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205 lines
24 KiB
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<document id="DFD4C15CEB05967B0F29A7340658B4E0" ID-CLB-Dataset="3167" ID-DOI="10.1111/j.1096-3642.2008.00494.x" ID-GBIF-Dataset="d59ef689-6666-421e-8e46-c3fa65865db4" ID-ISSN="0024-4082" ID-Zenodo-Dep="4687752" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618409491744" checkinUser="carolina" docAuthor="Antoñanzas, Raquel López" docDate="2009" docId="039C6B6DFF86FFAFD5DCC5BD4ADBF915" docLanguage="en" docName="ZoolJLinnSoc.156.664-679.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 156 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2008.00494.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Potwarmus primitivus" docType="treatment" docVersion="8" lastPageNumber="668" masterDocId="FFA51315FF82FFABD561C34B4E30FFA2" masterDocTitle="First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea)" masterLastPageNumber="679" masterPageNumber="664" pageNumber="668" updateTime="1698936486494" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="0E672ED18ACB0712131BE271589B3A4B">First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea)</mods:title>
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<mods:namePart id="DE87ED9DD388B740CF8033C117F7E90B">Antoñanzas, Raquel López</mods:namePart>
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<mods:typeOfResource id="CAA7062532EC11BDD34C7ED1233365D7">text</mods:typeOfResource>
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<mods:title id="3FC0EA163A91C724E1280A3E84D1FE78">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="72C19D634A84DBE5A82930B5430F5C41">
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<mods:date id="8BA0229DDCCC57EA3BAE274EC5EBF7CC">2009</mods:date>
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<mods:number id="2C9CA3B492B4979F166AEA0266F02E7C">2009-07-31</mods:number>
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<mods:number id="EFB7ABE978A1E9E0C3E9E1170C7DD85F">156</mods:number>
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<mods:classification id="F23E7DC8AAE343F7336F91594F04F91A">journal article</mods:classification>
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<mods:identifier id="331EE8FE273DD60A9FFCC915302FAC8A" type="CLB-Dataset">3167</mods:identifier>
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<mods:identifier id="2F269D09DFCC612A48EA7EBFA687B201" type="DOI">10.1111/j.1096-3642.2008.00494.x</mods:identifier>
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<treatment id="039C6B6DFF86FFAFD5DCC5BD4ADBF915" ID-DOI="http://doi.org/10.5281/zenodo.5747601" ID-GBIF-Taxon="190875390" ID-Zenodo-Dep="5747601" LSID="urn:lsid:plazi:treatment:039C6B6DFF86FFAFD5DCC5BD4ADBF915" httpUri="http://treatment.plazi.org/id/039C6B6DFF86FFAFD5DCC5BD4ADBF915" lastPageNumber="668" pageId="4" pageNumber="668">
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<subSubSection id="C32F89F0FF86FFAFD5DCC5BD4CC2F8A9" box="[189,754,1781,1803]" pageId="4" pageNumber="668" type="nomenclature">
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<paragraph id="8B8ADA7BFF86FFAFD5DCC5BD4CC2F8A9" blockId="4.[189,754,1781,1803]" box="[189,754,1781,1803]" pageId="4" pageNumber="668">
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<heading id="D0C26D17FF86FFAFD5DCC5BD4CC2F8A9" allCaps="true" box="[189,754,1781,1803]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="668" reason="9">
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<taxonomicName id="4C35A1F8FF86FFAFD5DCC5BD4FE6F8AB" authority="(WESSELS ET AL., 1982)" box="[189,470,1782,1803]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
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<emphasis id="B9410669FF86FFAFD5DCC5BD4FE6F8AB" box="[189,470,1782,1803]" italics="true" pageId="4" pageNumber="668">POTWARMUS PRIMITIVUS</emphasis>
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</taxonomicName>
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(WESSELS
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<emphasis id="B9410669FF86FFAFD73BC5B24C90F8A9" box="[602,672,1782,1803]" italics="true" pageId="4" pageNumber="668">ET AL.</emphasis>
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, 1982)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F89F0FF86FFAFD5C5C4574CF0F893" box="[164,704,1820,1842]" pageId="4" pageNumber="668" type="reference_group">
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<paragraph id="8B8ADA7BFF86FFAFD5C5C4574CF0F893" blockId="4.[164,779,1820,1903]" box="[164,704,1820,1842]" pageId="4" pageNumber="668">
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<emphasis id="B9410669FF86FFAFD5C5C4574F24F893" box="[164,276,1820,1841]" italics="true" pageId="4" pageNumber="668">Synonym:</emphasis>
