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<document id="6A51B3FB96D818EEB564A48A6946889C" ID-CLB-Dataset="281316" ID-DOI="10.5852/ejt.2023.910.2361" ID-GBIF-Dataset="43a5768f-e8c1-4484-81da-78217a053132" ID-ISSN="2118-9773" ID-Zenodo-Dep="10306494" ID-ZooBank="2B90DE6E-E70A-4199-B485-5A13FDD43056" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1702027259743" checkinUser="plazi" docAuthor="Kuwahara, Gregory K. &amp; Marshall, Stephen A." docDate="2023" docId="039C87E4FFE7FFB40D1ADD6BFBC9FC1C" docLanguage="en" docName="EJT.2023.910.69-97.pdf" docOrigin="European Journal of Taxonomy 910" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2361/10335" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pleuroseta monteithi Kuwahara &amp; Marshall 2023, sp. nov." docType="treatment" docUuid="9321FA5F-A43E-4B05-8C88-E661260B9E23" docUuidSource="ZooBank" docVersion="3" lastPageNumber="91" masterDocId="FFA5FF9CFFF6FFA20F71DE58FFE9FFC3" masterDocTitle="A revision of the Australian genus Pleuroseta Richards (Diptera, Sphaeroceridae)" masterLastPageNumber="97" masterPageNumber="69" pageNumber="86" updateTime="1702060955232" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="448CEC8FB03A4D338E7982A5DE82177A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="87BD6CD4AE494CBD26F8127342C912CC">
<mods:title id="1405ABAAC7ACCC536DBD91EEB32B5407">A revision of the Australian genus Pleuroseta Richards (Diptera, Sphaeroceridae)</mods:title>
</mods:titleInfo>
<mods:name id="9731C42E3B46101DF3B82036E3FDFE87" type="personal">
<mods:role id="93A1868A67F9A43F27484FCA3F5611F1">
<mods:roleTerm id="56CA77BB153D25D677198BDEFCDEB8C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C4AB76EAD5A952ADF60279233D5863F6">Kuwahara, Gregory K.</mods:namePart>
<mods:nameIdentifier id="67E51C72A0F2552B07A66BA96C5E37C2" type="LSID">2E2D313F-7249-4F76-AF5D-615516840188</mods:nameIdentifier>
<mods:affiliation id="C850ABAC3F05A94E13E5E4A32B36E92C">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
<mods:nameIdentifier id="A4279E18E23506AE3D46CC39CFC8EEC6" type="email">gkuwahar@uoguelph.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="4A3F638B5AAD72E29CFBC4B31305D2AB" type="personal">
<mods:role id="FDDFF05EA198FEC79526859FE25D86E7">
<mods:roleTerm id="FF1B17E416B1B8766075A6CE6808D89E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8CD97E8DAF2C0AF32447A438463A81B6">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="F7E0248A812D1BDE70EC2D2AB19BE3D3" type="LSID">6C1C1C9E-365F-48BF-B8C7-31163EC5F3A7</mods:nameIdentifier>
<mods:affiliation id="A88F26CC8B3EB6B349B002598F812AE7">University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.</mods:affiliation>
<mods:nameIdentifier id="CB2559BA536EFC5C84C6BFADFCDC70B5" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="30B9CDF8FE1D5D2E9145EFDCAAC37DEA">text</mods:typeOfResource>
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<mods:title id="10573EC7A8BDB03D7E9296798B02AF83">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="EED31930D13AF008044B79D3B11CF30D">
<mods:date id="45480795B1B43227DDCDCA7E96A5FFBA">2023</mods:date>
<mods:detail id="F833F7300634B59F5E4FAABD7B138870" type="pubDate">
<mods:number id="96810533C6202EF634C05F64981FB664">2023-12-07</mods:number>
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<mods:detail id="7ACCEBBF4417A498E1CC93CB307CF509" type="volume">
<mods:number id="A5B5298B8C3A8232AD9AF6C3A52337E0">910</mods:number>
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<mods:start id="D57AD226650A4F13A3A33FAC49379CAF">69</mods:start>
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<mods:url id="5718451615188C821898125F22C7A16A">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2361/10335</mods:url>
</mods:location>
<mods:classification id="7A44ED4D37D78BD3524386BA0EC7DF59">journal article</mods:classification>
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</mods:mods>
<treatment id="039C87E4FFE7FFB40D1ADD6BFBC9FC1C" LSID="urn:lsid:zoobank.org:act:9321FA5F-A43E-4B05-8C88-E661260B9E23" httpUri="http://treatment.plazi.org/id/039C87E4FFE7FFB40D1ADD6BFBC9FC1C" lastPageId="22" lastPageNumber="91" pageId="17" pageNumber="86">
<subSubSection id="C32F6579FFE7FFB30D1ADD6BFB57FCB1" pageId="17" pageNumber="86" type="nomenclature">
<paragraph id="8B8A36F2FFE7FFB30D1ADD6BFC20FC8D" blockId="17.[373,1214,819,917]" box="[619,969,819,846]" pageId="17" pageNumber="86">
<heading id="D0C2819EFFE7FFB30D1ADD6BFC20FC8D" box="[619,969,819,846]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="86" reason="2">
