treatments-xml/data/03/A0/36/03A03624FFF3B1044DA4ACEAFE9FF928.xml
2024-06-21 12:22:17 +02:00

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<document id="D3D0D94265C9F8FF9015BC5FBBD982C4" ID-DOI="10.11646/zootaxa.5437.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="10985633" ID-ZooBank="7114B49F-4A44-4165-9FC6-8A52A6E898F9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713347432768" checkinUser="plazi" docAuthor="Kasparyan, Dmitry R. &amp; Khalaim, Andrey I." docDate="2024" docId="03A03624FFF3B1044DA4ACEAFE9FF928" docLanguage="en" docName="zootaxa.5437.4.9.pdf" docOrigin="Zootaxa 5437 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5437.4.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.18:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="18" docTitle="Hylophasma aterrima Kasparyan &amp; Khalaim 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="579" masterDocId="FF994E5CFFF5B10C4D33A81EFFA1FF95" masterDocTitle="Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico" masterLastPageNumber="581" masterPageNumber="571" pageNumber="577" updateTime="1713383471488" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="5856A648B11472A867F983826DC10AD3">Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico</mods:title>
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<mods:namePart id="19B3C2D509B1BAD03309FFD7572D8243">Kasparyan, Dmitry R.</mods:namePart>
<mods:affiliation id="B68C439F73704B23481B48EA0C448B97">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
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<mods:namePart id="41698493A3CFE5E0A4DA916BC87ADFA6">Khalaim, Andrey I.</mods:namePart>
<mods:nameIdentifier id="CEE3D8A79F0123F7E9B33E2CF78E5FDB" type="ORCID">0000-0003-1802-2649</mods:nameIdentifier>
<mods:affiliation id="8B6693BF9857C1CB0F5E0C678303E007">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
<mods:nameIdentifier id="22A3ADE398ACE07FA32FC2907BAB4A51" type="email">akhalaim@gmail.com</mods:nameIdentifier>
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<mods:date id="802B0B691966463B0F510BD6409989CF">2024</mods:date>
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<treatment id="03A03624FFF3B1044DA4ACEAFE9FF928" ID-DOI="http://doi.org/10.5281/zenodo.10985673" ID-Zenodo-Dep="10985673" LSID="urn:lsid:plazi:treatment:03A03624FFF3B1044DA4ACEAFE9FF928" httpUri="http://treatment.plazi.org/id/03A03624FFF3B1044DA4ACEAFE9FF928" lastPageId="8" lastPageNumber="579" pageId="6" pageNumber="577">
<subSubSection id="C313D4B9FFF3B10A4DA4ACEAFCF1FA9B" box="[151,848,1268,1295]" pageId="6" pageNumber="577" type="nomenclature">
<paragraph id="8BB68732FFF3B10A4DA4ACEAFCF1FA9B" blockId="6.[151,848,1268,1330]" box="[151,848,1268,1295]" pageId="6" pageNumber="577">
<heading id="D0FE305EFFF3B10A4DA4ACEAFCF1FA9B" bold="true" box="[151,848,1268,1295]" fontSize="11" level="1" pageId="6" pageNumber="577" reason="1">
<emphasis id="B97D5B20FFF3B10A4DA4ACEAFCF1FA9B" bold="true" box="[151,848,1268,1295]" pageId="6" pageNumber="577">
2.