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<taxonomicName id="4C35A1F8FF86FFAFD47FC4574C8CF893" authority="Lindsay, 1988" authorityName="Lindsay" authorityYear="1988" box="[286,700,1820,1842]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="minimus">
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<emphasis id="B9410669FF86FFAFD47FC4574C3EF893" box="[286,526,1820,1841]" italics="true" pageId="4" pageNumber="668">Potwarmus minimus</emphasis>
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<bibRefCitation id="EFA4A78AFF86FFAFD779C4574C8CF893" author="Lindsay EH" box="[536,700,1820,1842]" pageId="4" pageNumber="668" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay, 1988</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F89F0FF86FFAFD5C5C4714B27FBF1" pageId="4" pageNumber="668" type="discussion">
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<paragraph id="8B8ADA7BFF86FFAFD5C5C4714B27FBF1" blockId="4.[164,779,1820,1903]" lastBlockId="4.[826,1442,198,1719]" pageId="4" pageNumber="668">
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<bibRefCitation id="EFA4A78AFF86FFAFD5C5C4714FBBF8F2" author="Wessels W & de Bruijn H & Hussain ST & Leinders J" box="[164,395,1850,1872]" pageId="4" pageNumber="668" pagination="337 - 364" refId="ref12338" refString="Wessels W, de Bruijn H, Hussain ST, Leinders J. 1982. Fossil rodents from the Chinji Formation, Banda Daud Shah, Kohat, Pakistan. Koninklijke Nederlandse Akademie van Wetenschappen Proceedings, Series B 85: 337 - 364." type="journal article" year="1982">
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Wessels
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<emphasis id="B9410669FF86FFAFD464C4704F0DF8ED" box="[261,317,1850,1872]" italics="true" pageId="4" pageNumber="668">et al.</emphasis>
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(1982)
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</bibRefCitation>
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erected the new species
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<taxonomicName id="4C35A1F8FF86FFAFD7C6C4714F2BF8CC" class="Mammalia" family="Muridae" genus="Antemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
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<emphasis id="B9410669FF86FFAFD7C6C4714F2BF8CC" italics="true" pageId="4" pageNumber="668">Antemus primitivus</emphasis>
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</taxonomicName>
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on the basis of eight complete and seven incomplete isolated cheek teeth from the Chinji Formation (near Banda Daud Shah),
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<collectingCountry id="F3229AEBFF86FFAFD18AC3AF4B64FF58" box="[1259,1364,228,250]" name="Pakistan" pageId="4" pageNumber="668">Pakistan</collectingCountry>
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. They assigned this taxon to the genus
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<taxonomicName id="4C35A1F8FF86FFAFD182C2484B77FEBA" authorityName="CHINJIENSIS JACOBS" authorityYear="1977" box="[1251,1351,259,280]" class="Mammalia" family="Muridae" genus="Antemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="genus">
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<emphasis id="B9410669FF86FFAFD182C2484B77FEBA" box="[1251,1351,259,280]" italics="true" pageId="4" pageNumber="668">Antemus</emphasis>
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</taxonomicName>
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. Later,
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<bibRefCitation id="EFA4A78AFF86FFAFD65BC2694A11FE9A" author="Jacobs LL & Flynn LJ & Downs WR & Barry JC" box="[826,1057,290,312]" pageId="4" pageNumber="668" pagination="157 - 177" refId="ref11169" refString="Jacobs LL, Flynn LJ, Downs WR, Barry JC. 1989. Neogene rodents of southern Asia. In: Black CC, Dawson MR, eds. Papers on fossil rodents in honor of Albert Elmer Wood. Los Angeles, CA: Natural History Museum of Los Angeles County, 157 - 177." type="book chapter" year="1989">
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Jacobs
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<emphasis id="B9410669FF86FFAFD6F5C2694DFCFE95" box="[916,972,290,311]" italics="true" pageId="4" pageNumber="668">et al.</emphasis>
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(1989)
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</bibRefCitation>
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excluded this species from the genus
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<taxonomicName id="4C35A1F8FF86FFAFD6E5C20A4DD8FEF4" authorityName="CHINJIENSIS JACOBS" authorityYear="1977" box="[900,1000,321,342]" class="Mammalia" family="Muridae" genus="Antemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="genus">
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<emphasis id="B9410669FF86FFAFD6E5C20A4DD8FEF4" box="[900,1000,321,342]" italics="true" pageId="4" pageNumber="668">Antemus</emphasis>
