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<document id="5602F03DF17F4378E0A0A62FE6B7D336" ID-CLB-Dataset="46430" ID-DOI="10.5281/zenodo.206103" ID-GBIF-Dataset="0b4037b0-5302-497a-ae49-bb2f074d6293" ID-ISSN="1175-5326" ID-Zenodo-Dep="206103" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460384793339" checkinUser="plazi" docAuthor="Imura, Jimpei" docDate="2011" docId="039CCB09FFE69D20439A75209694E8A4" docLanguage="en" docName="zt03012p047.pdf" docOrigin="Zootaxa 3012" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Chilocoris nitidulus Lis" docType="treatment" docVersion="9" lastPageNumber="40" masterDocId="FFA5B371FFEF9D29430D77679432EF08" masterDocTitle="New and obscure species of the genus Chilocorus Mayr in eastern Asia, with the proposal of a “ nitidus - group ” concept (Hemiptera: Heteroptera: Cydnidae)" masterLastPageNumber="47" masterPageNumber="31" pageNumber="40" updateTime="1698297366409" updateUser="ExternalLinkService">
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<mods:title id="C4EC9F34CA22045180905FA36A5E68EA">New and obscure species of the genus Chilocorus Mayr in eastern Asia, with the proposal of a “ nitidus - group ” concept (Hemiptera: Heteroptera: Cydnidae)</mods:title>
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<mods:namePart id="4A4297EA4D6400EF40B27C4BF04E4406">Imura, Jimpei</mods:namePart>
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<mods:date id="05794298436ACB75F3EE153BB8484F93">2011</mods:date>
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<treatment id="039CCB09FFE69D20439A75209694E8A4" ID-DOI="http://doi.org/10.5281/zenodo.6182568" ID-GBIF-Taxon="119415092" ID-Zenodo-Dep="6182568" LSID="urn:lsid:plazi:treatment:039CCB09FFE69D20439A75209694E8A4" httpUri="http://treatment.plazi.org/id/039CCB09FFE69D20439A75209694E8A4" lastPageNumber="40" pageId="9" pageNumber="40">
<subSubSection id="C32F2994FFE69D20439A752095ACED8A" pageId="9" pageNumber="40" type="nomenclature">
<paragraph id="8B8A7A1FFFE69D20439A75209587ED69" blockId="9.[151,437,583,642]" box="[151,437,583,609]" pageId="9" pageNumber="40">
<heading id="D0C2CD73FFE69D20439A75209587ED69" bold="true" box="[151,437,583,609]" fontSize="11" level="1" pageId="9" pageNumber="40" reason="1">
<taxonomicName id="4C35019CFFE69D20439A75209587ED69" authority="Lis" authorityName="Lis" box="[151,437,583,609]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B941A60DFFE69D20439A75209587ED69" bold="true" box="[151,437,583,609]" pageId="9" pageNumber="40">
<emphasis id="B941A60DFFE69D20439A752095B5ED69" bold="true" box="[151,391,583,609]" italics="true" pageId="9" pageNumber="40">Chilocoris nitidulus</emphasis>
Lis
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8A7A1FFFE69D20439A750D95ACED8A" blockId="9.[151,437,583,642]" box="[151,414,618,642]" pageId="9" pageNumber="40">
(
