treatments-xml/data/03/9C/6B/039C6B6DFF84FFADD5B9C7844B3EFDC8.xml
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<mods:title id="4C2A18727A3BF936ABD8D300CB0AA451">First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea)</mods:title>
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<heading id="D0C26D17FF84FFADD5B9C7844CE7FB41" allCaps="true" box="[216,727,1230,1252]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="670" reason="9">
<taxonomicName id="4C35A1F8FF84FFADD5B9C7844CE7FB41" authority="CHINJIENSIS LINDSAY, 1988" authorityName="CHINJIENSIS LINDSAY" authorityYear="1988" box="[216,727,1230,1252]" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF84FFADD5B9C7844C11FB40" box="[216,545,1231,1252]" italics="true" pageId="6" pageNumber="670">PARADAKKAMYS CHINJIENSIS</emphasis>
<bibRefCitation id="EFA4A78AFF84FFADD74DC7844CE7FB41" author="Lindsay EH" box="[556,727,1230,1252]" pageId="6" pageNumber="670" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">LINDSAY, 1988</bibRefCitation>
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<subSubSection id="C32F89F0FF84FFADD5C5C7BE4B3EFDC8" pageId="6" pageNumber="670" type="discussion">
<paragraph id="8B8ADA7BFF84FFADD5C5C7BE4FCFF915" blockId="6.[164,779,1269,1903]" pageId="6" pageNumber="670">
This species was erected by
<bibRefCitation id="EFA4A78AFF84FFADD4B8C7BE4CB1FAA9" author="Lindsay EH" box="[473,641,1269,1291]" pageId="6" pageNumber="670" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay (1988)</bibRefCitation>
on the basis of 23 isolated cheek teeth from the Middle Miocene localities YGSP 76, YGSP 491, YGSP 504, YGSP 634, YGSP 636, and YGSP
<quantity id="4CCD779EFF84FFADD4FAC61A4FD0FAC5" box="[411,480,1361,1383]" metricMagnitude="1" metricUnit="m" metricValue="1.84404" pageId="6" pageNumber="670" unit="in" value="726.0">726 in</quantity>
the Chinji Formation, and from the early Late Miocene YGSP 259 locality in the Nagri Formation, Potwar Plateau,
<collectingCountry id="F3229AEBFF84FFADD733C6C54C8BFA06" box="[594,699,1422,1444]" name="Pakistan" pageId="6" pageNumber="670">Pakistan</collectingCountry>
. Additional material has been found in the Nagri Formation at locality Y797 (
<bibRefCitation id="EFA4A78AFF84FFADD403C6804C57FA43" author="Flynn LJ &amp; Morgan ME" box="[354,615,1483,1505]" pageId="6" pageNumber="670" pagination="1 - 10" refId="ref10989" refString="Flynn LJ, Morgan ME. 2005. An unusual diatomyid rodent from an infrequently sampled late Miocene interval in the Siwaliks of Pakistan. Palaeontologia Electronica 8 17 A: 1 - 10." type="journal article" year="2005">Flynn &amp; Morgan, 2005</bibRefCitation>
). The
<typeStatus id="548E64D9FF84FFADD7CBC6804D3BFA43" box="[682,779,1483,1505]" pageId="6" pageNumber="670" type="holotype">holotype</typeStatus>
(YGSP 24656) of this species is an isolated right M1 from locality YGSP 76 (
<bibRefCitation id="EFA4A78AFF84FFADD4A2C5434C59F9BF" author="Lindsay EH" box="[451,617,1544,1566]" pageId="6" pageNumber="670" pagination="95 - 154" refId="ref11386" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay, 1988</bibRefCitation>
: pl. 8, fig. h).
<bibRefCitation id="EFA4A78AFF84FFADD5C5C56C4F7EF99F" author="Wessels W" box="[164,334,1575,1597]" pageId="6" pageNumber="670" pagination="253 - 312" refId="ref12311" refString="Wessels W. 1996. Myocricetodontinae from the Miocene of Pakistan. Koninklijke Nederlandse Akademie van Wetenschappen Proceedings, Series B 99: 253 - 312." type="journal article" year="1996">Wessels (1996)</bibRefCitation>
suggested that this species should be re-allocated to
<taxonomicName id="4C35A1F8FF84FFADD437C50E4FC9F9F8" box="[342,505,1605,1626]" class="Mammalia" family="Muridae" genus="Myocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF84FFADD437C50E4FC9F9F8" box="[342,505,1605,1626]" italics="true" pageId="6" pageNumber="670">Myocricetodon</emphasis>
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because, in particular, of its resemblance to
<taxonomicName id="4C35A1F8FF84FFADD4D2C52F4D34F9DB" box="[435,772,1636,1657]" class="Mammalia" family="Muridae" genus="Myocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="species" species="eskihisarensis">
<emphasis id="B9410669FF84FFADD4D2C52F4D34F9DB" box="[435,772,1636,1657]" italics="true" pageId="6" pageNumber="670">Myocricetodon eskihisarensis</emphasis>
</taxonomicName>
. However, the phylogenetic study below corroborates the validity of
<taxonomicName id="4C35A1F8FF84FFADD42CC5EA4FC8F914" authorityName="CHINJIENSIS LINDSAY" authorityYear="1988" box="[333,504,1697,1718]" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF84FFADD42CC5EA4FC8F914" box="[333,504,1697,1718]" italics="true" pageId="6" pageNumber="670">Paradakkamys</emphasis>
</taxonomicName>
.
</paragraph>
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The M1s of
<taxonomicName id="4C35A1F8FF84FFADD409C58B4CAFF977" box="[360,671,1728,1749]" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="species" species="chinjiensis">
<emphasis id="B9410669FF84FFADD409C58B4CAFF977" box="[360,671,1728,1749]" italics="true" pageId="6" pageNumber="670">Paradakkamys chinjiensis</emphasis>
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have a prominent enterostyle connected to the posterior side of the protocone, a paracone with a small posteriorly directed spur, a long anterior arm of the protocone joined with the anterocone near its midpoint, and a short lingual cingulum. In AJ7 the enterostyle is small and isolated, the spur of the paracone is lacking, the anterior arm of the protocone is only a relict, and the lingual cingulum is long. On the M2 of
<taxonomicName id="4C35A1F8FF84FFADD65BC2694DE1FE95" box="[826,977,290,311]" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="species" species="chinjiensis">
<emphasis id="B9410669FF84FFADD65BC2694DE1FE95" box="[826,977,290,311]" italics="true" pageId="6" pageNumber="670">P. chinjiensis</emphasis>
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the enterostyle is small, but connected to the posterior side of the protocone by a short loph, and the lingual anterior cingulum is normally absent. In AJ11 and AJ12, the enterostyle is isolated and the lingual anterior cingulum is strong. The m1s of
<taxonomicName id="4C35A1F8FF84FFADD0ECC2D64D83FE72" class="Mammalia" family="Muridae" genus="Paradakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="670" phylum="Chordata" rank="species" species="chinjiensis">
<emphasis id="B9410669FF84FFADD0ECC2D64D83FE72" italics="true" pageId="6" pageNumber="670">P. chinjiensis</emphasis>
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are not reduced anteriorly: they show a well-developed anteroconid, a large ectostylid, short anterior arms of the labial cusps, and a labial cingulum. In contrast, the anteroconid, the anterior arms of the labial cusps, and the labial cingulum are lacking in AJ13, and it has a minute ectostylid.
</paragraph>
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