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</taxonomicName>
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because it has a single lingual cusp on the M1 (the enterostyle), whereas
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<taxonomicName id="4C35A1F8FF86FFAFD1A7C2144B6FFED6" authorityName="Jacobs" authorityYear="1977" box="[1222,1375,351,372]" class="Mammalia" family="Muridae" genus="Antemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="chinjiensis">
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<emphasis id="B9410669FF86FFAFD1A7C2144B6FFED6" box="[1222,1375,351,372]" italics="true" pageId="4" pageNumber="668">A. chinjiensis</emphasis>
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</taxonomicName>
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(type species of the genus) has two (an isolated enterostyle and an anterostyle, t1, connected to the lingual anterocone, t2). The reassessment conducted by
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<bibRefCitation id="EFA4A78AFF86FFAFD65BC2924A2BFE4D" author="Jacobs LL & Flynn LJ & Downs WR & Barry JC" box="[826,1051,473,495]" pageId="4" pageNumber="668" pagination="157 - 177" refId="ref11169" refString="Jacobs LL, Flynn LJ, Downs WR, Barry JC. 1989. Neogene rodents of southern Asia. In: Black CC, Dawson MR, eds. Papers on fossil rodents in honor of Albert Elmer Wood. Los Angeles, CA: Natural History Museum of Los Angeles County, 157 - 177." type="book chapter" year="1989">
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Jacobs
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<emphasis id="B9410669FF86FFAFD6F1C2914DF8FE4C" box="[912,968,473,495]" italics="true" pageId="4" pageNumber="668">et al.</emphasis>
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(1989)
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</bibRefCitation>
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, then in press, together with the study of additional material from the Middle Miocene Pakistani localities YGSP 589, YGSP 709 (Chinji Formation) and YGSP 591 and YGSP 592 (Kamlial Formation), motivated
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<bibRefCitation id="EFA4A78AFF86FFAFD144C11F4AE5FDC8" author="Lindsay EH" box="[1061,1237,596,618]" pageId="4" pageNumber="668" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay (1988)</bibRefCitation>
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to re-allocate
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<taxonomicName id="4C35A1F8FF86FFAFD0E8C11F4D81FD2A" class="Mammalia" family="Muridae" genus="Antemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
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<emphasis id="B9410669FF86FFAFD0E8C11F4D81FD2A" italics="true" pageId="4" pageNumber="668">A. primitivus</emphasis>
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</taxonomicName>
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to the new genus
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<taxonomicName id="4C35A1F8FF86FFAFD1FCC1384B2EFD2A" box="[1181,1310,627,648]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="genus">
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<emphasis id="B9410669FF86FFAFD1FCC1384B2EFD2A" box="[1181,1310,627,648]" italics="true" pageId="4" pageNumber="668">Potwarmus</emphasis>
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</taxonomicName>
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, an action with which I agree.
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<bibRefCitation id="EFA4A78AFF86FFAFD17FC1DA4AF8FD05" author="Lindsay EH" box="[1054,1224,657,679]" pageId="4" pageNumber="668" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay (1988)</bibRefCitation>
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also erected a new species of
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<taxonomicName id="4C35A1F8FF86FFAFD6CFC1FB4A1FFD67" box="[942,1071,688,709]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="genus">
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<emphasis id="B9410669FF86FFAFD6CFC1FB4A1FFD67" box="[942,1071,688,709]" italics="true" pageId="4" pageNumber="668">Potwarmus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C35A1F8FF86FFAFD15FC1FB4A8FFD67" authorityName="Lindsay" authorityYear="1988" box="[1086,1215,688,709]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="minimus">
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<emphasis id="B9410669FF86FFAFD15FC1FB4A8FFD67" box="[1086,1215,688,709]" italics="true" pageId="4" pageNumber="668">P. minimus</emphasis>
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</taxonomicName>
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, the
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<typeStatus id="548E64D9FF86FFAFD19AC1E44B6CFD67" box="[1275,1372,687,709]" pageId="4" pageNumber="668" type="holotype">holotype</typeStatus>
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being an isolated left m1 (YGSP 19555) from the Middle Miocene locality YGSP 589 (Chinji Formation, Potwar Plateau,
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<collectingCountry id="F3229AEBFF86FFAFD6C2C0404A3AFC83" box="[931,1034,779,801]" name="Pakistan" pageId="4" pageNumber="668">Pakistan</collectingCountry>
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) (
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<bibRefCitation id="EFA4A78AFF86FFAFD143C0404AF6FC82" author="Lindsay EH" box="[1058,1222,779,801]" pageId="4" pageNumber="668" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay, 1988</bibRefCitation>
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: pl. 10, fig. l). This species was said to be also recorded in
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<collectingCountry id="F3229AEBFF86FFAFD073C0614B4BFCE2" box="[1298,1403,810,832]" name="Pakistan" pageId="4" pageNumber="668">Pakistan</collectingCountry>
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, in the Middle Miocene locality YGSP
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<quantity id="4CCD779EFF86FFAFD1A8C0034B21FCFC" box="[1225,1297,840,862]" metricMagnitude="1" metricUnit="m" metricValue="1.63068" pageId="4" pageNumber="668" unit="in" value="642.0">642 in</quantity>
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the Kamlial Formation and, possibly, in some HGSP localities in the Chinji Formation at Banda Daud Shah (
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<bibRefCitation id="EFA4A78AFF86FFAFD020C0CD4D44FC18" author="Lindsay EH" pageId="4" pageNumber="668" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay, 1988</bibRefCitation>
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). However, based on analysis of a larger sample from a single locality,
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<bibRefCitation id="EFA4A78AFF86FFAFD15AC0884AD7FC7B" author="Lindsay EH" box="[1083,1255,963,985]" pageId="4" pageNumber="668" pagination="131 - 147" refId="ref11418" refString="Lindsay EH. 1994. The evolution of murine rodents in Asia. National Science Museum Monographs 8: 131 - 147." type="journal article" year="1994">Lindsay (1994)</bibRefCitation>
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synonymized
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<taxonomicName id="4C35A1F8FF86FFAFD0ECC0884D81FC55" class="Mammalia" family="Muridae" genus="Antemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
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<emphasis id="B9410669FF86FFAFD0ECC0884D81FC55" italics="true" pageId="4" pageNumber="668">P. primitivus</emphasis>
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</taxonomicName>
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and
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<emphasis id="B9410669FF86FFAFD160C0A94AA4FC55" box="[1025,1172,994,1015]" italics="true" pageId="4" pageNumber="668">
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<taxonomicName id="4C35A1F8FF86FFAFD160C0A94ABFFC55" authorityName="Lindsay" authorityYear="1988" box="[1025,1167,994,1015]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="minimus">P. minimus</taxonomicName>
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.
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</emphasis>
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Incidentally,
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<bibRefCitation id="EFA4A78AFF86FFAFD024C0AA4D4CFBB7" author="Lindsay EH" pageId="4" pageNumber="668" pagination="131 - 147" refId="ref11418" refString="Lindsay EH. 1994. The evolution of murine rodents in Asia. National Science Museum Monographs 8: 131 - 147." type="journal article" year="1994">Lindsay (1994</bibRefCitation>
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: table 1) listed
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<taxonomicName id="4C35A1F8FF86FFAFD123C74A4AECFBB4" box="[1090,1244,1025,1046]" class="Mammalia" family="Muridae" genus="Antemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
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<emphasis id="B9410669FF86FFAFD123C74A4AECFBB4" box="[1090,1244,1025,1046]" italics="true" pageId="4" pageNumber="668">P. primitivus</emphasis>
|
|
</taxonomicName>
|
|
in the Kamlial Formation of
|
|
<collectingCountry id="F3229AEBFF86FFAFD6B8C7544A72FB97" box="[985,1090,1055,1077]" name="Pakistan" pageId="4" pageNumber="668">Pakistan</collectingCountry>
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|
, but also in the Manchar Formation and not in the Chinji Formation.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C32F89F0FF86FFAFD632C7174ADBF915" pageId="4" pageNumber="668" type="materials_examined">
|
|