<figureCitation id="130E669AFFE69D204392750D94DDED8A" box="[159,239,618,642]" captionStart="FIGURES 1 10" captionStartId="2.[151,264,1903,1926]" captionTargetBox="[156,1433,600,1870]" captionTargetId="figure@2.[151,1436,591,1879]" captionTargetPageId="2" captionText="FIGURES 1 10. Chilocoris spp.: C. alienus (1 3), C. birmanus (4 5), C. monticola n. sp. (6 8), C. nitidulus (9), and C. nigricans (10). 1 &amp; 4, head and pronotum; 2 &amp; 5 6, right half of head; 3, 8 &amp; 10, evaporatoria; 7, left hind wing; 9, pronotum. Abbreviations: Ant. Smg. L., anterior submarginal line; S. S., secondary setae of head. Scales: 0.5 mm for 1, 3, 4 &amp; 7 10; 0.2 mm for 2 &amp; 5 6." httpUri="https://zenodo.org/record/206104/files/figure.png" pageId="9" pageNumber="40">Figs. 9</figureCitation>
,
<figureCitation id="130E669AFFE69D2043F7750D9571ED8A" box="[250,323,618,642]" captionStart="FIGURES 21 30" captionStartId="5.[151,264,1827,1850]" captionTargetBox="[165,1411,740,1804]" captionTargetId="figure@5.[151,1436,717,1806]" captionTargetPageId="5" captionText="FIGURES 21 30. Terminalia of Chilocoris spp.: C. monticola n. sp. (21 22), C. birmanus (23 25), and C. nitidulus (26 28), C. nigricans (29 30). 21, pygophore, hair on left half omitted; 22 23, pygophoral openings; 24 &amp; 26, median projections; 25, proctiger, hair on left half omitted; 27 28, inflated phallus, lateral (27) and dorsal (28) views; 29 30, endosoma of inflated phallus, lateral (29) and ventral (direction of arrow in 29) (30) views. Abbreviations: Lat. If, lateral infolding of pygophoral opening; Med. Proj., median projection. Scales: 0.2 mm for 21 23, 25 &amp; 27 28; 0.1 mm for 24, 26 &amp; 29 30." httpUri="https://zenodo.org/record/206108/files/figure.png" pageId="9" pageNumber="40">2628</figureCitation>
,
<figureCitation id="130E669AFFE69D204243750D95A4ED8A" box="[334,406,618,642]" captionStart="FIGURES 31 42" captionStartId="6.[151,264,1142,1165]" captionTargetBox="[160,1430,197,1108]" captionTargetId="figure@6.[151,1436,193,1122]" captionTargetPageId="6" captionText="FIGURES 31 42. Left parameres of Chilocoris spp., mesal (31, 34, 37 &amp; 40), ventral (32, 35, 38 &amp; 41) and dorsal (33, 36, 39 &amp; 42) views: C. alienus (31 33), C. birmanus (34 36), C. monticola n. sp. (37 39), and C. nitidulus (40 42). Abbreviation: Hp.: hypophysis. Scales: 0.2 mm." httpUri="https://zenodo.org/record/206109/files/figure.png" pageId="9" pageNumber="40">4042</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32F2994FFE69D20439A75D596A4EDEF" pageId="9" pageNumber="40" type="reference_group">
<paragraph id="8B8A7A1FFFE69D20439A75D595C6EDC0" blockId="9.[151,662,690,743]" box="[151,500,690,712]" pageId="9" pageNumber="40">
<treatmentCitationGroup id="AB255D31FFE69D20439A75D595C6EDC0" box="[151,500,690,712]" pageId="9" pageNumber="40">
<taxonomicName id="4C35019CFFE69D20439A75D595C2EDC0" authority="Lis, 1994: 72" authorityName="Lis" authorityPageNumber="72" authorityYear="1994" box="[151,496,690,712]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B941A60DFFE69D20439A75D59553EDC0" box="[151,353,690,712]" italics="true" pageId="9" pageNumber="40">Chilocoris nitidulus</emphasis>
