132 lines
16 KiB
XML
132 lines
16 KiB
XML
<document id="8B165E5782F4F5BE34481751DD28ADEF" ID-CLB-Dataset="32201" ID-DOI="10.11646/zootaxa.4317.3.4" ID-GBIF-Dataset="ca0d653d-32de-4c75-a95d-d9a4690d6fda" ID-ISSN="1175-5326" ID-Zenodo-Dep="884766" ID-ZooBank="5Fd492A4-D41D-4F37-A121-Ffa680E7E778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504596385874" checkinUser="plazi" docAuthor="Fery, Hans" docDate="2017" docId="039C87CE0F30FF92FF1F6601CAB05606" docLanguage="en" docName="zootaxa.4317.3.4.pdf" docOrigin="Zootaxa 4317 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hygrotus sensu Stephens 1828" docType="treatment" docVersion="7" lastPageNumber="518" masterDocId="FFA5FFB60F23FF81FF886361C8305140" masterDocTitle="A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)" masterLastPageNumber="529" masterPageNumber="499" pageNumber="518" updateTime="1698467863718" updateUser="ExternalLinkService">
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<mods:title id="4331781736FD6CE3407AD67122EC6C2D">A new classification of the tribe Hygrotini Portevin, 1929 (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
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<mods:name id="EC999365CE159EAFFDA95B77A1408086" type="personal">
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<mods:namePart id="FC0589967EC5C82C9713F3861AB683E6">Fery, Hans</mods:namePart>
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<mods:typeOfResource id="1E6FEEA19980ED3235F4579F1F245576">text</mods:typeOfResource>
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<mods:title id="2AC128B80454CFC1CE719C3A5B41996E">Zootaxa</mods:title>
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<mods:part id="DB75338757170C6E8D3BF1512CEBCBBF">
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<mods:date id="BCECFC63D06F5AFA1D89119E5CB49072">2017</mods:date>
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<mods:number id="2B533D251F59D16707D6BCAA351B552F">2017-05-05</mods:number>
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<mods:number id="F29AE29BC024796E32B2739B92F437AB">4317</mods:number>
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<mods:end id="3ABA5A21BA5B82678DB6580764733502">529</mods:end>
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<mods:identifier id="5EE48C48F46034B0A447034F16D089BA" type="DOI">10.11646/zootaxa.4317.3.4</mods:identifier>
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<treatment id="039C87CE0F30FF92FF1F6601CAB05606" ID-DOI="http://doi.org/10.5281/zenodo.6026851" ID-GBIF-Taxon="133183478" ID-Zenodo-Dep="6026851" LSID="urn:lsid:plazi:treatment:039C87CE0F30FF92FF1F6601CAB05606" httpUri="http://treatment.plazi.org/id/039C87CE0F30FF92FF1F6601CAB05606" lastPageNumber="518" pageId="19" pageNumber="518">
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<subSubSection id="C32F65530F30FF92FF1F6601CA2E543A" box="[151,542,1375,1402]" pageId="19" pageNumber="518" type="nomenclature">
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<paragraph id="8B8A36D80F30FF92FF1F6601CA2E543A" blockId="19.[151,542,1375,1402]" box="[151,542,1375,1402]" pageId="19" pageNumber="518">
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<heading id="D0C281B40F30FF92FF1F6601CA2E543A" bold="true" box="[151,542,1375,1402]" fontSize="11" level="1" pageId="19" pageNumber="518" reason="1">
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<emphasis id="B941EACA0F30FF92FF1F6601CA2E543A" bold="true" box="[151,542,1375,1402]" pageId="19" pageNumber="518">
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Genus
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<taxonomicName id="4C354D5B0F30FF92FF65663ECA2E543A" ID-CoL="4ZYJ" authority="Stephens, 1828" authorityName="sensu Stephens" authorityYear="1828" box="[237,542,1375,1402]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EACA0F30FF92FF65663EC96D5439" bold="true" box="[237,349,1375,1401]" italics="true" pageId="19" pageNumber="518">Hygrotus</emphasis>
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<bibRefCitation id="EFA44B290F30FF92FEED6601CA2E543A" author="Stephens" box="[357,542,1376,1402]" pageId="19" pageNumber="518" refString="Stephens, J. F. (1828 - 1829) Illustrations of British entomology. Mandibulata. Vol. 2. Baldwin & Cradock, London, 200 pp. [pp. 1 - 112 in 1828, pp. 113 - 200 in 1829]" type="book chapter" year="1828">Stephens, 1828</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F65530F30FF92FF1F66C7CD5D570A" pageId="19" pageNumber="518" type="materials_examined">
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<paragraph id="8B8A36D80F30FF92FF1F66C7CCD454FC" blockId="19.[151,1252,1446,1469]" box="[151,1252,1446,1469]" pageId="19" pageNumber="518">
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<emphasis id="B941EACA0F30FF92FF1F66C7C92C54FC" bold="true" box="[151,284,1446,1468]" pageId="19" pageNumber="518">Type species</emphasis>
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:
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<taxonomicName id="4C354D5B0F30FF92FEA266C9CAFA54FC" authority="Fabricius, 1777: 239" authorityName="Fabricius" authorityPageNumber="239" authorityYear="1777" box="[298,714,1446,1469]" class="Insecta" family="Dytiscidae" genus="Dytiscus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="species" species="inaequalis">
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<emphasis id="B941EACA0F30FF92FEA266C9C9C154FC" box="[298,497,1446,1469]" italics="true" pageId="19" pageNumber="518">Dytiscus inaequalis</emphasis>
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<bibRefCitation id="EFA44B290F30FF92FE7F66C7CAA854FC" author="Fabricius" box="[503,664,1446,1468]" pageId="19" pageNumber="518" refString="Fabricius, J. C. (1777) Genera Insectorum eorumque characteres naturales secundum numerum, figuram, situm et proportionem omnium partium oris adiecta mantissa speciorum nuper detectarum. M. F. Bartschii, Chilonii, xvi + 310 pp." type="book" year="1777">Fabricius, 1777</bibRefCitation>
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: 239
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</taxonomicName>
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, by subsequent designation of
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<bibRefCitation id="EFA44B290F30FF92FB8A66C7CCD054FC" author="Curtis" box="[1026,1248,1446,1468]" pageId="19" pageNumber="518" refString="Curtis, J. (1835) British entomology; being illustrations and descriptions of the genera of insects found in Great Britain and Ireland. Vol. 12. J. Curtis, London, pls. 530 - 577." type="book chapter" year="1835">Curtis (1835: pl. 531)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8B8A36D80F30FF92FF1F6688CD5D570A" blockId="19.[151,1436,1513,1610]" pageId="19" pageNumber="518">
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<materialsCitation id="3B5D3C850F30FF92FF1F6688CD5D570A" ID-GBIF-Occurrence="1608130918" country="Mexico" location="Pacific region" pageId="19" pageNumber="518" specimenCount="1">
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The genus
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<taxonomicName id="4C354D5B0F30FF92FE95668BC9B45741" authorityName="sensu Stephens" authorityYear="1828" box="[285,388,1514,1537]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EACA0F30FF92FE95668BC9B45741" box="[285,388,1514,1537]" italics="true" pageId="19" pageNumber="518">Hygrotus</emphasis>
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</taxonomicName>
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as here defined includes 129 described species (two of them bitypic) distributed in the Nearctic (including the north of
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<collectingCountry id="F32276480F30FF92FD84656CCA575766" box="[524,615,1549,1574]" name="Mexico" pageId="19" pageNumber="518">MeXico</collectingCountry>
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), Palaearctic, Afrotropical and Oriental zoogeographical regions (
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<figureCitation id="130E2A5D0F30FF92FAD0656CCDAC5766" box="[1368,1436,1549,1574]" captionStart="FIGURE 2" captionStartId="12.[151,250,1941,1963]" captionTargetBox="[208,1378,197,1919]" captionTargetId="figure@12.[189,1396,193,1921]" captionTargetPageId="12" captionText="FIGURE 2. Dendrogram showing the new classification of Hygrotini. A number of species without molecular data (no voucher number) have been inserted near morphologically similar species with molecular data. Acronyms refer to biogeographical regions." httpUri="https://zenodo.org/record/884770/files/figure.png" pageId="19" pageNumber="518">Fig. 2</figureCitation>
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and
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<tableCitation id="C6B703630F30FF92FF4E6550C928570A" box="[198,280,1585,1610]" captionStart="TABLE 1" captionStartId="5.[125,200,512,532]" captionText="TABLE 1. Checklist of all 137 species of Hygrotini with original, new and former (Nilsson & Há ek 2017 a) generic and subgeneric names; last three columns indicating new combinations, distribution and methods applied to material studied. See Villastrigo et al. (in press) for details on the specimens used in the molecular phylogeny and accession numbers of the sequences." pageId="19" pageNumber="518">Table 1</tableCitation>
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). One Nearctic species was introduced to Hawaii (see below), which belongs to the
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<location id="8EEA60030F30FF92FB416550CD59570A" LSID="urn:lsid:plazi:treatment:039C87CE0F30FF92FF1F6601CAB05606:8EEA60030F30FF92FB416550CD59570A" box="[1225,1385,1585,1610]" country="Mexico" name="Pacific region" pageId="19" pageNumber="518">Pacific region</location>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F65530F30FF92FF4F6518CAB05606" pageId="19" pageNumber="518" type="diagnosis">
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<paragraph id="8B8A36D80F30FF92FF4F6518CCEB57F6" blockId="19.[151,1436,1657,1862]" pageId="19" pageNumber="518">
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<emphasis id="B941EACA0F30FF92FF4F6518C97157D2" bold="true" box="[199,321,1657,1682]" pageId="19" pageNumber="518">Diagnosis.</emphasis>
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In the new classification
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<taxonomicName id="4C354D5B0F30FF92FDF3651BCAD257D1" authorityName="sensu Stephens" authorityYear="1828" box="[635,738,1658,1681]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EACA0F30FF92FDF3651BCAD257D1" box="[635,738,1658,1681]" italics="true" pageId="19" pageNumber="518">Hygrotus</emphasis>
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</taxonomicName>
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is the second of two genera of the tribe
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<taxonomicName id="4C354D5B0F30FF92FB526518CD7B57D2" authorityName="Portevin" authorityYear="1929" box="[1242,1355,1657,1682]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="tribe" tribe="Hygrotini">Hygrotini</taxonomicName>
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. As in
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<taxonomicName id="4C354D5B0F30FF92FF1F65FCC93157F6" authority="Fery, 2017" authorityName="Fery" authorityYear="2017" box="[151,257,1693,1718]" class="Insecta" family="Dytiscidae" genus="Clemnius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B941EACA0F30FF92FF1F65FCC93157F6" box="[151,257,1693,1718]" italics="true" pageId="19" pageNumber="518">Clemnius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27257B10F30FF92FE8F65FCC96657F5" box="[263,342,1693,1718]" pageId="19" pageNumber="518" rank="genus">
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<emphasis id="B941EACA0F30FF92FE8F65FCC97F57F6" bold="true" box="[263,335,1693,1718]" pageId="19" pageNumber="518">n. gen</emphasis>
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.
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</taxonomicNameLabel>
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all species of the genus have two carinae in each antennal cavity (see
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<figureCitation id="130E2A5D0F30FF92FBE165FCCCFE57F6" box="[1129,1230,1693,1718]" captionStart-0="FIGURES 3 – 4" captionStart-1="FIGURES 5 – 6" captionStartId-0="13.[151,264,1658,1680]" captionStartId-1="14.[151,264,1930,1952]" captionTargetBox-0="[353,1132,193,1614]" captionTargetBox-1="[264,1323,403,1909]" captionTargetId-0="figure@13.[353,1233,193,1614]" captionTargetId-1="figure@14.[264,1323,403,1909]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 3 – 4. Antennal cavities of: (3) Hydroporus dorsalis (Hydroporini) and (4) Hygrotus (Leptolambus) lagari (Fery, 1992) (a: carina delimiting capsule for scape; b: fronto-lateral border of clypeus; c: second carina)." captionText-1="FIGURES 5 – 6. SEM micrographs with details of head and antennal cavities of: (5) Hygrotus (Leptolambus) impressopunctatus (frontal view) and (6) Hygrotus (s. str.) inaequalis (oblique ventral view) (a, b and c as in Figs 3 – 4; d: clypeal stripe; e: labrum)." httpUri-0="https://zenodo.org/record/884772/files/figure.png" httpUri-1="https://zenodo.org/record/884774/files/figure.png" pageId="19" pageNumber="518">Figs 4–6</figureCitation>
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).
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</paragraph>
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<paragraph id="8B8A36D80F30FF92FF4F65A0CAB05606" blockId="19.[151,1436,1657,1862]" pageId="19" pageNumber="518">
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According to the two main clades in the phylogeny of
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<taxonomicName id="4C354D5B0F30FF92FCCF65A0CB84579A" authorityName="Portevin" authorityYear="1929" box="[839,948,1729,1754]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="tribe" tribe="Hygrotini">Hygrotini</taxonomicName>
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and several morphological characters, the genus (which corresponds to Clade B in
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<figureCitation id="130E2A5D0F30FF92FDEA6584CA9257BE" box="[610,674,1765,1790]" captionStart="FIGURE 1" captionStartId="11.[151,250,1975,1997]" captionTargetBox="[188,1357,201,1928]" captionTargetId="figure@11.[185,1388,195,1940]" captionTargetPageId="11" captionText="FIGURE 1. Phylogeny of Hygrotini, modified from Villastrigo et al. (in press). Numbers above nodes are bootstrap support values. Note that here the former generic and subgeneric names are still used." httpUri="https://zenodo.org/record/884768/files/figure.png" pageId="19" pageNumber="518">Fig. 1</figureCitation>
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) is subdivided into four clades corresponding to the following four subgenera: clade B1 = subgenus
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<taxonomicName id="4C354D5B0F30FF92FD846468CABA5662" box="[524,650,1801,1826]" class="Insecta" family="Dytiscidae" genus="Coelambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EACA0F30FF92FD846468CABA5662" box="[524,650,1801,1826]" italics="true" pageId="19" pageNumber="518">Coelambus</emphasis>
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</taxonomicName>
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, clade B2 = subgenus
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<taxonomicName id="4C354D5B0F30FF92FC056468CC285662" box="[909,1048,1801,1826]" class="Insecta" family="Dytiscidae" genus="Hyphoporus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus">
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<emphasis id="B941EACA0F30FF92FC056468CC285662" box="[909,1048,1801,1826]" italics="true" pageId="19" pageNumber="518">Hyphoporus</emphasis>
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</taxonomicName>
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, clade B3 = subgenus
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<taxonomicName id="4C354D5B0F30FF92FA93646BCDB25661" authority="Fery, 2017" authorityName="Fery" authorityYear="2017" box="[1307,1410,1802,1825]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="518" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B941EACA0F30FF92FA93646BCDB25661" box="[1307,1410,1802,1825]" italics="true" pageId="19" pageNumber="518">Hygrotus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27257B10F30FF92FA02646BC8895605" pageId="19" pageNumber="518" sensu="stricto">s. str.</taxonomicNameLabel>
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and clade B4 =
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<emphasis id="B941EACA0F30FF92FEE7644CCA305606" box="[367,512,1837,1862]" italics="true" pageId="19" pageNumber="518">Leptolambus</emphasis>
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<emphasis id="B941EACA0F30FF92FD8F644CCAB05606" bold="true" box="[519,640,1837,1862]" pageId="19" pageNumber="518">n. subgen.</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |