 - 100." type="journal article" year="1966">Kessel and Kessel (1966)</bibRefCitation>
and
<bibRefCitation id="EFE3D1AAFFEDFFEDFDFCF9C26D92F95B" author="Chandler" box="[545,730,1661,1685]" pageId="1" pageNumber="62" refString="Chandler, P. J. (1974) Additions and corrections to the British list of Platypezidae (Diptera), incorporating a revision of the Palaearctic species of Callomyia Meigen. Proceedings and Transactions of the British Entomological Society, 7, 1 - 32. Chandler, P. J. (1991) Platypezidae. In: Soos, A. &amp; Papp, L. (Eds.), Catalogue of Palaearctic Diptera, Volume 7, Dolichopodidae - Platypezidae. Elsevier Science Publishing, pp. 205 - 217." type="journal article" year="1974">Chandler (1974)</bibRefCitation>
both commented, respectively, on how rarely
<taxonomicName id="4C72D7D8FFEDFFEDFB35F9C16A06F958" box="[1256,1358,1662,1686]" class="Insecta" family="Platypezidae" genus="Clythia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="dymka">
<emphasis id="B9067049FFEDFFEDFB35F9C16A06F958" box="[1256,1358,1662,1686]" italics="true" pageId="1" pageNumber="62">S. dymka</emphasis>
</taxonomicName>
and
<taxonomicName id="4C72D7D8FFEDFFEDFA5AF9C16E7CF974" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B9067049FFEDFFEDFA5AF9C16E7CF974" italics="true" pageId="1" pageNumber="62">S. obscuripennis</emphasis>
</taxonomicName>
appear to be collected, suggesting that each species may be overlooked by specialists because when resting they tend to resemble small muscid flies. However, with increased use of passive trapping techniques such as Malaise traps and pan traps during the last 40 years, many more specimens of
<taxonomicName id="4C72D7D8FFEDFFEDFBF8F9556B1AF8CF" box="[1061,1106,1770,1793]" class="Insecta" family="Platypezidae" genus="Seri" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B9067049FFEDFFEDFBF8F9556B1AF8CF" box="[1061,1106,1770,1793]" italics="true" pageId="1" pageNumber="62">Seri</emphasis>
</taxonomicName>
would be expected in collections if these flies were more common.
</paragraph>
</subSubSection>
</treatment>
</document>