<treatmentCitation id="0A945C0EFFE69D20426A75D595C2EDC0" author="Lis" box="[359,496,690,712]" page="72" pageId="9" pageNumber="40" year="1994">
<bibRefCitation id="EFA407EEFFE69D20426A75D595F8EDC0" author="Lis" box="[359,458,690,712]" pageId="9" pageNumber="40" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae), Upper Silesian Museum, Bytom, 1 - 349 pp." type="book chapter" year="1994">Lis, 1994</bibRefCitation>
: 72
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8A7A1FFFE69D20439A75B696A4EDEF" blockId="9.[151,662,690,743]" box="[151,662,721,743]" pageId="9" pageNumber="40">
<treatmentCitationGroup id="AB255D31FFE69D20439A75B696A4EDEF" box="[151,662,721,743]" pageId="9" pageNumber="40">
<taxonomicName id="4C35019CFFE69D20439A75B6957FEDEF" box="[151,333,721,743]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B941A60DFFE69D20439A75B6957FEDEF" box="[151,333,721,743]" italics="true" pageId="9" pageNumber="40">Chilocoris nitidus</emphasis>
</taxonomicName>
[part],
<emphasis id="B941A60DFFE69D20429575B5958CEDEF" box="[408,446,722,743]" italics="true" pageId="9" pageNumber="40">non</emphasis>
Mayr:
<treatmentCitation id="0A945C0EFFE69D20410475B696A0EDEF" author="Lis" box="[521,658,721,743]" page="73" pageId="9" pageNumber="40" year="1994">
<bibRefCitation id="EFA407EEFFE69D20410475B6965FEDEF" author="Lis" box="[521,621,721,743]" pageId="9" pageNumber="40" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae), Upper Silesian Museum, Bytom, 1 - 349 pp." type="book chapter" year="1994">Lis, 1994</bibRefCitation>
: 73
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32F2994FFE69D20439A74749716E917" pageId="9" pageNumber="40" type="diagnosis">
<paragraph id="8B8A7A1FFFE69D20439A747496BFEC9F" blockId="9.[151,1437,787,1964]" pageId="9" pageNumber="40">
<emphasis id="B941A60DFFE69D20439A74749523EC24" bold="true" box="[151,273,787,812]" pageId="9" pageNumber="40">Diagnosis.</emphasis>
Total length 3.905.15; total width 2.202.75. Arrangement of paraclypeal secondary setae, in order from mesalmost to lateralmost: 3 pegs - 1 hair - 2 pegs - 1 hair - 1 peg. Ocular index 3.23.6. Antennal segment III 2.73.2× as long as II. Pronotum (
<figureCitation id="130E669AFFE69D20412A743B965FEC7C" box="[551,621,860,884]" captionStart="FIGURES 1 10" captionStartId="2.[151,264,1903,1926]" captionTargetBox="[156,1433,600,1870]" captionTargetId="figure@2.[151,1436,591,1879]" captionTargetPageId="2" captionText="FIGURES 1 10. Chilocoris spp.: C. alienus (1 3), C. birmanus (4 5), C. monticola n. sp. (6 8), C. nitidulus (9), and C. nigricans (10). 1 &amp; 4, head and pronotum; 2 &amp; 5 6, right half of head; 3, 8 &amp; 10, evaporatoria; 7, left hind wing; 9, pronotum. Abbreviations: Ant. Smg. L., anterior submarginal line; S. S., secondary setae of head. Scales: 0.5 mm for 1, 3, 4 &amp; 7 10; 0.2 mm for 2 &amp; 5 6." httpUri="https://zenodo.org/record/206104/files/figure.png" pageId="9" pageNumber="40">Fig. 9</figureCitation>
) nearly trapezoid, 1.731.85× as wide as long, rather strongly tapered anterad, with 56 setae on lateral submargin.
</paragraph>
<paragraph id="8B8A7A1FFFE69D2043C874C39571EB78" blockId="9.[151,1437,787,1964]" pageId="9" pageNumber="40">
This species is closely related to
<emphasis id="B941A60DFFE69D20413774C2969DECB4" box="[570,687,932,956]" italics="true" pageId="9" pageNumber="40">
<taxonomicName id="4C35019CFFE69D20413774C29699ECB4" box="[570,683,932,956]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidus">C. nitidus</taxonomicName>
,
</emphasis>
but was distinguished from it by
<bibRefCitation id="EFA407EEFFE69D20472074C39091ECB4" author="Lis" box="[1069,1187,932,956]" pageId="9" pageNumber="40" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae), Upper Silesian Museum, Bytom, 1 - 349 pp." type="book chapter" year="1994">Lis (1994)</bibRefCitation>
who used the relative length of antennal segment III to II, the shape of pronotum, and the shape of the paramere. However, Lis based his conclusions on only
<specimenCount id="9D33B196FFE69D204270748B95FAEB0C" box="[381,456,1004,1028]" pageId="9" pageNumber="40" type="male">1 male</specimenCount>
<typeStatus id="548EC4BDFFE69D2042C2748B9603EB0C" box="[463,561,1004,1028]" pageId="9" pageNumber="40" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C35019CFFE69D20415B748A96EBEB0C" box="[598,729,1004,1028]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B941A60DFFE69D20415B748A96EBEB0C" box="[598,729,1004,1028]" italics="true" pageId="9" pageNumber="40">C. nitidulus</emphasis>
</taxonomicName>
. Our investigation into the species, based on additional materials, showed that the former 2 structures are insufficient to separate these 2 species precisely. Therefore, at present, the male genital structure appears to be the only reliable characteristic to distinguish these species; this is described in detail below.
</paragraph>
<paragraph id="8B8A7A1FFFE69D2043C8731B900FEA4F" blockId="9.[151,1437,787,1964]" pageId="9" pageNumber="40">
3
<emphasis id="B941A60DFFE69D2043EF731C956CEB9C" bold="true" box="[226,350,1147,1172]" pageId="9" pageNumber="40">terminalia</emphasis>
—Pygophoral opening rounded; median projection truncate, and maybe slightly concave in apical margin (
<figureCitation id="130E669AFFE69D20422C73F89545EBBF" box="[289,375,1183,1207]" captionStart="FIGURES 21 30" captionStartId="5.[151,264,1827,1850]" captionTargetBox="[165,1411,740,1804]" captionTargetId="figure@5.[151,1436,717,1806]" captionTargetPageId="5" captionText="FIGURES 21 30. Terminalia of Chilocoris spp.: C. monticola n. sp. (21 22), C. birmanus (23 25), and C. nitidulus (26 28), C. nigricans (29 30). 21, pygophore, hair on left half omitted; 22 23, pygophoral openings; 24 &amp; 26, median projections; 25, proctiger, hair on left half omitted; 27 28, inflated phallus, lateral (27) and dorsal (28) views; 29 30, endosoma of inflated phallus, lateral (29) and ventral (direction of arrow in 29) (30) views. Abbreviations: Lat. If, lateral infolding of pygophoral opening; Med. Proj., median projection. Scales: 0.2 mm for 21 23, 25 &amp; 27 28; 0.1 mm for 24, 26 &amp; 29 30." httpUri="https://zenodo.org/record/206108/files/figure.png" pageId="9" pageNumber="40">Fig. 26</figureCitation>
). Phallus (