<taxonomicName id="4C354D71FFE7FFB30D1ADD6BFC8EFC8D" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[619,871,819,846]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="86" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B941EAE0FFE7FFB30D1ADD6BFC8EFC8D" bold="true" box="[619,871,819,846]" italics="true" pageId="17" pageNumber="86">Pleuroseta monteithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE7FFB30C1FDD6CFC20FC8D" box="[878,969,820,846]" pageId="17" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8A36F2FFE7FFB30E04DD0FFB57FCB1" blockId="17.[373,1214,819,917]" box="[373,1214,855,882]" pageId="17" pageNumber="86">
<uri id="FFA43AF0FFE7FFB30E04DD0FFB57FCB1" box="[373,1214,855,882]" pageId="17" pageNumber="86">
urn:lsid:zoobank.org:act:
<uuid id="FF930C27FFE7FFB30DE4DD0FFB57FCB1" box="[661,1214,855,882]" pageId="17" pageNumber="86">9321FA5F-A43E-4B05-8C88-E661260B9E23</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32F6579FFE7FFB30DD0DD22FC7AFC57" box="[673,915,890,917]" pageId="17" pageNumber="86" type="description">
<paragraph id="8B8A36F2FFE7FFB30DD0DD22FC7AFC57" blockId="17.[373,1214,819,917]" box="[673,915,890,917]" pageId="17" pageNumber="86">
<figureCitation id="130E2A77FFE7FFB30DD0DD22FD17FC56" box="[673,766,890,917]" captionStart="Fig" captionStartId="10.[189,230,1857,1883]" captionTargetBox="[243,1343,265,1815]" captionTargetId="figure-65@10.[243,1343,265,1820]" captionTargetPageId="10" captionText="Fig. 5. A. Pleuroseta occidentalis sp. nov. paratype, left wing, ventral view (debu00244165). B. Pleuroseta occidentalis paratype, hypandrium, dorsal view (debu00245797). C. Pleuroseta wentworthi (Richards, 1973), hypandrium, dorsal view (debu00264733). D. Pleuroseta ingens sp. nov. paratype, hypandrium, dorsal view (debu00160984). E. Pleuroseta monteithi sp. nov. paratype, hypandrium, dorsal view (debu00160892). Scale bars: A=0.50 mm, B–E=0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.10306508" httpUri="https://zenodo.org/record/10306508/files/figure.png" pageId="17" pageNumber="86">Figs 5E</figureCitation>
,
<figureCitation id="130E2A77FFE7FFB30C7BDD22FCBCFC57" box="[778,853,890,916]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[189,232,1892,1918]" captionStartId-1="19.[189,232,1928,1954]" captionTargetBox-0="[243,1340,265,1814]" captionTargetBox-1="[236,1343,278,1882]" captionTargetId-0="figure-54@18.[243,1343,265,1820]" captionTargetId-1="figure-1@19.[223,1365,272,1886]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 10. Pleuroseta monteithi sp. nov. A. Male holotype habitus, lateral view (debu00160870). B. Male paratype terminalia, lateral view (debu00160892). C. Male paratype terminalia, posterolateral view (debu00160892). Abbreviations: epa =epandrium; hyp =hypandrium; se =subepandrial sclerite; ss =surstylus. Scale bars: A = 1.0 mm; B–C =0.40 mm." captionText-1="Fig. 11. Pleuroseta monteithi sp. nov. paratype (debu00160892). A. Aedeagus and associated structures, lateral view. B. Male S5–8, ventral view. Abbreviations: bp = basiphallus; dp =distiphallus; pg =postgonite; phal=phallapodeme; rs =ring sclerite. Scale bars: A=0.40 mm; B= 0.60 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10306520" figureDoi-1="http://doi.org/10.5281/zenodo.10306522" httpUri-0="https://zenodo.org/record/10306520/files/figure.png" httpUri-1="https://zenodo.org/record/10306522/files/figure.png" pageId="17" pageNumber="86">10–11</figureCitation>
,
<figureCitation id="130E2A77FFE7FFB30C10DD22FC7AFC57" box="[865,915,890,916]" captionStart="Fig" captionStartId="26.[189,231,1928,1954]" captionTargetBox="[227,1360,292,1886]" captionTargetId="figure-36@26.[227,1360,289,1890]" captionTargetPageId="26" captionText="Fig. 16. Pleuroseta left wings, ventral view. A. Pleuroseta ingens sp. nov. paratype (debu00162128). B. Pleuroseta monteithi sp. nov. paratype (debu00160892). C. Pleuroseta wentworthi (Richards, 1973) (debu402972). Scale bar: A–C= 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10306534" httpUri="https://zenodo.org/record/10306534/files/figure.png" pageId="17" pageNumber="86">16B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F6579FFE7FFB30FCCDDE4FC6AFBEB" pageId="17" pageNumber="86" type="etymology">
<paragraph id="8B8A36F2FFE7FFB30FCCDDE4FEADFC15" blockId="17.[189,324,956,982]" box="[189,324,956,982]" pageId="17" pageNumber="86">
<heading id="D0C2819EFFE7FFB30FCCDDE4FEADFC15" bold="true" box="[189,324,956,982]" fontSize="11" level="3" pageId="17" pageNumber="86" reason="3">
<emphasis id="B941EAE0FFE7FFB30FCCDDE4FEADFC15" bold="true" box="[189,324,956,982]" pageId="17" pageNumber="86">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A36F2FFE7FFB30FCCDDB2FC6AFBEB" blockId="17.[189,1399,1002,1064]" pageId="17" pageNumber="86">