<taxonomicName id="4C09FCB1FFF3B10A4D86ACEAFD44FA9B" authority="Kasparyan &amp; Khalaim" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[181,741,1268,1295]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="aterrima" status="sp. nov.">
<emphasis id="B97D5B20FFF3B10A4D86ACEAFE1CFA9B" bold="true" box="[181,445,1268,1295]" italics="true" pageId="6" pageNumber="577">Hylophasma aterrima</emphasis>
Kasparyan &amp; Khalaim
</taxonomicName>
,
<taxonomicNameLabel id="A24EE65BFFF3B10A4FC1ACEAFCF1FA9B" box="[754,848,1268,1294]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C313D4B9FFF3B10A4DA4AD06FE86FAA7" box="[151,295,1304,1330]" pageId="6" pageNumber="577" type="description">
<paragraph id="8BB68732FFF3B10A4DA4AD06FE86FAA7" blockId="6.[151,848,1268,1330]" box="[151,295,1304,1330]" pageId="6" pageNumber="577">
(
<figureCitation id="13329BB7FFF3B10A4DACAD06FEBEFAA7" box="[159,287,1304,1330]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="6" pageNumber="577">Figs 1217</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFF3B10A4DA4AD7EFE8EF99E" pageId="6" pageNumber="577" type="materials_examined">
<paragraph id="8BB68732FFF3B10A4DA4AD7EFF4AFA57" blockId="6.[151,1437,1376,1906]" pageId="6" pageNumber="577">
<emphasis id="B97D5B20FFF3B10A4DA4AD7EFE28FAEF" bold="true" box="[151,393,1376,1402]" pageId="6" pageNumber="577">Material examined.</emphasis>
<materialsCitation id="3B618D6FFFF3B10A4CABAD7EFF4AFA57" collectingDate="2018-03" collectionCode="UNAM, USBI" collectorName="de Informacion &amp; M. Lopez-Ortega." country="Mexico" county="State" elevation="1279" latitude="19.5" location="Unidad de Servicios Bibliotecarios" longLatPrecision="1271" longitude="-96.916664" municipality="Xalapa" pageId="6" pageNumber="577" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<emphasis id="B97D5B20FFF3B10A4CABAD7EFDA4FAEF" bold="true" box="[408,517,1376,1402]" pageId="6" pageNumber="577">
<typeStatus id="54B23990FFF3B10A4CABAD7EFDA4FAEF" box="[408,517,1376,1402]" pageId="6" pageNumber="577" type="holotype">Holotype</typeStatus>
</emphasis>
female (
<collectionCode id="ED181FF7FFF3B10A4F4BAD7EFD72FAEF" box="[632,723,1376,1402]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="6" pageNumber="577">UNAM</collectionCode>
),
<collectingCountry id="F31EC7A2FFF3B10A4FDCAD7EFCE6FAEF" box="[751,839,1376,1402]" name="Mexico" pageId="6" pageNumber="577">Mexico</collectingCountry>
,
<collectingCounty id="62D7FFBEFFF3B10A4E68AD7EFC32FAEF" box="[859,915,1376,1402]" pageId="6" pageNumber="577">State</collectingCounty>
of
<collectingRegion id="49CD49D0FFF3B10A4EFAAD7FFB91FAEE" box="[969,1072,1377,1403]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="577">Veracruz</collectingRegion>
,
<collectingMunicipality id="6BD21D48FFF3B10A4970AD7EFB34FAEF" box="[1091,1173,1376,1402]" pageId="6" pageNumber="577">Xalapa</collectingMunicipality>
,
<location id="8ED6D1E9FFF3B10A499BAD7EFE95FA0B" LSID="urn:lsid:plazi:treatment:03A03624FFF3B1044DA4ACEAFE9FF928:8ED6D1E9FFF3B10A499BAD7EFE95FA0B" country="Mexico" county="State" latitude="19.5" longLatPrecision="1271" longitude="-96.916664" municipality="Xalapa" name="Unidad de Servicios Bibliotecarios" pageId="6" pageNumber="577" stateProvince="Veracruz de Ignacio de la Llave">Unidad de Servicios Bibliotecarios</location>
y
<collectorName id="26FCE2E4FFF3B10A4C60AD9AFDA1FA0B" box="[339,512,1412,1438]" pageId="6" pageNumber="577">de Información</collectorName>
(
<collectionCode id="ED181FF7FFF3B10A4F22AD9AFDF7FA0B" box="[529,598,1412,1438]" pageId="6" pageNumber="577">USBI</collectionCode>