<figureCitation id="130E669AFFE69D2042FC73F89648EBBF" box="[497,634,1183,1207]" captionStart="FIGURES 21 30" captionStartId="5.[151,264,1827,1850]" captionTargetBox="[165,1411,740,1804]" captionTargetId="figure@5.[151,1436,717,1806]" captionTargetPageId="5" captionText="FIGURES 21 30. Terminalia of Chilocoris spp.: C. monticola n. sp. (21 22), C. birmanus (23 25), and C. nitidulus (26 28), C. nigricans (29 30). 21, pygophore, hair on left half omitted; 22 23, pygophoral openings; 24 &amp; 26, median projections; 25, proctiger, hair on left half omitted; 27 28, inflated phallus, lateral (27) and dorsal (28) views; 29 30, endosoma of inflated phallus, lateral (29) and ventral (direction of arrow in 29) (30) views. Abbreviations: Lat. If, lateral infolding of pygophoral opening; Med. Proj., median projection. Scales: 0.2 mm for 21 23, 25 &amp; 27 28; 0.1 mm for 24, 26 &amp; 29 30." httpUri="https://zenodo.org/record/206108/files/figure.png" pageId="9" pageNumber="40">Figs. 2728</figureCitation>
) relatively small; endosoma tapered, slightly pigmented distal to conjunctival appendages; conjunctival appendages totally membranous, lobe-like, composed of slender dorsal pair and stout ventral pair, not reflexed in inflated position; endophallic duct relatively short, coiled in 1 time from its turning point, and protruding from dorsum of endosoma. Paramere (
<figureCitation id="130E669AFFE69D20406C726B97D5EA2C" box="[865,999,1292,1316]" captionStart="FIGURES 31 42" captionStartId="6.[151,264,1142,1165]" captionTargetBox="[160,1430,197,1108]" captionTargetId="figure@6.[151,1436,193,1122]" captionTargetPageId="6" captionText="FIGURES 31 42. Left parameres of Chilocoris spp., mesal (31, 34, 37 &amp; 40), ventral (32, 35, 38 &amp; 41) and dorsal (33, 36, 39 &amp; 42) views: C. alienus (31 33), C. birmanus (34 36), C. monticola n. sp. (37 39), and C. nitidulus (40 42). Abbreviation: Hp.: hypophysis. Scales: 0.2 mm." httpUri="https://zenodo.org/record/206109/files/figure.png" pageId="9" pageNumber="40">Figs. 4042</figureCitation>
) slender, rounded apically; hypophysis moderately stout, sharpened. Proctiger moderately shortened, rounded-pentangular.
</paragraph>
<paragraph id="8B8A7A1FFFE69D2043C872349716E917" blockId="9.[151,1437,787,1964]" pageId="9" pageNumber="40">
<emphasis id="B941A60DFFE69D2043C8723495F4EA64" bold="true" box="[197,454,1363,1388]" pageId="9" pageNumber="40">Specimens examined.</emphasis>
432Ƥ—VIETNAM: Deo
<collectingCountry id="F3223A8FFFE69D2041F77233973CEA64" box="[762,782,1364,1388]" name="Iceland" pageId="9" pageNumber="40">O</collectingCountry>
Quy Ho,
<quantity id="4CCDD7FAFFE69D20408F723397FDEA64" box="[898,975,1364,1388]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="9" pageNumber="40" unit="m" value="1750.0">1750m</quantity>
alt., Sapa,
<collectingCountry id="F3223A8FFFE69D20475C7232904EEA64" box="[1105,1148,1365,1388]" name="Laos" pageId="9" pageNumber="40">Lao</collectingCountry>
Cai Prov.: 13,
<date id="FF8B5CDFFFE69D20463E723391A5EA64" box="[1331,1431,1364,1388]" pageId="9" pageNumber="40" value="1995-10-03">3.x.1995</date>
, M. Tomokuni leg. (NSMT-I-He 36463); 1Ƥ,
<date id="FF8B5CDFFFE69D20419472109721EA87" box="[665,787,1399,1423]" pageId="9" pageNumber="40" value="1997-06-28">28.vi.1997</date>
, M. Tomokuni leg. (NSMT-I-He 36461). Deo Tram Ton,
<quantity id="4CCDD7FAFFE69D20439A72FB94D7EABC" box="[151,229,1436,1460]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="9" pageNumber="40" unit="m" value="1950.0">1950m</quantity>
alt., Sapa,
<collectingCountry id="F3223A8FFFE69D20426A72FA95A0EABC" box="[359,402,1437,1460]" name="Laos" pageId="9" pageNumber="40">Lao</collectingCountry>
Cai Prov.: 131Ƥ,
<date id="FF8B5CDFFFE69D20416172FB96E3EABC" box="[620,721,1436,1460]" pageId="9" pageNumber="40" value="1995-10-04">4.x.1995</date>
, M. Tomokuni leg. (NSMT-I-He 36460, 36462).
<collectingCountry id="F3223A8FFFE69D20460E72FB91AAEABC" box="[1283,1432,1436,1460]" name="Thailand" pageId="9" pageNumber="40">THAILAND</collectingCountry>
: Pakia,
<quantity id="4CCDD7FAFFE69D2043EB72D89507EADF" box="[230,309,1471,1495]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="40" unit="m" value="1500.0">1500m</quantity>
alt., Chiang
<collectingCountry id="F3223A8FFFE69D2042C372A795CDEADF" box="[462,511,1472,1495]" name="Iceland" pageId="9" pageNumber="40">Dao</collectingCountry>
, Chiang Mai:
<date id="FF8B5CDFFFE69D2041A572D8972EEADF" box="[680,796,1471,1495]" pageId="9" pageNumber="40" value="1913-05-05" valueMax="1913-05-08" valueMin="1913-05-05">13, 58.v.</date>
2002, T. Ishizaki leg. by Malaise trap (TUA).
<collectingCountry id="F3223A8FFFE69D20463072A791ABEAD0" box="[1341,1433,1472,1496]" name="Nepal" pageId="9" pageNumber="40">NEPAL</collectingCountry>
: Phakding,
<quantity id="4CCDD7FAFFE69D2042037283956EEAF4" box="[270,348,1508,1532]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="9" pageNumber="40" unit="m" value="2580.0">2580m</quantity>
alt., Solukhumbu: 13,
<date id="FF8B5CDFFFE69D20416E728396EEEAF4" box="[611,732,1508,1532]" pageId="9" pageNumber="40" value="1979-09-29">29.ix.1979</date>
, M. Sato leg., det. as
<taxonomicName id="4C35019CFFE69D2040C172839070EAF4" box="[972,1090,1508,1532]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B941A60DFFE69D2040C172839070EAF4" box="[972,1090,1508,1532]" italics="true" pageId="9" pageNumber="40">Chilocoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2721B76FFE69D204747728290B6EAF4" box="[1098,1156,1509,1532]" pageId="9" pageNumber="40" sensu="stricto">s. str.</taxonomicNameLabel>
<taxonomicName id="4C35019CFFE69D20478672839129EAF5" authority="Mayr" authorityName="Mayr" box="[1163,1307,1508,1533]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B941A60DFFE69D204786728390E5EAF4" box="[1163,1239,1508,1532]" italics="true" pageId="9" pageNumber="40">nitidus</emphasis>
Mayr
</taxonomicName>
by J.A. Lis (
<emphasis id="B941A60DFFE69D2043AD716F948EE928" box="[160,188,1544,1568]" italics="true" pageId="9" pageNumber="40">cf.</emphasis>
<bibRefCitation id="EFA407EEFFE69D2043CF71609514E917" author="Lis" box="[194,294,1543,1567]" pageId="9" pageNumber="40" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae), Upper Silesian Museum, Bytom, 1 - 349 pp." type="book chapter" year="1994">Lis 1994</bibRefCitation>
) (
<figureCitation id="130E669AFFE69D204233716095BFE917" box="[318,397,1543,1567]" captionStart="FIGURES 1 10" captionStartId="2.[151,264,1903,1926]" captionTargetBox="[156,1433,600,1870]" captionTargetId="figure@2.[151,1436,591,1879]" captionTargetPageId="2" captionText="FIGURES 1 10. Chilocoris spp.: C. alienus (1 3), C. birmanus (4 5), C. monticola n. sp. (6 8), C. nitidulus (9), and C. nigricans (10). 1 &amp; 4, head and pronotum; 2 &amp; 5 6, right half of head; 3, 8 &amp; 10, evaporatoria; 7, left hind wing; 9, pronotum. Abbreviations: Ant. Smg. L., anterior submarginal line; S. S., secondary setae of head. Scales: 0.5 mm for 1, 3, 4 &amp; 7 10; 0.2 mm for 2 &amp; 5 6." httpUri="https://zenodo.org/record/206104/files/figure.png" pageId="9" pageNumber="40">Figs. 9</figureCitation>
,