This species name is in honour of Dr Geoff Monteith, who collected the
<typeStatus id="548E8850FFE7FFB30B65DDB3FBAEFBC6" box="[1044,1095,1003,1029]" pageId="17" pageNumber="86">type</typeStatus>
specimens of this species as well as most of the other known specimens of
<taxonomicName id="4C354D71FFE7FFB30D8FDA55FC95FBE4" authorityName="Richards" authorityYear="1973" box="[766,892,1037,1063]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B941EAE0FFE7FFB30D8FDA55FC95FBE4" box="[766,892,1037,1063]" italics="true" pageId="17" pageNumber="86">Pleuroseta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32F6579FFE7FFB30FCCDA16FB85FA80" pageId="17" pageNumber="86" type="materials_examined">
<paragraph id="8B8A36F2FFE7FFB30FCCDA16FE43FBAB" blockId="17.[189,426,1102,1128]" box="[189,426,1102,1128]" pageId="17" pageNumber="86">
<heading id="D0C2819EFFE7FFB30FCCDA16FE43FBAB" bold="true" box="[189,426,1102,1128]" fontSize="11" level="3" pageId="17" pageNumber="86" reason="3">
<emphasis id="B941EAE0FFE7FFB30FCCDA16FE43FBAB" bold="true" box="[189,426,1102,1128]" pageId="17" pageNumber="86">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A36F2FFE7FFB30F9BDA25FEB5FB54" blockId="17.[189,1399,1149,1246]" box="[234,348,1149,1175]" pageId="17" pageNumber="86">
<emphasis id="B941EAE0FFE7FFB30F9BDA25FEB5FB54" bold="true" box="[234,348,1149,1175]" pageId="17" pageNumber="86">
<typeStatus id="548E8850FFE7FFB30F9BDA25FEB5FB54" box="[234,348,1149,1175]" pageId="17" pageNumber="86" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8A36F2FFE7FFB30FCCDAF9FC3FFB1E" blockId="17.[189,1399,1149,1246]" pageId="17" pageNumber="86">
<materialsCitation id="3B5D3CAFFFE7FFB30FCCDAF9FC3FFB1E" collectingDate="1998-11-18" collectingMethod="dung trap" collectionCode="QMBA" collectorName="G. B. Monteith" country="Australia" location="Cape Tribulation, transect site 8" pageId="17" pageNumber="86" specimenCode="QMBA debu00160870" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="holotype">
<collectingCountry id="F3227662FFE7FFB30FCCDAF9FE8BFB78" box="[189,354,1185,1211]" name="Australia" pageId="17" pageNumber="86">AUSTRALIA</collectingCountry>
•
<specimenCount id="9D33FD7BFFE7FFB30E0EDAF9FE71FB78" box="[383,408,1185,1211]" count="1" pageId="17" pageNumber="86" type="male">♂</specimenCount>
;
<collectingRegion id="49F1F810FFE7FFB30ED7DAF8FDDDFB79" box="[422,564,1184,1210]" country="Australia" name="Queensland" pageId="17" pageNumber="86">Queensland</collectingRegion>
,
<location id="8EEA6029FFE7FFB30D33DAF9FC2BFB78" LSID="urn:lsid:plazi:treatment:039C87E4FFE7FFB40D1ADD6BFBC9FC1C:8EEA6029FFE7FFB30D33DAF9FC2BFB78" box="[578,962,1184,1211]" country="Australia" name="Cape Tribulation, transect site 8" pageId="17" pageNumber="86" stateProvince="Queensland">Cape Tribulation, transect site 8</location>
;
<geoCoordinate id="EE015035FFE7FFB30CA3DAF8FBDEFB78" box="[978,1079,1184,1211]" pageId="17" pageNumber="86">16°05' S</geoCoordinate>
,
<geoCoordinate id="EE015035FFE7FFB30B37DAF8FB54FB78" box="[1094,1213,1184,1211]" pageId="17" pageNumber="86">145°26' E</geoCoordinate>
;
<date id="FF8B1032FFE7FFB30BBADAF9FA9BFB78" box="[1227,1394,1185,1211]" pageId="17" pageNumber="86" value="1998-11-18">
<collectingDate id="EFCFE9DAFFE7FFB30BBADAF9FA9BFB78" box="[1227,1394,1185,1211]" pageId="17" pageNumber="86" value="1998-11-18">18 Nov. 1998</collectingDate>
</date>
;
<collectorName id="26C05324FFE7FFB30FCCDA9CFE81FB1E" box="[189,360,1219,1246]" pageId="17" pageNumber="86">G.B. Monteith</collectorName>
leg.;
<collectingMethod id="52744EE5FFE7FFB30ED8DA9BFDF5FB1D" box="[425,540,1219,1246]" pageId="17" pageNumber="86">dung trap</collectingMethod>
, 6 am –6 pm;
<specimenCode id="DB939E89FFE7FFB30DB3DA9CFC3BFB1E" box="[706,978,1219,1246]" pageId="17" pageNumber="86">QMBA debu00160870</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8A36F2FFE7FFB30F9BDB5DFEB5FADC" blockId="17.[189,1132,1285,1347]" box="[234,348,1285,1311]" pageId="17" pageNumber="86">
<emphasis id="B941EAE0FFE7FFB30F9BDB5DFEB5FADC" bold="true" box="[234,348,1285,1311]" pageId="17" pageNumber="86">
<typeStatus id="548E8850FFE7FFB30F9BDB5DFEB5FADC" box="[234,348,1285,1311]" pageId="17" pageNumber="86" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8A36F2FFE7FFB30FCCDB70FB85FA80" blockId="17.[189,1132,1285,1347]" box="[189,1132,1320,1347]" pageId="17" pageNumber="86">
<materialsCitation id="3B5D3CAFFFE7FFB30FCCDB70FB81FA80" box="[189,1128,1320,1347]" collectingDate="1998-11-18" collectingMethod="dung trap" collectionCode="DEBU" collectorName="G. B. Monteith" country="Australia" location="Queensland" pageId="17" pageNumber="86" specimenCode="QMBA debu00160870" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
<collectingCountry id="F3227662FFE7FFB30FCCDB70FE8BFA81" box="[189,354,1320,1346]" name="Australia" pageId="17" pageNumber="86">AUSTRALIA</collectingCountry>
–
<collectingRegion id="49F1F810FFE7FFB30E0EDB70FDE0FA81" box="[383,521,1320,1346]" country="Australia" name="Queensland" pageId="17" pageNumber="86">Queensland</collectingRegion>
•
<specimenCount id="9D33FD7BFFE7FFB30D53DB70FDB8FA80" box="[546,593,1320,1347]" count="1" pageId="17" pageNumber="86" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED24AE37FFE7FFB30B65DB71FB81FA80" box="[1044,1128,1321,1347]" collectionName="Canada, Ontario, Guelph, University of Guelph" pageId="17" pageNumber="86">DEBU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32F6579FFE7FFB40FCCDB31FD8CFDFB" lastPageId="22" lastPageNumber="91" pageId="17" pageNumber="86" type="description">
<paragraph id="8B8A36F2FFE7FFB30FCCDB31FEA7FA40" blockId="17.[189,334,1385,1411]" box="[189,334,1385,1411]" pageId="17" pageNumber="86">
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<emphasis id="B941EAE0FFE7FFB30FCCDB31FEA7FA40" bold="true" box="[189,334,1385,1411]" pageId="17" pageNumber="86">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A36F2FFE7FFB30FCCDBC1FABAF9DF" blockId="17.[187,1399,1432,1565]" pageId="17" pageNumber="86">
<smallCapsWord id="8D6CA02EFFE7FFB30FCCDBC1FF14FA72" baselines="1452,1452" box="[189,253,1433,1459]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="17" pageNumber="86">BODY</smallCapsWord>
(
<figureCitation id="130E2A77FFE7FFB30E7ADBC0FE9EFA71" box="[267,375,1432,1458]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[243,1340,265,1814]" captionTargetId="figure-54@18.[243,1343,265,1820]" captionTargetPageId="18" captionText="Fig. 10. Pleuroseta monteithi sp. nov. A. Male holotype habitus, lateral view (debu00160870). B. Male paratype terminalia, lateral view (debu00160892). C. Male paratype terminalia, posterolateral view (debu00160892). Abbreviations: epa =epandrium; hyp =hypandrium; se =subepandrial sclerite; ss =surstylus. Scale bars: A = 1.0 mm; B–C =0.40 mm." figureDoi="http://doi.org/10.5281/zenodo.10306520" httpUri="https://zenodo.org/record/10306520/files/figure.png" pageId="17" pageNumber="86">Fig. 10A</figureCitation>
). As described for
<taxonomicName id="4C354D71FFE7FFB30D38DBC1FD58FA71" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[585,689,1433,1458]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="86" phylum="Arthropoda" rank="species" species="ingens" status="sp. nov.">
<emphasis id="B941EAE0FFE7FFB30D38DBC1FD58FA71" box="[585,689,1433,1458]" italics="true" pageId="17" pageNumber="86">P. ingens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE7FFB30DC9DBC1FCFAFA70" box="[696,787,1433,1459]" pageId="17" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
, except as follows: length
<quantity id="4CCD9B17FFE7FFB30B33DBC0FB39FA70" box="[1090,1232,1432,1459]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="17" pageNumber="86" unit="mm" value="3.75" valueMax="3.8" valueMin="3.7">3.7–3.8 mm</quantity>
. Frontal width 2.4–2.5 ×interfrontal height. Gena with 11–14 setulae. First postsutural pair of dorsocentral bristles separated by 9–10 rows of fine acrostichal setulae. Anepisternum with posterodorsal patch of 5–6 setae and 10–12 setulae. Wing pattern (
<figureCitation id="130E2A77FFE7FFB30D39D85AFD59F9DE" box="[584,688,1538,1565]" captionStart="Fig" captionStartId="25.[188,231,1926,1952]" captionTargetBox="[189,1399,540,1879]" captionTargetId="figure-119@25.[189,1399,540,1890]" captionTargetPageId="25" captionText="Fig. 15. A. Pleuroseta wentworthi (Richards, 1973) holotype, ♂, habitus, lateral view. B. Pleuroseta dorrigonis (Richards, 1973) holotype, ♂, habitus, lateral view. Scale bar: A–B =1.0 mm. Photo credit: N. Tees and R. Cox (AMSA)." figureDoi="http://doi.org/10.5281/zenodo.10306530" httpUri="https://zenodo.org/record/10306530/files/figure.png" pageId="17" pageNumber="86">Fig. 15B</figureCitation>
) slightly paler than in
<taxonomicName id="4C354D71FFE7FFB30CCBD85BFBCDF9DF" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[954,1060,1539,1564]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="86" phylum="Arthropoda" rank="species" species="ingens">
<emphasis id="B941EAE0FFE7FFB30CCBD85BFBCDF9DF" box="[954,1060,1539,1564]" italics="true" pageId="17" pageNumber="86">P. ingens</emphasis>
</taxonomicName>
. CS2 1.3×length of CS3.
</paragraph>
<paragraph id="8B8A36F2FFE7FFB40FCCD81CFD33FE35" blockId="17.[187,1401,1603,2017]" lastBlockId="22.[188,1399,265,502]" lastPageId="22" lastPageNumber="91" pageId="17" pageNumber="86">
<smallCapsWord id="8D6CA02EFFE7FFB30FCCD81CFF16F99F" baselines="1623,1623" box="[189,255,1604,1630]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="17" pageNumber="86">MALE</smallCapsWord>
<smallCapsWord id="8D6CA02EFFE7FFB30E78D811FE98F99F" baselines="1623" box="[265,369,1609,1628]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="17" pageNumber="86">ABDOMEN</smallCapsWord>
(
<figureCitation id="130E2A77FFE7FFB30EF2D81BFE0CF99D" box="[387,485,1603,1630]" captionStart="Fig" captionStartId="10.[189,230,1857,1883]" captionTargetBox="[243,1343,265,1815]" captionTargetId="figure-65@10.[243,1343,265,1820]" captionTargetPageId="10" captionText="Fig. 5. A. Pleuroseta occidentalis sp. nov. paratype, left wing, ventral view (debu00244165). B. Pleuroseta occidentalis paratype, hypandrium, dorsal view (debu00245797). C. Pleuroseta wentworthi (Richards, 1973), hypandrium, dorsal view (debu00264733). D. Pleuroseta ingens sp. nov. paratype, hypandrium, dorsal view (debu00160984). E. Pleuroseta monteithi sp. nov. paratype, hypandrium, dorsal view (debu00160892). Scale bars: A=0.50 mm, B–E=0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.10306508" httpUri="https://zenodo.org/record/10306508/files/figure.png" pageId="17" pageNumber="86">Figs 5E</figureCitation>
,
<figureCitation id="130E2A77FFE7FFB30E85D81BFDA0F99E" box="[500,585,1603,1629]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[243,1340,265,1814]" captionTargetId="figure-54@18.[243,1343,265,1820]" captionTargetPageId="18" captionText="Fig. 10. Pleuroseta monteithi sp. nov. A. Male holotype habitus, lateral view (debu00160870). B. Male paratype terminalia, lateral view (debu00160892). C. Male paratype terminalia, posterolateral view (debu00160892). Abbreviations: epa =epandrium; hyp =hypandrium; se =subepandrial sclerite; ss =surstylus. Scale bars: A = 1.0 mm; B–C =0.40 mm." figureDoi="http://doi.org/10.5281/zenodo.10306520" httpUri="https://zenodo.org/record/10306520/files/figure.png" pageId="17" pageNumber="86">10B–C</figureCitation>
,
<figureCitation id="130E2A77FFE7FFB30D29D81BFD9EF99E" box="[600,631,1603,1629]" captionStart="Fig" captionStartId="19.[189,232,1928,1954]" captionTargetBox="[236,1343,278,1882]" captionTargetId="figure-1@19.[223,1365,272,1886]" captionTargetPageId="19" captionText="Fig. 11. Pleuroseta monteithi sp. nov. paratype (debu00160892). A. Aedeagus and associated structures, lateral view. B. Male S5–8, ventral view. Abbreviations: bp = basiphallus; dp =distiphallus; pg =postgonite; phal=phallapodeme; rs =ring sclerite. Scale bars: A=0.40 mm; B= 0.60 mm." figureDoi="http://doi.org/10.5281/zenodo.10306522" httpUri="https://zenodo.org/record/10306522/files/figure.png" pageId="17" pageNumber="86">11</figureCitation>
). S5 broad, subrectangular with large, triangular, posteromedial emargination, posterior surface of each lateral lobe with a dense tuft of fine, curved setae. S6+7 large (almost subequal to epandrium) and complex: S6 extending ventrally under (dorsal to) S5 and with dark, T-shaped posterior extension, the apex of which has a rippled margin; dorsal to this lies a tubular, basally constricted sclerite with heavily sclerotized, flattened, disc-like apex; S7 with elongate, dextral extension reaching across to ring sclerite; S8 with dorsal, anteromedial semicircular emargination; ring sclerite well-developed. Epandrium very large (only slightly smaller than head); cercus indistinct from epandrium; subepandrial sclerite thick, X-shaped. Hypandrium triangular, anteromedial apodeme curved upwards anteriorly. Surstylus short and complex: anterior half composed to two lobes: a semicircular dorsal lobe which is bent inwards anterodorsally and a thick, triangular, beak-like, ventrally setose ventral lobe; posterior half subtriangular, bulging laterally, with a broad anterior point, an elongate posterior ‘arm’, and a broad, rounded ventral lobe which has many long setae on the outer surface and a patch of thickened setae along the inner surface. Postgonite elongate, triangular and slightly bent forwards apically into a small hook. Phallapodeme elongate and curved; basiphallus large and saddle-shaped (often visible between surstyli in dried specimens). Distiphallus extremely large (~11% total body length), divided into basal and apical sections: basal section elongate, sclerotized and curved, with small lower knob articulating with apical section and spinose, membranous upper apex; apical section composed of a bent, trough-like, sinuate lower sclerite and a pair of flattened, curved lateral sclerites which fit along inner surface of lower sclerite.
</paragraph>
<caption id="DF4A667AFFE4FFB00FCCD93CFCD2F82A" ID-DOI="http://doi.org/10.5281/zenodo.10306520" ID-Zenodo-Dep="10306520" httpUri="https://zenodo.org/record/10306520/files/figure.png" pageId="18" pageNumber="87" startId="18.[189,232,1892,1918]" targetBox="[243,1340,265,1814]" targetPageId="18">
<paragraph id="8B8A36F2FFE4FFB00FCCD93CFCD2F82A" blockId="18.[185,1398,1892,2025]" pageId="18" pageNumber="87">
<emphasis id="B941EAE0FFE4FFB00FCCD93CFEF3F8BC" bold="true" box="[189,282,1892,1919]" pageId="18" pageNumber="87">Fig. 10.</emphasis>
<taxonomicName id="4C354D71FFE4FFB00E50D93DFDFFF8BC" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[289,534,1893,1919]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B941EAE0FFE4FFB00E50D93DFDFFF8BC" box="[289,534,1893,1919]" italics="true" pageId="18" pageNumber="87">Pleuroseta monteithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE4FFB00D6CD93DFD9EF8BC" box="[541,631,1893,1919]" pageId="18" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B941EAE0FFE4FFB00D0FD93CFD7AF8BD" bold="true" box="[638,659,1892,1918]" pageId="18" pageNumber="87">A</emphasis>
. Male holotype habitus, lateral view (debu00160870).
<emphasis id="B941EAE0FFE4FFB00A69D93DFAC5F8BC" bold="true" box="[1304,1324,1893,1919]" pageId="18" pageNumber="87">B</emphasis>
. Male paratype terminalia, lateral view (debu00160892).
<emphasis id="B941EAE0FFE4FFB00C56D9D0FCD5F861" bold="true" box="[807,828,1928,1954]" pageId="18" pageNumber="87">C</emphasis>
. Male paratype terminalia, posterolateral view (debu00160892). Abbreviations: epa =epandrium; hyp =hypandrium; se =subepandrial sclerite; ss =surstylus. Scale bars: A = 1.0 mm; B–C =0.40 mm.
</paragraph>
</caption>
<caption id="DF4A667AFFE5FFB10FCCD9D0FAEFF82A" ID-DOI="http://doi.org/10.5281/zenodo.10306522" ID-Zenodo-Dep="10306522" httpUri="https://zenodo.org/record/10306522/files/figure.png" pageId="19" pageNumber="88" startId="19.[189,232,1928,1954]" targetBox="[236,1343,278,1882]" targetPageId="19">
<paragraph id="8B8A36F2FFE5FFB10FCCD9D0FAEFF82A" blockId="19.[189,1399,1928,2025]" pageId="19" pageNumber="88">
<emphasis id="B941EAE0FFE5FFB10FCCD9D0FECAF861" bold="true" box="[189,291,1928,1954]" pageId="19" pageNumber="88">Fig. 11.</emphasis>
<taxonomicName id="4C354D71FFE5FFB10E44D9D0FDDDF861" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[309,564,1928,1954]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="88" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B941EAE0FFE5FFB10E44D9D0FDDDF861" box="[309,564,1928,1954]" italics="true" pageId="19" pageNumber="88">Pleuroseta monteithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE5FFB10D34D9D0FD42F861" box="[581,683,1928,1954]" pageId="19" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
paratype (debu00160892).
<emphasis id="B941EAE0FFE5FFB10B7FD9D0FBCAF861" bold="true" box="[1038,1059,1928,1954]" pageId="19" pageNumber="88">A</emphasis>
. Aedeagus and associated structures, lateral view.
<emphasis id="B941EAE0FFE5FFB10EAAD9F3FE04F806" bold="true" box="[475,493,1963,1989]" pageId="19" pageNumber="88">B</emphasis>
. Male S5–8, ventral view. Abbreviations: bp = basiphallus; dp =distiphallus; pg =postgonite; phal=phallapodeme; rs =ring sclerite. Scale bars: A=0.40 mm; B= 0.60 mm.
</paragraph>
</caption>
<caption id="DF4A667AFFE2FFB60FCCD93CFDE6F82A" ID-DOI="http://doi.org/10.5281/zenodo.10306524" ID-Zenodo-Dep="10306524" httpUri="https://zenodo.org/record/10306524/files/figure.png" pageId="20" pageNumber="89" startId="20.[189,232,1892,1918]" targetBox="[238,1349,265,1832]" targetPageId="20">
<paragraph id="8B8A36F2FFE2FFB60FCCD93CFDE6F82A" blockId="20.[185,1399,1892,2025]" pageId="20" pageNumber="89">
<emphasis id="B941EAE0FFE2FFB60FCCD93CFEFEF8BC" bold="true" box="[189,279,1892,1919]" pageId="20" pageNumber="89">Fig. 12.</emphasis>
<taxonomicName id="4C354D71FFE2FFB60E6AD93DFD19F8BC" authority="(Richards, 1973)" authorityName="NSW, QLD" baseAuthorityName="Richards" baseAuthorityYear="1973" box="[283,752,1893,1919]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="89" phylum="Arthropoda" rank="species" species="wentworthi">
<emphasis id="B941EAE0FFE2FFB60E6AD93DFDCDF8BC" box="[283,548,1893,1919]" italics="true" pageId="20" pageNumber="89">Pleuroseta wentworthi</emphasis>
(
<bibRefCitation id="EFA44B03FFE2FFB60D40D93DFD0EF8BC" author="Richards O. W." box="[561,743,1893,1919]" pageId="20" pageNumber="89" pagination="297 - 401" refId="ref10624" refString="Richards O. W. 1973. The Sphaeroceridae (= Borboridae or Cypselidae; Diptera Cyclorrhapha) of the Australasian region. Australian Journal of Zoology Supplemental Series 22: 297 - 401." type="book chapter" year="1973">Richards, 1973</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B941EAE0FFE2FFB60D86D93CFCE5F8BD" bold="true" box="[759,780,1892,1918]" pageId="20" pageNumber="89">A</emphasis>
. Male habitus, lateral view (debu00403539).
<emphasis id="B941EAE0FFE2FFB60A69D93DFAC5F8BC" bold="true" box="[1304,1324,1893,1919]" pageId="20" pageNumber="89">B</emphasis>
. Male terminalia, lateral view (debu00264733).
<emphasis id="B941EAE0FFE2FFB60DCDD9D0FD38F861" bold="true" box="[700,721,1928,1954]" pageId="20" pageNumber="89">C</emphasis>
. Male terminalia, posterolateral view (debu00264733). Abbreviations: epa =epandrium; hyp=hypandrium; se = subepandrial sclerite; ss =surstylus. Scale bars: A =1.0 mm; B–C =0.30 mm.
</paragraph>
</caption>
<caption id="DF4A667AFFE3FFB70FCCD9D0FB1AF82A" ID-DOI="http://doi.org/10.5281/zenodo.10306526" ID-Zenodo-Dep="10306526" httpUri="https://zenodo.org/record/10306526/files/figure.png" pageId="21" pageNumber="90" startId="21.[189,232,1928,1954]" targetBox="[236,1351,268,1875]" targetPageId="21">
<paragraph id="8B8A36F2FFE3FFB70FCCD9D0FB1AF82A" blockId="21.[189,1399,1928,2025]" pageId="21" pageNumber="90">
<emphasis id="B941EAE0FFE3FFB70FCCD9D0FECDF861" bold="true" box="[189,292,1928,1954]" pageId="21" pageNumber="90">Fig. 13.</emphasis>
<taxonomicName id="4C354D71FFE3FFB70E44D9D0FCCEF861" authority="(Richards, 1973)" authorityName="NSW, QLD" baseAuthorityName="Richards" baseAuthorityYear="1973" box="[309,807,1928,1954]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="90" phylum="Arthropoda" rank="species" species="wentworthi">
<emphasis id="B941EAE0FFE3FFB70E44D9D0FDA1F861" box="[309,584,1928,1954]" italics="true" pageId="21" pageNumber="90">Pleuroseta wentworthi</emphasis>
(
<bibRefCitation id="EFA44B03FFE3FFB70D10D9D0FCF6F861" author="Richards O. W." box="[609,799,1928,1954]" pageId="21" pageNumber="90" pagination="297 - 401" refId="ref10624" refString="Richards O. W. 1973. The Sphaeroceridae (= Borboridae or Cypselidae; Diptera Cyclorrhapha) of the Australasian region. Australian Journal of Zoology Supplemental Series 22: 297 - 401." type="book chapter" year="1973">Richards, 1973</bibRefCitation>
)
</taxonomicName>
(debu00264733).
<emphasis id="B941EAE0FFE3FFB70B63D9D0FBCEF861" bold="true" box="[1042,1063,1928,1954]" pageId="21" pageNumber="90">A</emphasis>
. Aedeagus and associated structures, lateral view.
<emphasis id="B941EAE0FFE3FFB70EAAD9F3FE04F806" bold="true" box="[475,493,1963,1989]" pageId="21" pageNumber="90">B</emphasis>
. Male S5–8, ventral view. Abbreviations: bp = basiphallus; dp =distiphallus; ea= ejaculatory apodeme; pg =postgonite; phal=phallapodeme. Scale bars: A–B =0.30 mm.
</paragraph>
</caption>
<paragraph id="8B8A36F2FFE0FFB40FCCDC46FD8CFDFB" blockId="22.[189,613,542,568]" box="[189,613,542,568]" pageId="22" pageNumber="91">
<smallCapsWord id="8D6CA02EFFE0FFB40FCCDC46FEFDFDF5" baselines="561,561" box="[189,276,542,568]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="22" pageNumber="91">FEMALE</smallCapsWord>
<smallCapsWord id="8D6CA02EFFE0FFB40E6ADC7BFE6AFDF5" baselines="561" box="[283,387,547,566]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="abdomen" pageId="22" pageNumber="91">ABDOMEN</smallCapsWord>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32F6579FFE0FFB40FCCDC07FBC9FC1C" pageId="22" pageNumber="91" type="discussion">
<paragraph id="8B8A36F2FFE0FFB40FCCDC07FEC6FDBA" blockId="22.[189,303,607,633]" box="[189,303,607,633]" pageId="22" pageNumber="91">
<heading id="D0C2819EFFE0FFB40FCCDC07FEC6FDBA" bold="true" box="[189,303,607,633]" fontSize="11" level="3" pageId="22" pageNumber="91" reason="3">
<emphasis id="B941EAE0FFE0FFB40FCCDC07FEC6FDBA" bold="true" box="[189,303,607,633]" pageId="22" pageNumber="91">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A36F2FFE0FFB40FCCDCD5FE06FC94" blockId="22.[189,1399,653,855]" pageId="22" pageNumber="91">
<taxonomicName id="4C354D71FFE0FFB40FCCDCD5FE58FD64" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[189,433,653,679]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="91" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B941EAE0FFE0FFB40FCCDCD5FE58FD64" box="[189,433,653,679]" italics="true" pageId="22" pageNumber="91">Pleuroseta monteithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE0FFB40EC9DCD6FDFBFD6B" box="[440,530,654,680]" pageId="22" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
is externally almost indistinguishable from, and internally very similar to,
<taxonomicName id="4C354D71FFE0FFB40FCCDCEAFECEFD08" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[189,295,690,715]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="91" phylum="Arthropoda" rank="species" species="ingens" status="sp. nov.">
<emphasis id="B941EAE0FFE0FFB40FCCDCEAFECEFD08" box="[189,295,690,715]" italics="true" pageId="22" pageNumber="91">P. ingens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE0FFB40E41DCE9FE62FD08" box="[304,395,689,715]" pageId="22" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
As such, the male genitalia are required to identify this species: S5 with a triangular posteromedial emargination, S6 with a flattened, T-shaped posterior extension, anterior lobe of surstylus with a more elongate triangular ventral lobe and a semicircular dorsal lobe, postgonite thicker and straighter along the anterior margin, and basal part of the distiphallus more evenly curved without an elongate apical extension.
</paragraph>
<paragraph id="8B8A36F2FFE0FFB40FCCDD27FBC9FC1C" blockId="22.[189,1399,895,992]" pageId="22" pageNumber="91">
Both known specimens of
<taxonomicName id="4C354D71FFE0FFB40D79DD27FCE8FC5A" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[520,769,895,921]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="91" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B941EAE0FFE0FFB40D79DD27FCE8FC5A" box="[520,769,895,921]" italics="true" pageId="22" pageNumber="91">Pleuroseta monteithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE0FFB40C7DDDD8FC85FC59" box="[780,876,896,922]" pageId="22" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
were collected some
<quantity id="4CCD9B17FFE0FFB40B0ADD27FB31FC5A" box="[1147,1240,895,922]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="91" unit="km" value="150.0">150 km</quantity>
north of the northern-most collection locality of the similar
<taxonomicName id="4C354D71FFE0FFB40D8ADDFBFC81FC7F" authority="Kuwahara &amp; Marshall, 2023" authorityName="Kuwahara &amp; Marshall" authorityYear="2023" box="[763,872,931,956]" class="Insecta" family="Sphaeroceridae" genus="Pleuroseta" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="91" phylum="Arthropoda" rank="species" species="ingens" status="sp. nov.">
<emphasis id="B941EAE0FFE0FFB40D8ADDFBFC81FC7F" box="[763,872,931,956]" italics="true" pageId="22" pageNumber="91">P. ingens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272579BFFE0FFB40C03DDFBFC39FC7E" box="[882,976,931,957]" pageId="22" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
(from Mount Hypipamee National Park), suggesting that they might be allopatric or parapatric sister species.
</paragraph>
</subSubSection>
</treatment>
</document>
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