),
<collectingMethod id="5248FF25FFF3B10A4F58AD9AFD5AFA0A" box="[619,763,1412,1439]" pageId="6" pageNumber="577">Malaise trap</collectingMethod>
,
<geoCoordinate id="EE3DE1F5FFF3B10A4E3BAD9AFCCAFA0B" box="[776,875,1412,1438]" degrees="19" direction="north" minutes="30" orientation="latitude" pageId="6" pageNumber="577" precision="925" value="19.5">19°30N</geoCoordinate>
,
<geoCoordinate id="EE3DE1F5FFF3B10A4E4BAD9AFC7EFA0A" box="[888,991,1412,1439]" degrees="96" direction="west" minutes="55" orientation="longitude" pageId="6" pageNumber="577" precision="925" value="-96.916664">96°55W</geoCoordinate>
,
<quantity id="4CF12AD7FFF3B10A4EDFAD9AFBE5FA0A" box="[1004,1092,1412,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.279" pageId="6" pageNumber="577" unit="m" value="1279.0">
<elevation id="00246001FFF3B10A4EDFAD9AFBE5FA0A" box="[1004,1092,1412,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.279" pageId="6" pageNumber="577" unit="m" value="1279.0">1279 m</elevation>
</quantity>
,
<date id="FFB7A1F2FFF3B10A4961AD9AFB7FFA0B" box="[1106,1246,1412,1438]" pageId="6" pageNumber="577" value="2018-03">
<collectingDate id="EFF3581AFFF3B10A4961AD9AFB7FFA0B" box="[1106,1246,1412,1438]" pageId="6" pageNumber="577" value="2018-03">March 2018</collectingDate>
</date>
, coll.
<collectorName id="26FCE2E4FFF3B10A4816AD9BFF4AFA57" pageId="6" pageNumber="577">M. López-Ortega.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BB68732FFF3B10A4DF4ADD2FE8EF99E" blockId="6.[151,1437,1376,1906]" pageId="6" pageNumber="577">
<materialsCitation id="3B618D6FFFF3B10A4DF4ADD2FDDAFA73" box="[199,635,1484,1512]" collectionCode="UNAM" country="Mexico" location="MEXICO" pageId="6" pageNumber="577" specimenCount="1" typeStatus="paratype">
<emphasis id="B97D5B20FFF3B10A4DF4ADD2FE71FA73" bold="true" box="[199,464,1484,1510]" pageId="6" pageNumber="577">
<typeStatus id="54B23990FFF3B10A4DF4ADD2FEE0FA73" box="[199,321,1484,1510]" pageId="6" pageNumber="577" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F31EC7A2FFF3B10A4C63ADD2FE6AFA73" box="[336,459,1484,1510]" name="Mexico" pageId="6" pageNumber="577">MEXICO</collectingCountry>
:
</emphasis>
5 ♂ (
<collectionCode id="ED181FF7FFF3B10A4F29ADD2FDD2FA73" box="[538,627,1484,1510]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="6" pageNumber="577">UNAM</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B618D6FFFF3B10A4FB6ADD3FB73FA73" box="[645,1234,1484,1512]" collectingDate="2016-04-13" collectionCode="ZISP" collectorName="de Informacion &amp; M. Lopez-Ortega." country="Mexico" county="State" elevation="1279" latitude="19.5" location="Unidad de Servicios Bibliotecarios" longLatPrecision="1271" longitude="-96.916664" municipality="Xalapa" pageId="6" pageNumber="577" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
same data as holotype, but
<date id="FFB7A1F2FFF3B10A4EF5ADD2FB9EFA73" box="[966,1087,1484,1510]" pageId="6" pageNumber="577" value="2016-04-13">
<collectingDate id="EFF3581AFFF3B10A4EF5ADD2FB9EFA73" box="[966,1087,1484,1510]" pageId="6" pageNumber="577" value="2016-04-13">13.iv.2016</collectingDate>
</date>
. 1 ♂ (
<collectionCode id="ED181FF7FFF3B10A49BEADD2FB6BFA73" box="[1165,1226,1484,1510]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="577">ZISP</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B618D6FFFF3B10A49EFADD3FE8AF99E" collectingDate="2015-08-07" collectionCode="ZISP" collectorName="de Informacion &amp; M. Lopez-Ortega." country="Mexico" county="State" elevation="1279" latitude="19.5" location="Unidad de Servicios Bibliotecarios" longLatPrecision="1271" longitude="-96.916664" municipality="Xalapa" pageId="6" pageNumber="577" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
same data, but
<date id="FFB7A1F2FFF3B10A48BDADD3FE8AF99E" pageId="6" pageNumber="577" value="2015-08-07">
<collectingDate id="EFF3581AFFF3B10A48BDADD3FE8AF99E" pageId="6" pageNumber="577" value="2015-08-07">7 August 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFF3B1044DF4AE0AFBA0FAA4" lastPageId="8" lastPageNumber="579" pageId="6" pageNumber="577" type="description">
<paragraph id="8BB68732FFF3B10A4DF4AE0AFC72F9BA" blockId="6.[151,1437,1376,1906]" box="[199,979,1556,1583]" pageId="6" pageNumber="577">
<emphasis id="B97D5B20FFF3B10A4DF4AE0AFEF8F9BB" bold="true" box="[199,345,1556,1582]" pageId="6" pageNumber="577">Description.</emphasis>
<emphasis id="B97D5B20FFF3B10A4C53AE0AFE15F9BB" box="[352,436,1556,1582]" italics="true" pageId="6" pageNumber="577">Female</emphasis>
. Body length
<quantity id="4CF12AD7FFF3B10A4F61AE0AFD0BF9BA" box="[594,682,1556,1583]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="577" unit="mm" value="4.8">4.8 mm</quantity>
, fore wing length
<quantity id="4CF12AD7FFF3B10A4E45AE0AFC6EF9BA" box="[886,975,1556,1583]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="577" unit="mm" value="4.1">4.1 mm</quantity>
.
</paragraph>
<paragraph id="8BB68732FFF3B10A4DF4AE26FB93F8E7" blockId="6.[151,1437,1376,1906]" pageId="6" pageNumber="577">
Antenna with 24 flagellomeres; flagellum distinctly thickened in apical 0.4, about ten subapical flagellomeres 1.51.8× as broad as basal flagellomeres (
<figureCitation id="13329BB7FFF3B10A4FB0AE42FD7BF9E3" box="[643,730,1628,1654]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="6" pageNumber="577">Fig. 12</figureCitation>
). Head polished, with sparse fine setiferous punctures; frons highly polished, with very fine and shallow granulation on eye orbits (
<figureCitation id="13329BB7FFF3B10A4EFFAE9EFB83F90F" box="[972,1058,1664,1690]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="6" pageNumber="577">Fig. 14</figureCitation>
); median part of frons, posterior half of vertex and gena almost impunctate (
<figureCitation id="13329BB7FFF3B10A4FA6AEBAFCBCF92B" box="[661,797,1700,1726]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="6" pageNumber="577">Figs 13, 14</figureCitation>
). Clypeus evenly convex in profile, with lower margin evenly truncated, laterally with small sharp lobe (
<figureCitation id="13329BB7FFF3B10A4FF0AED6FCB6F977" box="[707,791,1736,1762]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="6" pageNumber="577">Fig. 13</figureCitation>
). Lower tooth of mandible slightly shorter than the upper tooth; lower margin of mandible with distinct sharp flange extending to base of teeth (
<figureCitation id="13329BB7FFF3B10A4951AEF2FB17F893" box="[1122,1206,1772,1798]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="6" pageNumber="577">Fig. 13</figureCitation>
). Malar space finely granulate, about 0.6× as long as basal mandibular width. Lower part of gena convex, without concavity before base of mandible (
<figureCitation id="13329BB7FFF3B10A4C00AF2AFE28F8DB" box="[307,393,1844,1870]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="6" pageNumber="577">Fig. 13</figureCitation>
). Occipital carina joining to hypostomal carina very close to base of mandible; portion of hypostomal carina between this junction and mandible shorter than width of palpi.
</paragraph>
<caption id="DF76D7BAFFF2B10B4DA4AF00FF4CF8E8" ID-DOI="http://doi.org/10.5281/zenodo.10985643" ID-Zenodo-Dep="10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="7" pageNumber="578" startId="7.[151,264,1822,1846]" targetBox="[151,1435,181,1798]" targetPageId="7" targetType="figure">
<paragraph id="8BB68732FFF2B10B4DA4AF00FF4CF8E8" blockId="7.[151,1436,1822,1917]" pageId="7" pageNumber="578">
<emphasis id="B97D5B20FFF2B10B4DA4AF00FEF6F8A3" bold="true" box="[151,343,1822,1846]" pageId="7" pageNumber="578">FIGURES 1217.</emphasis>
<taxonomicName id="4C09FCB1FFF2B10B4C53AF00FDE3F8A3" authority="Kasparyan &amp; Khalaim, 2024" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[352,578,1822,1846]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="aterrima" status="sp. nov.">
<emphasis id="B97D5B20FFF2B10B4C53AF00FDE3F8A3" box="[352,578,1822,1846]" italics="true" pageId="7" pageNumber="578">Hylophasma aterrima</emphasis>
</taxonomicName>
<emphasis id="B97D5B20FFF2B10B4F78AF00FD3EF8A3" bold="true" box="[587,671,1822,1846]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A24EE65BFFF2B10B4F78AF00FD3EF8A3" box="[587,671,1822,1846]" pageId="7" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀, holotype (1215) and ♂, paratype (16, 17):
<emphasis id="B97D5B20FFF2B10B4992AF00FB42F8A3" bold="true" box="[1185,1251,1822,1846]" pageId="7" pageNumber="578">12, 16</emphasis>
—habitus, lateral;
<emphasis id="B97D5B20FFF2B10B4DA4AF5CFF11F8CF" bold="true" box="[151,176,1858,1882]" pageId="7" pageNumber="578">13</emphasis>
—head, front;
<emphasis id="B97D5B20FFF2B10B4C7BAF5CFEC0F8CF" bold="true" box="[328,353,1858,1882]" pageId="7" pageNumber="578">14</emphasis>
—head and anterior part of mesosoma, dorsal;
<emphasis id="B97D5B20FFF2B10B4E75AF5CFCFEF8CF" bold="true" box="[838,863,1858,1882]" pageId="7" pageNumber="578">15</emphasis>
—head, mesosoma and base of metasoma, lateral;
<emphasis id="B97D5B20FFF2B10B4859AF5CFA22F8CF" bold="true" box="[1386,1411,1858,1882]" pageId="7" pageNumber="578">17</emphasis>
— antenna.
</paragraph>
</caption>
<paragraph id="8BB68732FFFDB1044DF4A889FDD1FE44" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">
Mesosoma polished with fine setiferous punctures (
<figureCitation id="13329BB7FFFDB1044E34A889FCFBFF24" box="[775,858,150,177]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 15</figureCitation>
); pronotum and median part of mesopleuron almost impunctate; puncture on anterior part of mesoscutum denser (
<figureCitation id="13329BB7FFFDB1044E5EA8A5FC64FF40" box="[877,965,187,213]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 14</figureCitation>
); notaulus deep and sharp, extending to posterior 0.7 of mesoscutum (
<figureCitation id="13329BB7FFFDB1044CEEA8C1FD8FFF6C" box="[477,558,223,249]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 14</figureCitation>
). Scuto-scutellar groove with 68 short transverse rugae (
<figureCitation id="13329BB7FFFDB1044996A8C1FB57FF6C" box="[1189,1270,223,249]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 14</figureCitation>
). Mesopleuron between subtegular ridge and mesopleural fovea somewhat impressed and with five short carinae. Sternaulus short and sharp, almost reaching midlength of mesopleuron (
<figureCitation id="13329BB7FFFDB1044E2AA939FCCEFED4" box="[793,879,294,321]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 15</figureCitation>
). Metapleuron with fine and short setae (denser than on mesopleuron). Juxtacoxal carina absent. Propodeum predominantly polished, pleural and all apical areas densely and finely punctato-rugose; lateromedian and lateral longitudinal carinae of propodeum absent; pleural carina behind anterior transverse carina obliterated; posterior transverse carina almost straight medially, without projections (apophyses or crests) laterally.
</paragraph>
<paragraph id="8BB68732FFFDB1044DF4A9C5FECDFDF4" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">Fore wing with areolet fore wing of medium size, as high as portion of vein 2m-cu above bulla; vein 1cu-a (nervulus) slightly reclivous; postnervulus intercepted at upper 0.35. Hind wing with nervellus intercepted at lower 0.4; posterior section of AA (brachiella) entirely absent. Proportion of length of tarsomeres 15 of hind tarsus 5.5: 2.1: 1.5: 0.8: 1.4.</paragraph>
<paragraph id="8BB68732FFFDB1044DF4AA75FF0DFD64" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">
First metasomal segment slender, about 4.0× as long as wide posteriorly, without longitudinal carinae, ridges, basolateral tooth or flange. Second tergite strongly widened towards apex, 2.5× wider posteriorly than at base. Ovipositor sheath about 1.1× as long as hind tibia (
<figureCitation id="13329BB7FFFDB1044FF5AAADFCB9FD58" box="[710,792,691,717]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 12</figureCitation>
); upper valve behind nodus 1.8× as long as hind tarsomere 2.
</paragraph>
<paragraph id="8BB68732FFFDB1044DF4AAE5FCC4FBA0" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">
Body predominantly black (
<figureCitation id="13329BB7FFFDB1044F34AAE5FDFDFC80" box="[519,604,763,789]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 12</figureCitation>
). Antenna with scape and pedicel blackish dorsally and yellowish brown ventrally; flagellum black, flagellomeres 711 white dorsally. Head black; lower half of clypeus and mandibles (except for teeth) reddish brown; palpi yellowish white. Mesosoma black; propleuron, entire anterior margin and extreme dorso-posterior corner of pronotum, upper part of mesepimeron and posteroventral part of mesopleuron yellowish to pale reddish (
<figureCitation id="13329BB7FFFDB1044CF7AB95FDB8FC30" box="[452,537,906,933]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 15</figureCitation>
); dorsal mark on collar and tegula whitish. Metasoma brownish black to black, tergites
<quantity id="4CF12AD7FFFDB1044DC2ABB1FE99FC5C" box="[241,312,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.524" metricValueMin="0.508" pageId="8" pageNumber="579" unit="in" value="4.0" valueMax="6.0" valueMin="2.0">26 in</quantity>
posterior 0.20.3 brown (brown marks wider medially and narrower laterally). Legs predominantly reddish yellow; fore and mid coxae and fore trochanter and trochantellus white; hind coxa with brown mark dorsally, hind trochanter and trochantellus marked with brown, all femora narrowly brownish dorsally, hind tibia entirely brown, hind and partly mid tarsi darkened with brown (
<figureCitation id="13329BB7FFFDB1044E37AC05FCF9FBA0" box="[772,856,1051,1077]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BB68732FFFDB1044DF4AC21FBA0FAA4" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">
<emphasis id="B97D5B20FFFDB1044DF4AC21FEA0FBCC" box="[199,257,1087,1113]" italics="true" pageId="8" pageNumber="579">Male</emphasis>
(
<figureCitation id="13329BB7FFFDB1044C21AC21FEC9FBCC" box="[274,360,1087,1113]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 16</figureCitation>
). Fore wing length
<quantity id="4CF12AD7FFFDB1044F78AC20FD76FBCC" box="[587,727,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="8" pageNumber="579" unit="mm" value="4.6" valueMax="5.1" valueMin="4.1">4.15.1 mm</quantity>
. Antenna with 2829 flagellomeres (
<figureCitation id="13329BB7FFFDB104494DAC21FB73FBCC" box="[1150,1234,1087,1113]" captionStart="FIGURES 1217" captionStartId="7.[151,264,1822,1846]" captionTargetBox="[151,1435,181,1798]" captionTargetId="figure-19@7.[151,1435,181,1798]" captionTargetPageId="7" captionText="FIGURES 1217. Hylophasma aterrima sp. nov., ♀, holotype (1215) and ♂, paratype (16, 17): 12, 16—habitus, lateral; 13—head, front; 14—head and anterior part of mesosoma, dorsal; 15—head, mesosoma and base of metasoma, lateral; 17— antenna." figureDoi="http://doi.org/10.5281/zenodo.10985643" httpUri="https://zenodo.org/record/10985643/files/figure.png" pageId="8" pageNumber="579">Fig. 17</figureCitation>
), flagellum about 1.2× as long as fore wing; flagellomeres 1214 with linear tyloid along their entire length. First tergite very long and slender, 5.6× as long as wide posteriorly. Paler than female; scape and pedicel ventrally white; flagellomeres 914 almost completely white, slightly brownish ventrally; clypeus, mandible (except teeth), propleuron, anterior margin of pronotum, mid trochanter and trochantellus, fore and mid femora in ventro-basal halves, and hind coxa ventrally white; mesepimeron almost entirely dull whitish; parameres whitish. Posterior pale markings on tergites 25 medially expanded towards centre of tergite. Otherwise similar to female.
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFFDB1044DF4AD25FF74FAEC" pageId="8" pageNumber="579" type="etymology">
<paragraph id="8BB68732FFFDB1044DF4AD25FF74FAEC" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">
<emphasis id="B97D5B20FFFDB1044DF4AD25FEEFFAC0" bold="true" box="[199,334,1339,1365]" pageId="8" pageNumber="579">Etymology.</emphasis>
The new species is named from the Latin
<emphasis id="B97D5B20FFFDB1044E75AD25FC12FAC1" box="[838,947,1339,1364]" italics="true" pageId="8" pageNumber="579">aterrimus</emphasis>
(black, dark) after its predominantly dark body.
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFFDB1044DF4AD9CFE9FF928" pageId="8" pageNumber="579" type="distribution">
<paragraph id="8BB68732FFFDB1044DF4AD9CFDE3FA08" blockId="8.[151,1437,150,1725]" box="[199,578,1410,1437]" pageId="8" pageNumber="579">
<emphasis id="B97D5B20FFFDB1044DF4AD9CFEC0FA09" bold="true" box="[199,353,1410,1436]" pageId="8" pageNumber="579">Distribution.</emphasis>
<collectingCountry id="F31EC7A2FFFDB1044C54AD9DFE1CFA08" box="[359,445,1411,1437]" name="Mexico" pageId="8" pageNumber="579">Mexico</collectingCountry>
(
<collectingRegion id="49CD49D0FFFDB1044CFFAD9DFD96FA08" box="[460,567,1411,1437]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="8" pageNumber="579">Veracruz</collectingRegion>
).