<figureCitation id="130E669AFFE69D204294716095D3E917" box="[409,481,1543,1567]" captionStart="FIGURES 21 30" captionStartId="5.[151,264,1827,1850]" captionTargetBox="[165,1411,740,1804]" captionTargetId="figure@5.[151,1436,717,1806]" captionTargetPageId="5" captionText="FIGURES 21 30. Terminalia of Chilocoris spp.: C. monticola n. sp. (21 22), C. birmanus (23 25), and C. nitidulus (26 28), C. nigricans (29 30). 21, pygophore, hair on left half omitted; 22 23, pygophoral openings; 24 &amp; 26, median projections; 25, proctiger, hair on left half omitted; 27 28, inflated phallus, lateral (27) and dorsal (28) views; 29 30, endosoma of inflated phallus, lateral (29) and ventral (direction of arrow in 29) (30) views. Abbreviations: Lat. If, lateral infolding of pygophoral opening; Med. Proj., median projection. Scales: 0.2 mm for 21 23, 25 &amp; 27 28; 0.1 mm for 24, 26 &amp; 29 30." httpUri="https://zenodo.org/record/206108/files/figure.png" pageId="9" pageNumber="40">2628</figureCitation>
,
<figureCitation id="130E669AFFE69D2042E071609606E917" box="[493,564,1543,1567]" captionStart="FIGURES 31 42" captionStartId="6.[151,264,1142,1165]" captionTargetBox="[160,1430,197,1108]" captionTargetId="figure@6.[151,1436,193,1122]" captionTargetPageId="6" captionText="FIGURES 31 42. Left parameres of Chilocoris spp., mesal (31, 34, 37 &amp; 40), ventral (32, 35, 38 &amp; 41) and dorsal (33, 36, 39 &amp; 42) views: C. alienus (31 33), C. birmanus (34 36), C. monticola n. sp. (37 39), and C. nitidulus (40 42). Abbreviation: Hp.: hypophysis. Scales: 0.2 mm." httpUri="https://zenodo.org/record/206109/files/figure.png" pageId="9" pageNumber="40">4042</figureCitation>
) (NSMT-I-He 9124).
</paragraph>
</subSubSection>
<subSubSection id="C32F2994FFE69D2043C8714C962CE96F" pageId="9" pageNumber="40" type="distribution">
<paragraph id="8B8A7A1FFFE69D2043C8714C970BE94C" blockId="9.[151,1437,787,1964]" box="[197,825,1579,1604]" pageId="9" pageNumber="40">
<emphasis id="B941A60DFFE69D2043C8714C956CE94C" bold="true" box="[197,350,1579,1604]" pageId="9" pageNumber="40">Distribution.</emphasis>
<collectingCountry id="F3223A8FFFE69D204268714B95F6E94C" box="[357,452,1580,1604]" name="Vietnam" pageId="9" pageNumber="40">Vietnam</collectingCountry>
(north);
<collectingCountry id="F3223A8FFFE69D20412A714B96B9E94C" box="[551,651,1580,1604]" name="Thailand" pageId="9" pageNumber="40">Thailand</collectingCountry>
(north);
<collectingCountry id="F3223A8FFFE69D2041E3714B9707E94C" box="[750,821,1580,1604]" name="Nepal" pageId="9" pageNumber="40">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8B8A7A1FFFE69D2043C87136962CE96F" blockId="9.[151,1437,787,1964]" box="[197,542,1615,1640]" pageId="9" pageNumber="40">
New to
<collectingCountry id="F3223A8FFFE69D2042137128954FE96F" box="[286,381,1615,1639]" name="Vietnam" pageId="9" pageNumber="40">Vietnam</collectingCountry>
and
<collectingCountry id="F3223A8FFFE69D2042BE71289628E96F" box="[435,538,1615,1639]" name="Thailand" pageId="9" pageNumber="40">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32F2994FFE69D2043C871139694E8A4" pageId="9" pageNumber="40" type="discussion">
<paragraph id="8B8A7A1FFFE69D2043C87113962AE848" blockId="9.[151,1437,787,1964]" pageId="9" pageNumber="40">
<emphasis id="B941A60DFFE69D2043C87113950BE984" bold="true" box="[197,313,1652,1676]" pageId="9" pageNumber="40">Remarks.</emphasis>
The original description of this species from Central
<collectingCountry id="F3223A8FFFE69D2040AA711397D8E984" box="[935,1002,1652,1676]" name="Nepal" pageId="9" pageNumber="40">Nepal</collectingCountry>
was based solely on a male
<typeStatus id="548EC4BDFFE69D204637711391AEE984" box="[1338,1436,1652,1676]" pageId="9" pageNumber="40" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFA407EEFFE69D20439271F0953FE9A7" author="Lis" box="[159,269,1687,1711]" pageId="9" pageNumber="40" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae), Upper Silesian Museum, Bytom, 1 - 349 pp." type="book chapter" year="1994">Lis, 1994</bibRefCitation>
), and no other specimens have been reported until now. In this study, a male specimen from
<collectingCountry id="F3223A8FFFE69D20460371F09163E9A7" box="[1294,1361,1687,1711]" name="Nepal" pageId="9" pageNumber="40">Nepal</collectingCountry>
which was identified as
<taxonomicName id="4C35019CFFE69D20425771DA95F4E9DC" box="[346,454,1724,1748]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B941A60DFFE69D20425771DA95F4E9DC" box="[346,454,1724,1748]" italics="true" pageId="9" pageNumber="40">C. nitidus</emphasis>
</taxonomicName>
by J. A. Lis was examined, but it corresponded with the diagnostic characters of
<emphasis id="B941A60DFFE69D20465971DA94D0E9F0" italics="true" pageId="9" pageNumber="40">
<taxonomicName id="4C35019CFFE69D20465971DA94ECE9F0" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidulus">C. nitidulus</taxonomicName>
,
</emphasis>
rather than
<taxonomicName id="4C35019CFFE69D204267718795E5E9F0" box="[362,471,1760,1784]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B941A60DFFE69D204267718795E5E9F0" box="[362,471,1760,1784]" italics="true" pageId="9" pageNumber="40">C. nitidus</emphasis>
</taxonomicName>
, in the shape of the pronotum and paramere. Hence, this specimen was determined to be
<taxonomicName id="4C35019CFFE69D2043B57062950BE814" box="[184,313,1796,1820]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B941A60DFFE69D2043B57062950BE814" box="[184,313,1796,1820]" italics="true" pageId="9" pageNumber="40">C. nitidulus</emphasis>
</taxonomicName>
. In addition, some specimens collected from other countries of southeastern Asia were considered to be conspecific with this specimen.
</paragraph>
<paragraph id="8B8A7A1FFFE69D2043C8702B9694E8A4" blockId="9.[151,1437,787,1964]" pageId="9" pageNumber="40">
This study demonstrates that this species is more broadly distributed over the southeastern Asia than previously thought. It is likely that some records of
<taxonomicName id="4C35019CFFE69D2041AE70179722E880" box="[675,784,1904,1928]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B941A60DFFE69D2041AE70179722E880" box="[675,784,1904,1928]" italics="true" pageId="9" pageNumber="40">C. nitidus</emphasis>
</taxonomicName>
from this region may actually pertain to
<taxonomicName id="4C35019CFFE69D2047D17017916CE880" box="[1244,1374,1904,1928]" class="Insecta" family="Cydnidae" genus="Chilocoris" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="B941A60DFFE69D2047D17017916CE880" box="[1244,1374,1904,1928]" italics="true" pageId="9" pageNumber="40">C. nitidulus</emphasis>
</taxonomicName>
, and, therefore, relevant records should be reviewed.
</paragraph>
</subSubSection>
</treatment>
</document>