<paragraph id="8B8ADA7BFF86FFAFD632C7174ADBF915" blockId="4.[826,1442,198,1719]" pageId="4" pageNumber="668">
|
|
The
|
|
<typeStatus id="548E64D9FF86FFAFD6EFC7174DDFFBD0" box="[910,1007,1116,1138]" pageId="4" pageNumber="668" type="holotype">holotype</typeStatus>
|
|
of this species (H-GSP 5521) is a right M1 from Chinji locality H-GSP 107, which is possibly Middle Miocene in age (
|
|
<bibRefCitation id="EFA4A78AFF86FFAFD192C7D24D44FB6C" author="Wessels W & de Bruijn H & Hussain ST & Leinders J" pageId="4" pageNumber="668" pagination="337 - 364" refId="ref12338" refString="Wessels W, de Bruijn H, Hussain ST, Leinders J. 1982. Fossil rodents from the Chinji Formation, Banda Daud Shah, Kohat, Pakistan. Koninklijke Nederlandse Akademie van Wetenschappen Proceedings, Series B 85: 337 - 364." type="journal article" year="1982">
|
|
Wessels
|
|
<emphasis id="B9410669FF86FFAFD002C7D14BAAFB0C" box="[1379,1434,1177,1199]" italics="true" pageId="4" pageNumber="668">et al.</emphasis>
|
|
, 1982
|
|
</bibRefCitation>
|
|
). The morphology of the first lower molar of
|
|
<taxonomicName id="4C35A1F8FF86FFAFD65BC79C4A0DFB4E" box="[826,1085,1239,1260]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
|
|
<emphasis id="B9410669FF86FFAFD65BC79C4A0DFB4E" box="[826,1085,1239,1260]" italics="true" pageId="4" pageNumber="668">Potwarmus primitivus</emphasis>
|
|
</taxonomicName>
|
|
is very distinct from
|
|
<taxonomicName id="4C35A1F8FF86FFAFD020C79C4B91FB49" authority="Antoñanzas, 2009" authorityName="Antoñanzas" authorityYear="2009" box="[1345,1441,1238,1260]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="flynni" status="sp. nov.">
|
|
<emphasis id="B9410669FF86FFAFD020C79C4B91FB49" box="[1345,1441,1238,1260]" italics="true" pageId="4" pageNumber="668">P. flynni</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A272BB12FF86FFAFD65BC7BE4DA1FAA8" box="[826,913,1269,1290]" pageId="4" pageNumber="668" rank="species">sp. nov.</taxonomicNameLabel>
|
|
The m1s of
|
|
<taxonomicName id="4C35A1F8FF86FFAFD144C7BE4A8AFAA8" box="[1061,1210,1269,1290]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
|
|
<emphasis id="B9410669FF86FFAFD144C7BE4A8AFAA8" box="[1061,1210,1269,1290]" italics="true" pageId="4" pageNumber="668">P. primitivus</emphasis>
|
|
</taxonomicName>
|
|
have an unreduced anterior portion: they have a distinct single anteroconid and a metaconid, which are not distinct in
|
|
<taxonomicName id="4C35A1F8FF86FFAFD0ECC6784D4CFAC4" authority="Antoñanzas, 2009" authorityName="Antoñanzas" authorityYear="2009" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="flynni" status="sp. nov.">
|
|
<emphasis id="B9410669FF86FFAFD0ECC6784D4CFAC4" italics="true" pageId="4" pageNumber="668">P. flynni</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A272BB12FF86FFAFD6EBC61A4DD7FAC4" box="[906,999,1361,1382]" pageId="4" pageNumber="668" rank="species">sp. nov.</taxonomicNameLabel>
|
|
<taxonomicName id="4C35A1F8FF86FFAFD695C61A4ACBFAC4" box="[1012,1275,1361,1382]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
|
|
<emphasis id="B9410669FF86FFAFD695C61A4ACBFAC4" box="[1012,1275,1361,1382]" italics="true" pageId="4" pageNumber="668">Potwarmus primitivus</emphasis>
|
|
</taxonomicName>
|
|
has a larger enterostyle and a better developed longitudinal crest on the M1s than AJ7. On the M2s of
|
|
<taxonomicName id="4C35A1F8FF86FFAFD065C6C54BAAFA01" box="[1284,1434,1422,1443]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
|
|
<emphasis id="B9410669FF86FFAFD065C6C54BAAFA01" box="[1284,1434,1422,1443]" italics="true" pageId="4" pageNumber="668">P. primitivus</emphasis>
|
|
</taxonomicName>
|
|
, the longitudinal crest is also better developed than in the Arabian specimens and, the strong, long lingual branch of the anterior cingulum descends lingually continuing posteriorly as a prominent lingual cingulum. In contrast, these cingula are less prominent, much shorter and not in continuity both in AJ11 and AJ12. In addition, the hypocone joins with the metacone more anteriorly on the M2 of
|
|
<taxonomicName id="4C35A1F8FF86FFAFD0ECC5C84D81F915" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="primitivus">
|
|
<emphasis id="B9410669FF86FFAFD0ECC5C84D81F915" italics="true" pageId="4" pageNumber="668">P. primitivus</emphasis>
|
|
</taxonomicName>
|
|
than in
|
|
<taxonomicName id="4C35A1F8FF86FFAFD143C5E94AB4F914" authority="Antoñanzas, 2009" authorityName="Antoñanzas" authorityYear="2009" box="[1058,1156,1697,1719]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="668" phylum="Chordata" rank="species" species="flynni" status="sp. nov.">
|
|
<emphasis id="B9410669FF86FFAFD143C5E94AB4F914" box="[1058,1156,1697,1719]" italics="true" pageId="4" pageNumber="668">P. flynni</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A272BB12FF86FFAFD1F1C5E94ADBF915" box="[1168,1259,1698,1719]" pageId="4" pageNumber="668" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |