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<document ID-DOI="10.11646/zootaxa.5263.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804268" ID-ZooBank="20CB3CA0-BEF9-474C-8931-6A7948B9CA61" approvalRequired="5" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="4" checkinTime="1680760231104" checkinUser="plazi" docAuthor="Řezáč, Milan, Cardoso, Pedro &amp; Řezáčová, Veronika" docDate="2023" docId="039CE028E87DFFCFFF12FEEAFC346328" docLanguage="en" docName="zootaxa.5263.3.2.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpactea korenkoi Rezac 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="348" masterDocId="FFA59850E870FFC2FF85FFA1FFD2667D" masterDocTitle="Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal" masterLastPageNumber="364" masterPageNumber="335" pageNumber="348" updateTime="1680760785065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Řezáč, Milan</mods:namePart>
<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cardoso, Pedro</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8119-9960</mods:nameIdentifier>
<mods:affiliation>Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History Luomus, University of Helsinki, P. O. Box 17, FI- 00014 Helsinki, Finland. pedro. cardoso @ helsinki. fi; https: // orcid. org / 0000 - 0001 - 8119 - 9960</mods:affiliation>
<mods:nameIdentifier type="email">pedro.cardoso@helsinki.fi</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Řezáčová, Veronika</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1749-0355</mods:nameIdentifier>
<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia &amp; strver @ seznam. cz; https: // orcid. org / 0000 - 0002 - 1749 - 0355</mods:affiliation>
<mods:nameIdentifier type="email">strver@seznam.cz</mods:nameIdentifier>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-06</mods:number>
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<mods:detail type="volume">
<mods:number>5263</mods:number>
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<mods:detail type="issue">
<mods:number>3</mods:number>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.3.2</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5263.3.2</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="ZooBank">20CB3CA0-BEF9-474C-8931-6A7948B9CA61</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7804300" ID-Zenodo-Dep="7804300" LSID="urn:lsid:plazi:treatment:039CE028E87DFFCFFF12FEEAFC346328" httpUri="http://treatment.plazi.org/id/039CE028E87DFFCFFF12FEEAFC346328" lastPageNumber="348" pageId="13" pageNumber="348">
<subSubSection box="[151,576,330,360]" pageId="13" pageNumber="348" type="nomenclature">
<paragraph blockId="13.[151,576,330,393]" box="[151,576,330,360]" pageId="13" pageNumber="348">
<heading bold="true" box="[151,576,330,360]" fontSize="11" level="1" pageId="13" pageNumber="348" reason="1">
<emphasis bold="true" box="[151,576,330,360]" pageId="13" pageNumber="348">
<taxonomicName authority="Rezac" authorityName="Rezac" authorityYear="2023" box="[151,476,330,360]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="korenkoi" status="sp. nov.">
<emphasis bold="true" box="[151,394,330,357]" italics="true" pageId="13" pageNumber="348">Harpactea korenkoi</emphasis>
Řezáč
</taxonomicName>
<taxonomicNameLabel box="[483,576,331,357]" pageId="13" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,479,367,393]" pageId="13" pageNumber="348" type="description">
<paragraph blockId="13.[151,576,330,393]" box="[151,479,367,393]" pageId="13" pageNumber="348">
(
<figureCitation box="[159,251,367,393]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="13" pageNumber="348">Figs 1H</figureCitation>
,
<figureCitation box="[264,300,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[250,1344,181,1799]" captionTargetId="figure-24@4.[244,1344,181,1799]" captionTargetPageId="4" captionText="FIGURE 2. Portuguese Harpactea spiders. Right male chelicera.Above anterior (G, J, L, M) or posterior view (A, B, C, D, E, F, H, I, K, N); below median view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804274" httpUri="https://zenodo.org/record/7804274/files/figure.png" pageId="13" pageNumber="348">2H</figureCitation>
,
<figureCitation box="[312,348,367,393]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="13" pageNumber="348">3H</figureCitation>
,
<figureCitation box="[360,397,367,393]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="13" pageNumber="348">4H</figureCitation>
,
<figureCitation box="[409,444,367,393]" captionStart="FIGURE 8" captionStartId="16.[151,250,1822,1846]" captionTargetBox="[169,1418,182,1795]" captionTargetId="figure-24@16.[169,1418,181,1798]" captionTargetPageId="16" captionText="FIGURE 8. Portuguese Harpactea spiders. Vulva. A. H. magnibulbi. Portugal: Fóia; B. H. korenkoi sp. nov. Portugal: Carrapateira; C. H. crespoi sp. nov. Portugal: Sobral da Adiça; D. H. algarvensis. Portugal: Barranco do Velho; E. H. tavirensis. Portugal: Barranco do Velho. Views: dor.—dorsal, fro.—anterior, hin.—posterior, ven.—ventral. Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue." figureDoi="http://doi.org/10.5281/zenodo.7804292" httpUri="https://zenodo.org/record/7804292/files/figure.png" pageId="13" pageNumber="348">8B</figureCitation>
,
<figureCitation box="[455,470,367,393]" captionStart="FIGURE 9" captionStartId="22.[151,250,1892,1916]" captionTargetBox="[279,1342,195,1818]" captionTargetId="figure-24@22.[231,1356,181,1869]" captionTargetPageId="22" captionText="FIGURE 9. Known distribution of Portuguese Harpactea species. In each map the first species is represented by a transparent circle and the second species by a black diamond:A) H. adicensis &amp; H. algarvensis; B) H. crespoi &amp; H. dolanskyi; C) H. fageli &amp; H. gaditana; D) H. henriquesi &amp; H. hombergii; E) H. korenkoi &amp; H. krejcii; F) H. magnibulbi &amp; H. minoccii; G) H. pekari &amp; H. proxima; H) H. stalitoides &amp; H. subiasi; I) H. tavirensis." figureDoi="http://doi.org/10.5281/zenodo.7804296" httpUri="https://zenodo.org/record/7804296/files/figure.png" pageId="13" pageNumber="348">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="348" type="materials_examined">
<paragraph blockId="13.[151,1437,438,1365]" pageId="13" pageNumber="348">
<emphasis bold="true" box="[151,325,438,465]" pageId="13" pageNumber="348">Type material.</emphasis>
<materialsCitation collectingDate="2013-04-02" collectorName="M. Rezac &amp; National Natural History Museum" latitude="37.184" location="Prague" longLatPrecision="70" longitude="-8.895" pageId="13" pageNumber="348" specimenCount="1" stateProvince="Praha" typeStatus="holotype">
<emphasis box="[332,442,439,465]" italics="true" pageId="13" pageNumber="348">
<typeStatus box="[332,438,439,465]" pageId="13" pageNumber="348">Holotype</typeStatus>
.
</emphasis>
♁, Aljezur, Carrapateira,
<geoCoordinate box="[732,812,439,465]" degrees="37.184" direction="north" orientation="latitude" pageId="13" pageNumber="348" precision="55" value="37.184">37.184</geoCoordinate>
,
<geoCoordinate box="[823,898,438,465]" degrees="8.895" direction="west" orientation="longitude" pageId="13" pageNumber="348" precision="55" value="-8.895">-8.895</geoCoordinate>
,
<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[910,1115,439,465]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="348" phylum="Tracheophyta" rank="species" species="coccifera">
<emphasis box="[910,1115,439,465]" italics="true" pageId="13" pageNumber="348">Quercus coccifera</emphasis>
</taxonomicName>
forest,
<date box="[1200,1345,439,465]" pageId="13" pageNumber="348" value="2013-04-02">
<collectingDate box="[1200,1345,439,465]" pageId="13" pageNumber="348" value="2013-04-02">2 April 2013</collectingDate>
</date>
, leg.
<collectorName pageId="13" pageNumber="348">M. Řezáč</collectorName>
, coll.
<collectorName box="[290,676,475,501]" pageId="13" pageNumber="348">National Natural History Museum</collectorName>
,
<collectingRegion box="[687,768,475,501]" country="Czech Republic" name="Praha" pageId="13" pageNumber="348">Prague</collectingRegion>
, code P6A 7387.
</materialsCitation>
</paragraph>
<paragraph blockId="13.[151,1437,438,1365]" pageId="13" pageNumber="348">
<materialsCitation collectingDate="2008-04-07" collectingDateMax="2013-04-02" collectingDateMin="2008-04-07" collectorName="M. Rezac &amp; Crop Research Institute" latitude="37.184" location="Prague" longLatPrecision="70" longitude="-8.895" pageId="13" pageNumber="348" specimenCount="22" specimenCount-female="22" stateProvince="Praha" typeStatus="paratype">
<emphasis box="[199,320,511,536]" italics="true" pageId="13" pageNumber="348">
<typeStatus box="[199,316,511,536]" pageId="13" pageNumber="348">Paratypes</typeStatus>
.
</emphasis>
Aljezur, Carrapateira,
<geoCoordinate box="[578,658,511,537]" degrees="37.184" direction="north" orientation="latitude" pageId="13" pageNumber="348" precision="55" value="37.184">37.184</geoCoordinate>
,
<geoCoordinate box="[670,745,510,537]" degrees="8.895" direction="west" orientation="longitude" pageId="13" pageNumber="348" precision="55" value="-8.895">-8.895</geoCoordinate>
,
<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[757,963,511,537]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="348" phylum="Tracheophyta" rank="species" species="coccifera">
<emphasis box="[757,963,511,537]" italics="true" pageId="13" pageNumber="348">Quercus coccifera</emphasis>
</taxonomicName>
forest, 3 ♁♁,
<specimenCount box="[1129,1193,511,537]" pageId="13" pageNumber="348" type="female">7 ♀♀</specimenCount>
,
<date box="[1207,1353,511,537]" pageId="13" pageNumber="348" value="2008-04-07">
<collectingDate box="[1207,1353,511,537]" pageId="13" pageNumber="348" value="2008-04-07">7 April 2008</collectingDate>
</date>
, 7 ♁♁,
<specimenCount box="[151,229,546,573]" pageId="13" pageNumber="348" type="female">15 ♀♀</specimenCount>
,
<date box="[242,387,547,573]" pageId="13" pageNumber="348" value="2013-04-02">
<collectingDate box="[242,387,547,573]" pageId="13" pageNumber="348" value="2013-04-02">2 April 2013</collectingDate>
</date>
, leg.
<collectorName box="[446,556,547,575]" pageId="13" pageNumber="348">M. Řezáč</collectorName>
, coll.
<collectorName box="[623,891,547,573]" pageId="13" pageNumber="348">Crop Research Institute</collectorName>
,
<collectingRegion box="[902,983,547,573]" country="Czech Republic" name="Praha" pageId="13" pageNumber="348">Prague</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="348" type="etymology">
<paragraph blockId="13.[151,1437,438,1365]" pageId="13" pageNumber="348">
<emphasis bold="true" box="[199,334,583,609]" pageId="13" pageNumber="348">Etymology.</emphasis>
Named after the Czechoslovak arachnologist Stanislav Korenko, our friend who helped us to collect material for this study.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="348" type="diagnosis">
<paragraph blockId="13.[151,1437,438,1365]" pageId="13" pageNumber="348">
<emphasis bold="true" box="[199,322,654,680]" pageId="13" pageNumber="348">Diagnosis.</emphasis>
It differs from most Iberian
<taxonomicName box="[661,872,655,681]" pageId="13" pageNumber="348">
<emphasis box="[661,781,655,680]" italics="true" pageId="13" pageNumber="348">Harpactea</emphasis>
species
</taxonomicName>
by almost straight dorsal side of cymbium (
<figureCitation captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="13" pageNumber="348">Fig. 3H</figureCitation>
, like in
<taxonomicName authorityName="Řezáč &amp; Cardoso &amp; Řezáčová" authorityYear="2023" box="[277,440,691,717]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="magnibulbi">
<emphasis box="[277,440,691,717]" italics="true" pageId="13" pageNumber="348">H. magnibulbi</emphasis>
</taxonomicName>
), and relatively short embolus (approximately of the same length as conductor,
<figureCitation box="[1334,1424,691,717]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="13" pageNumber="348">Fig. 4H</figureCitation>
). It resembles
<taxonomicName authorityName="Řezáč &amp; Cardoso &amp; Řezáčová" authorityYear="2023" box="[297,461,727,753]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="magnibulbi">
<emphasis box="[297,461,727,753]" italics="true" pageId="13" pageNumber="348">H. magnibulbi</emphasis>
</taxonomicName>
; males differ by the distance between the anterior cheliceral teeth, that is 1.5× longer than the distance between the teeth in the posterior row (
<figureCitation box="[786,875,763,789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[250,1344,181,1799]" captionTargetId="figure-24@4.[244,1344,181,1799]" captionTargetPageId="4" captionText="FIGURE 2. Portuguese Harpactea spiders. Right male chelicera.Above anterior (G, J, L, M) or posterior view (A, B, C, D, E, F, H, I, K, N); below median view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804274" httpUri="https://zenodo.org/record/7804274/files/figure.png" pageId="13" pageNumber="348">Fig. 2H</figureCitation>
, in
<taxonomicName authorityName="Řezáč &amp; Cardoso &amp; Řezáčová" authorityYear="2023" box="[916,1079,763,789]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="magnibulbi">
<emphasis box="[916,1079,763,789]" italics="true" pageId="13" pageNumber="348">H. magnibulbi</emphasis>
</taxonomicName>
the distance between the dorsal teeth is the same as the distance between the ventral teeth,
<figureCitation box="[826,917,799,825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[250,1344,181,1799]" captionTargetId="figure-24@4.[244,1344,181,1799]" captionTargetPageId="4" captionText="FIGURE 2. Portuguese Harpactea spiders. Right male chelicera.Above anterior (G, J, L, M) or posterior view (A, B, C, D, E, F, H, I, K, N); below median view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804274" httpUri="https://zenodo.org/record/7804274/files/figure.png" pageId="13" pageNumber="348">Fig. 2G</figureCitation>
); the position of the dorsal distal tooth that is between the ventral teeth (in
<taxonomicName authorityName="Řezáč &amp; Cardoso &amp; Řezáčová" authorityYear="2023" box="[487,651,835,861]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="magnibulbi">
<emphasis box="[487,651,835,861]" italics="true" pageId="13" pageNumber="348">H. magnibulbi</emphasis>
</taxonomicName>
it is next to the posterior basal tooth), and by the shape of embolus, which is thinner (
<figureCitation box="[349,439,871,897]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="13" pageNumber="348">Fig. 4H</figureCitation>
), S-shaped on its distal end; females differ by anterior arc with lobes bent backwards on its prolateral edges and absence of posterior diverticle rudiment (
<figureCitation box="[879,966,907,933]" captionStart="FIGURE 8" captionStartId="16.[151,250,1822,1846]" captionTargetBox="[169,1418,182,1795]" captionTargetId="figure-24@16.[169,1418,181,1798]" captionTargetPageId="16" captionText="FIGURE 8. Portuguese Harpactea spiders. Vulva. A. H. magnibulbi. Portugal: Fóia; B. H. korenkoi sp. nov. Portugal: Carrapateira; C. H. crespoi sp. nov. Portugal: Sobral da Adiça; D. H. algarvensis. Portugal: Barranco do Velho; E. H. tavirensis. Portugal: Barranco do Velho. Views: dor.—dorsal, fro.—anterior, hin.—posterior, ven.—ventral. Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue." figureDoi="http://doi.org/10.5281/zenodo.7804292" httpUri="https://zenodo.org/record/7804292/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="348" type="description">
<paragraph blockId="13.[151,1437,438,1365]" pageId="13" pageNumber="348">
<emphasis bold="true" box="[199,345,942,968]" pageId="13" pageNumber="348">Description.</emphasis>
<emphasis box="[354,544,943,969]" italics="true" pageId="13" pageNumber="348">
Male (
<typeStatus box="[430,532,943,969]" pageId="13" pageNumber="348">holotype</typeStatus>
).
</emphasis>
Carapace yellow, matting (
<figureCitation box="[861,953,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="13" pageNumber="348">Fig. 1H</figureCitation>
). Sternum yellow, matting, with long hair on the edges. Chelicerae yellow brown. Legs yellow brown, pedipalps ferruginous. Measurements and spination are shown in the
<tableCitation box="[353,439,1015,1041]" captionStart="TABLE 1" captionStartId="17.[152,244,151,175]" captionText="TABLE 1. Measurements and spination of holotypes of the new Portuguese Harpactea species and Harpactea tavirensis." pageId="13" pageNumber="348">Table 1</tableCitation>
. Relative leg length: IV&gt;I&gt;II&gt;III. Cymbium with very elongated distal part (
<figureCitation box="[1331,1423,1015,1041]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="13" pageNumber="348">Fig. 3H</figureCitation>
), and very concave prolateral side. Tegulum large, longer than wide (
<figureCitation box="[919,1011,1051,1077]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="13" pageNumber="348">Fig. 4H</figureCitation>
). Conductor regularly bent, flattened, tapering. Embolus dark, parallel to conductor but more bent, only slightly longer than conductor, flattened, its end is S-shaped. Opisthosoma cylindrical, whitish.
</paragraph>
<paragraph blockId="13.[151,1437,438,1365]" pageId="13" pageNumber="348">
<emphasis box="[199,291,1159,1185]" italics="true" pageId="13" pageNumber="348">Female.</emphasis>
All somatic characters as described for the male. Vulva is of the
<typeStatus box="[1035,1083,1159,1185]" pageId="13" pageNumber="348">type</typeStatus>
<taxonomicName authorityName="Ferrandez" authorityYear="1990" box="[1091,1220,1159,1185]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="algarvensis">
<emphasis box="[1091,1220,1159,1185]" italics="true" pageId="13" pageNumber="348">algarvensis</emphasis>
</taxonomicName>
(see Materials and Methods) (
<figureCitation box="[274,362,1195,1221]" captionStart="FIGURE 8" captionStartId="16.[151,250,1822,1846]" captionTargetBox="[169,1418,182,1795]" captionTargetId="figure-24@16.[169,1418,181,1798]" captionTargetPageId="16" captionText="FIGURE 8. Portuguese Harpactea spiders. Vulva. A. H. magnibulbi. Portugal: Fóia; B. H. korenkoi sp. nov. Portugal: Carrapateira; C. H. crespoi sp. nov. Portugal: Sobral da Adiça; D. H. algarvensis. Portugal: Barranco do Velho; E. H. tavirensis. Portugal: Barranco do Velho. Views: dor.—dorsal, fro.—anterior, hin.—posterior, ven.—ventral. Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue." figureDoi="http://doi.org/10.5281/zenodo.7804292" httpUri="https://zenodo.org/record/7804292/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph blockId="13.[151,1437,438,1365]" box="[199,893,1230,1257]" pageId="13" pageNumber="348">
<emphasis box="[199,324,1231,1257]" italics="true" pageId="13" pageNumber="348">Variability.</emphasis>
Carapace length
<quantity box="[518,653,1230,1257]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="13" pageNumber="348" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.52.2 mm</quantity>
(both sexes) (N=33).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="348" type="biology_ecology">
<paragraph blockId="13.[151,1437,438,1365]" pageId="13" pageNumber="348">
<emphasis bold="true" box="[199,300,1267,1293]" pageId="13" pageNumber="348">Ecology.</emphasis>
It lives in leaf litter in shaded
<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[628,832,1267,1293]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="348" phylum="Tracheophyta" rank="species" species="coccifera">
<emphasis box="[628,832,1267,1293]" italics="true" pageId="13" pageNumber="348">Quercus coccifera</emphasis>
</taxonomicName>
forests. Females lay eggs in early spring. The presence of independent juveniles together with mothers with egg sacs suggests a possible two-year life cycle.
</paragraph>
</subSubSection>
<subSubSection box="[199,998,1338,1365]" pageId="13" pageNumber="348" type="distribution">
<paragraph blockId="13.[151,1437,438,1365]" box="[199,998,1338,1365]" pageId="13" pageNumber="348">
<emphasis bold="true" box="[199,353,1338,1364]" pageId="13" pageNumber="348">Distribution.</emphasis>
Forests around the south Portuguese city of Carrapateira.
</paragraph>
</subSubSection>
</treatment>
</document>