</paragraph>
<paragraph id="8BB68732FFFDB1044DF4ADB8FD78F9C4" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">
<emphasis id="B97D5B20FFFDB1044DF4ADB8FEC3FA55" bold="true" box="[199,354,1446,1472]" pageId="8" pageNumber="579">Comparison.</emphasis>
The new species belongs to the
<taxonomicName id="4C09FCB1FFFDB1044FD0ADB9FC8FFA54" authorityName="Townes" authorityYear="1962" box="[739,814,1447,1473]" class="Insecta" family="Ichneumonidae" genus="Hemisphragia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B97D5B20FFFDB1044FD0ADB9FC8FFA54" box="[739,814,1447,1473]" italics="true" pageId="8" pageNumber="579">debilis</emphasis>
</taxonomicName>
species-group as it has occipital carina complete, not interrupted ventrally. Within this species-group it resembles
<taxonomicName id="4C09FCB1FFFDB1044E51ADD5FC53FA71" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[866,1010,1483,1508]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B97D5B20FFFDB1044E51ADD5FC53FA71" box="[866,1010,1483,1508]" italics="true" pageId="8" pageNumber="579">H. nigriceps</emphasis>
</taxonomicName>
but differs from this species by its predominantly black mesosoma with scarce pale markings, polished frons (distinctly granulate in
<taxonomicName id="4C09FCB1FFFDB10449CCADF1FA2DF99D" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[1279,1420,1519,1544]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B97D5B20FFFDB10449CCADF1FA2DF99D" box="[1279,1420,1519,1544]" italics="true" pageId="8" pageNumber="579">H. nigriceps</emphasis>
</taxonomicName>
), shorter sternaulus, metapleuron without juxtacoxal carina (present in
<taxonomicName id="4C09FCB1FFFDB1044EAAAE0DFB83F9B9" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[921,1058,1555,1580]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B97D5B20FFFDB1044EAAAE0DFB83F9B9" box="[921,1058,1555,1580]" italics="true" pageId="8" pageNumber="579">H. nigriceps</emphasis>
</taxonomicName>
), and hind wing without posterior section of AA (brachiella) (present in
<taxonomicName id="4C09FCB1FFFDB1044F0DAE29FD69F9C5" authorityName="Kasparyan &amp; Ruiz-Cancino" authorityYear="2005" box="[574,712,1591,1616]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B97D5B20FFFDB1044F0DAE29FD69F9C5" box="[574,712,1591,1616]" italics="true" pageId="8" pageNumber="579">H. nigriceps</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB68732FFFDB1044DF4AE45FE9FF928" blockId="8.[151,1437,150,1725]" pageId="8" pageNumber="579">
<taxonomicName id="4C09FCB1FFFDB1044DF4AE45FE61F9E1" authority="Kasparyan &amp; Khalaim, 2024" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[199,448,1627,1653]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="aterrima" status="sp. nov.">
<emphasis id="B97D5B20FFFDB1044DF4AE45FE61F9E1" box="[199,448,1627,1653]" italics="true" pageId="8" pageNumber="579">Hylophasma aterrima</emphasis>
</taxonomicName>
<emphasis id="B97D5B20FFFDB1044CFAAE45FD85F9E0" bold="true" box="[457,548,1627,1653]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A24EE65BFFFDB1044CFAAE45FD85F9E0" box="[457,548,1627,1653]" pageId="8" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs from the Nearctic
<taxonomicName id="4C09FCB1FFFDB1044EBBAE45FBEFF9E0" authority="Townes" authorityName="Townes" box="[904,1102,1627,1653]" class="Insecta" family="Ichneumonidae" genus="Hylophasma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="debils">
<emphasis id="B97D5B20FFFDB1044EBBAE45FC4EF9E0" box="[904,1007,1627,1653]" italics="true" pageId="8" pageNumber="579">H. debils</emphasis>
Townes
</taxonomicName>
by the ovipositor with upper valve behind nodus 1.8× as long as hind tarsomere 2, while in
<taxonomicName id="4C09FCB1FFFDB1044E5AAE61FC79F90C" authorityName="Townes" authorityYear="1962" box="[873,984,1663,1689]" class="Insecta" family="Ichneumonidae" genus="Hemisphragia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="B97D5B20FFFDB1044E5AAE61FC79F90C" box="[873,984,1663,1689]" italics="true" pageId="8" pageNumber="579">H. debilis</emphasis>
</taxonomicName>
this proportion is 1.2× (see Fig.
<quantity id="4CF12AD7FFFDB1044867AE61FA3DF90C" box="[1364,1436,1663,1689]" metricMagnitude="0" metricUnit="m" metricValue="6.0198" pageId="8" pageNumber="579" unit="in" value="237.0">237 in</quantity>
<bibRefCitation id="EF98FAC3FFFDB1044DA4AEBDFE90F928" author="Townes, H. K." box="[151,305,1699,1725]" pageId="8" pageNumber="579" pagination="1 - 537" refId="ref7309" refString="Townes, H. K. (1970) The genera of Ichneumonidae. Part 2. Memoirs of the American Entomological Institute, 12, 1 - 537." type="journal article" year="1970">Townes 1970</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>