treatments-xml/data/03/9C/1A/039C1A521C5EA558FE827A8DCEA34673.xml
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<document id="38931A265AE10A72EDF2EFE7A42F185E" ID-CLB-Dataset="4220" ID-DOI="10.12782/sd.22_29" ID-GBIF-Dataset="111f6060-421e-4f53-bfb2-63a7d605f1a4" ID-ISSN="2189-7301" ID-Zenodo-Dep="4583717" ID-ZooBank="EFF0CA4B-AD0A-44B4-99BA-79446785ED0A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614892972232" checkinUser="felipe" docAuthor="Okumura, Ken-ichi" docDate="2017" docId="039C1A521C5EA558FE827A8DCEA34673" docLanguage="en" docName="SpeciesDiversity.22.1.29-36.pdf" docOrigin="Species Diversity 22 (1)" docSource="http://dx.doi.org/10.12782/sd.22_29" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Dichodactylus shinshuensis Okumura 2017, sp. nov." docType="treatment" docUuid="D476485E-EEEA-4983-8941-A6AA3401638C" docUuidSource="ZooBank" docVersion="10" lastPageNumber="31" masterDocId="FFA5622A1C5CA55AFF947C13CD204006" masterDocTitle="Dichodactylus gen. nov. (Araneae: Agelenidae: Coelotinae) from Japan" masterLastPageNumber="36" masterPageNumber="29" pageNumber="31" updateTime="1698932642149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E62540B39AFEDC98AAE5C3A3663473AF">Dichodactylus gen. nov. (Araneae: Agelenidae: Coelotinae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="2ED40B3992F8ECFD4724E8B6B0B1A32D" type="personal">
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<mods:namePart id="D835AF9DADBADD497AB3406E55839EC6">Okumura, Ken-ichi</mods:namePart>
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<mods:title id="ACFC141E2FBECD68BC23822F2AC66833">Species Diversity</mods:title>
</mods:titleInfo>
<mods:part id="0257CEFF360580FC7CBD2BF74765CF3F">
<mods:date id="016F16B0748460FE65EE199B61B1D0FE">2017</mods:date>
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<mods:number id="FA40E684D8B527CB984F128F29D620A3">2017-05-25</mods:number>
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<treatment id="039C1A521C5EA558FE827A8DCEA34673" ID-DOI="http://doi.org/10.5281/zenodo.4584018" ID-GBIF-Taxon="188493171" ID-Zenodo-Dep="4584018" LSID="urn:lsid:zoobank.org:act:D476485E-EEEA-4983-8941-A6AA3401638C" httpUri="http://treatment.plazi.org/id/039C1A521C5EA558FE827A8DCEA34673" lastPageNumber="31" pageId="2" pageNumber="31">
<subSubSection id="C32FF8CF1C5EA558FE827A8DCFA546B0" box="[278,645,1694,1718]" pageId="2" pageNumber="31" type="nomenclature">
<paragraph id="8B8AAB441C5EA558FE827A8DCFA546B0" blockId="2.[278,645,1694,1718]" box="[278,645,1694,1718]" pageId="2" pageNumber="31">
<heading id="D0C21C281C5EA558FE827A8DCFA546B0" box="[278,645,1694,1718]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="31" reason="2">
<taxonomicName id="4C35D0C71C5EA558FE827A8DCF1346B0" ID-CoL="35M3M" authority="Okumura, 2017" authorityName="Okumura" authorityYear="2017" box="[278,563,1694,1718]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="shinshuensis" status="sp. nov.">
<emphasis id="B94177561C5EA558FE827A8DCF1346B0" bold="true" box="[278,563,1694,1718]" italics="true" pageId="2" pageNumber="31">Dichodactylus shinshuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272CA2D1C5EA558FDAD7A8CCFA546B0" box="[569,645,1695,1718]" pageId="2" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32FF8CF1C5EA558FEF47AACCF1B46D1" box="[352,571,1727,1751]" pageId="2" pageNumber="31" type="description">
<paragraph id="8B8AAB441C5EA558FEF47AACCF1B46D1" blockId="2.[352,571,1727,1751]" box="[352,571,1727,1751]" pageId="2" pageNumber="31">
<heading id="D0C21C281C5EA558FEF47AACCF1B46D1" box="[352,571,1727,1751]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="31" reason="2">
(
<figureCitation id="130EB7C11C5EA558FEF37AACCC8446D1" box="[359,420,1727,1751]" captionStart="Fig" captionStartId="3.[156,187,1637,1659]" captionTargetBox="[278,1319,206,1597]" captionTargetId="figure-161@3.[265,1323,205,1598]" captionTargetPageId="3" captionText="Fig. 2. Dichodactylus shinshuensis sp. nov., male (A, B: holotype, NSMT-Ar 13339), female (C, D: paratype, NSMT-Ar 13341). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CD, copulatory duct; CO, conductor; CY, cymbial furrow; EM, embolus; LAM, lamellas; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583722" httpUri="https://zenodo.org/record/4583722/files/figure.png" pageId="2" pageNumber="31">Figs 2</figureCitation>
AD, 5A, D, 6)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32FF8CF1C5EA558FF087B13CF04471E" box="[156,548,1792,1816]" pageId="2" pageNumber="31" type="reference_group">
<paragraph id="8B8AAB441C5EA558FF087B13CF04471E" blockId="2.[156,548,1792,1816]" box="[156,548,1792,1816]" pageId="2" pageNumber="31">
<treatmentCitationGroup id="AB258C6A1C5EA558FF087B13CF04471E" box="[156,548,1792,1816]" pageId="2" pageNumber="31">
<taxonomicName id="4C35D0C71C5EA558FF087B13CDCD471E" authorityName="Blackwall" authorityYear="1841" box="[156,237,1792,1816]" class="Arachnida" family="Amaurobiidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B94177561C5EA558FF087B13CDCD471E" box="[156,237,1792,1816]" italics="true" pageId="2" pageNumber="31">Coelotes</emphasis>
</taxonomicName>
sp.:
<treatmentCitation id="0A948D551C5EA558FE8E7B13CCF6471E" author="Chikuni, Y." box="[282,470,1792,1816]" page="103" pageId="2" pageNumber="31" year="1989">
<bibRefCitation id="EFA4D6B51C5EA558FE8E7B13CCF6471E" author="Chikuni, Y." box="[282,470,1792,1816]" pageId="2" pageNumber="31" refId="ref4580" refString="Chikuni, Y. 1989. Pictorial Encyclopedia of Spiders in Japan. Kaiseisha, Tokyo, 310 pp. [In Japanese]" type="book" year="1989">Chikuni 1989: 103</bibRefCitation>
</treatmentCitation>
, fig. 25.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32FF8CF1C5EA558FF217B52CDC8481B" pageId="2" pageNumber="31" type="materials_examined">
<paragraph id="8B8AAB441C5EA558FF217B52CDC8481B" blockId="2.[156,768,1857,2077]" pageId="2" pageNumber="31">
<emphasis id="B94177561C5EA558FF217B52CC73475F" bold="true" box="[181,339,1857,1881]" pageId="2" pageNumber="31">Type material.</emphasis>
<materialsCitation id="3B5DA1191C5EA558FEF87B52CCBD47BD" ID-GBIF-Occurrence="3357537335" collectingDate="2009-10" collectingDateMax="2010-10-21" collectingDateMin="2009-10" county="Shimo-ina County" elevation="840" latitude="35.950558" location="Tochibaradani" longLatPrecision="19" longitude="137.56694" municipality="Takamori Town" pageId="2" pageNumber="31" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nagano" typeStatus="holotype">
<collectorName id="26C0CE921C5EA558FEF87B52CCAB475F" box="[364,395,1857,1881]" pageId="2" pageNumber="31">All</collectorName>
collected by
<collectorName id="26C0CE921C5EA558FD8A7B52CFAC475F" box="[542,652,1857,1881]" pageId="2" pageNumber="31">Yoh Ihara.</collectorName>
<emphasis id="B94177561C5EA558FD037B52CFD9475F" bold="true" box="[663,761,1857,1881]" pageId="2" pageNumber="31">
<typeStatus id="548E15E61C5EA558FD037B52CFD9475F" box="[663,761,1857,1881]" pageId="2" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<collectorName id="26C0CE921C5EA558FF757B70CC65477B" box="[225,325,1891,1917]" pageId="2" pageNumber="31">Ōjimasan</collectorName>
,
<quantity id="4CCD06A11C5EA558FECC7B71CCB5477F" box="[344,405,1890,1914]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="2" pageNumber="31" unit="m" value="840.0">
<elevation id="00184C771C5EA558FECC7B71CCB5477F" box="[344,405,1890,1914]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="2" pageNumber="31" unit="m" value="840.0">840 m</elevation>
</quantity>
alt.,
<collectingMunicipality id="6BEE313E1C5EA558FE437B71CFA2477F" box="[471,642,1890,1914]" pageId="2" pageNumber="31">Takamori Town</collectingMunicipality>
,
<collectingCounty id="62EBD3C81C5EA558FD007B71CDC8479C" pageId="2" pageNumber="31">Shimo-ina County</collectingCounty>
,
<collectingRegion id="49F165A61C5EA558FF6A7B90CCAE479C" box="[254,398,1922,1947]" country="Japan" name="Nagano" pageId="2" pageNumber="31">Nagano Pref.</collectingRegion>
(
<geoCoordinate id="EE01CD831C5EA558FE327B91CF32479C" box="[422,530,1921,1946]" degrees="35" direction="north" minutes="57" orientation="latitude" pageId="2" pageNumber="31" precision="15" seconds="2" value="35.950558">35°572″N</geoCoordinate>
,
<geoCoordinate id="EE01CD831C5EA558FDBC7B91CFBB479C" box="[552,667,1921,1946]" pageId="2" pageNumber="31">137°841″E</geoCoordinate>
),
<date id="FF8B8D841C5EA558FD2D7B91CDED47BD" pageId="2" pageNumber="31" value="2010-10-21">
<collectingDate id="EFCF746C1C5EA558FD2D7B91CDED47BD" pageId="2" pageNumber="31" value="2010-10-21">21. X. 2010</collectingDate>
</date>
(NSMT-Ar 13339).
</materialsCitation>
<materialsCitation id="3B5DA1191C5EA558FE327BB0CDC8481B" ID-GBIF-Occurrence="3357537371" collectingDate="2009-10" collectingDateMax="2010-10-21" collectingDateMin="2009-10" county="Shimo-ina County" elevation="840" latitude="35.950558" location="Tochibaradani" longLatPrecision="19" longitude="137.56694" municipality="Takamori Town" pageId="2" pageNumber="31" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nagano" typeStatus="paratype">
<emphasis id="B94177561C5EA558FE327BB0CF2E47BD" bold="true" box="[422,526,1955,1979]" pageId="2" pageNumber="31">
<typeStatus id="548E15E61C5EA558FE327BB0CF2E47BD" box="[422,526,1955,1979]" pageId="2" pageNumber="31" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D3360CD1C5EA558FD8A7BB0CF4547BD" box="[542,613,1955,1979]" pageId="2" pageNumber="31" type="male">1 male</specimenCount>
and
<specimenCount id="9D3360CD1C5EA558FD347BB0CFDB47BD" box="[672,763,1955,1979]" pageId="2" pageNumber="31" type="female">1 female</specimenCount>
,
<location id="8EEAFD9F1C5EA558FF087BD0CC1047DD" LSID="urn:lsid:plazi:treatment:039C1A521C5EA558FE827A8DCEA34673:8EEAFD9F1C5EA558FF087BD0CC1047DD" box="[156,304,1987,2011]" county="Shimo-ina County" latitude="35.950558" longLatPrecision="19" longitude="137.56694" municipality="Takamori Town" name="Tochibaradani" pageId="2" pageNumber="31" stateProvince="Nagano">Tochibaradani</location>
,
<location id="8EEAFD9F1C5EA558FEAA7BD0CCA447DD" LSID="urn:lsid:plazi:treatment:039C1A521C5EA558FE827A8DCEA34673:8EEAFD9F1C5EA558FEAA7BD0CCA447DD" box="[318,388,1987,2011]" county="Shimo-ina County" latitude="35.950558" longLatPrecision="19" longitude="137.56694" municipality="Takamori Town" name="Ochiai" pageId="2" pageNumber="31" stateProvince="Nagano">Ochiai</location>
,
<collectingRegion id="49F165A61C5EA558FE057BD0CCF047DD" box="[401,464,1987,2011]" country="Japan" name="Osaka" pageId="2" pageNumber="31">Osaka</collectingRegion>
<location id="8EEAFD9F1C5EA558FE4E7BD7CF3547DD" LSID="urn:lsid:plazi:treatment:039C1A521C5EA558FE827A8DCEA34673:8EEAFD9F1C5EA558FE4E7BD7CF3547DD" box="[474,533,1988,2011]" county="Shimo-ina County" latitude="35.950558" longLatPrecision="19" longitude="137.56694" municipality="Takamori Town" name="Town" pageId="2" pageNumber="31" stateProvince="Nagano">Town</location>
,
<location id="8EEAFD9F1C5EA558FDB77BD7CFAB47DD" LSID="urn:lsid:plazi:treatment:039C1A521C5EA558FE827A8DCEA34673:8EEAFD9F1C5EA558FDB77BD7CFAB47DD" box="[547,651,1987,2012]" county="Shimo-ina County" latitude="35.950558" longLatPrecision="19" longitude="137.56694" municipality="Takamori Town" name="Gero City" pageId="2" pageNumber="31" stateProvince="Nagano">Gero City</location>
,
<collectingRegion id="49F165A61C5EA558FD0C7BD0CE2047DD" box="[664,768,1987,2011]" country="Japan" name="Gifu" pageId="2" pageNumber="31">Gifu Pref.</collectingRegion>
(
<geoCoordinate id="EE01CD831C5EA558FF377BF7CC2F47FD" box="[163,271,2019,2044]" pageId="2" pageNumber="31">35°957″N</geoCoordinate>
,
<geoCoordinate id="EE01CD831C5EA558FEB67BF7CCB547FA" box="[290,405,2019,2044]" degrees="137" direction="east" minutes="34" orientation="longitude" pageId="2" pageNumber="31" precision="15" seconds="1" value="137.56694">137°341″E</geoCoordinate>
), 14,
<date id="FF8B8D841C5EA558FE4F7BF7CF1147FA" box="[475,561,2020,2044]" pageId="2" pageNumber="31" value="2009-10">
<collectingDate id="EFCF746C1C5EA558FE4F7BF7CF1147FA" box="[475,561,2020,2044]" pageId="2" pageNumber="31" value="2009-10">X. 2009</collectingDate>
</date>
(NSMT-Ar 13340, 13341).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32FF8CF1C5EA558FCDA7CD5C90141A7" pageId="2" pageNumber="31" type="diagnosis">
<paragraph id="8B8AAB441C5EA558FCDA7CD5C90141A7" blockId="2.[820,1432,198,1653]" pageId="2" pageNumber="31">
<emphasis id="B94177561C5EA558FCDA7CD5CE9E40D8" bold="true" box="[846,958,198,222]" pageId="2" pageNumber="31">Diagnosis.</emphasis>
<taxonomicName id="4C35D0C71C5EA558FC437CD5C9D240D8" authority="Okumura, 2017" authorityName="Okumura" authorityYear="2017" box="[983,1266,198,222]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="shinshuensis" status="sp. nov.">
<emphasis id="B94177561C5EA558FC437CD5C9D240D8" box="[983,1266,198,222]" italics="true" pageId="2" pageNumber="31">Dichodactylus shinshuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272CA2D1C5EA558FA927CD4C84140D8" box="[1286,1377,199,222]" pageId="2" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from all the other
<taxonomicName id="4C35D0C71C5EA558FB457CF4C87E40F9" authorityName="Okumura" authorityYear="2017" box="[1233,1374,231,255]" class="Arachnida" family="Agelenidae" genus="Dichodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B94177561C5EA558FB457CF4C87E40F9" box="[1233,1374,231,255]" italics="true" pageId="2" pageNumber="31">Dichodactylus</emphasis>
</taxonomicName>
species by having the tip of the male palpal conductor twisted proximally at a right angle (
<figureCitation id="130EB7C11C5EA558FBC17D3BC9854146" box="[1109,1189,296,320]" captionStart="Fig" captionStartId="3.[156,187,1637,1659]" captionTargetBox="[278,1319,206,1597]" captionTargetId="figure-161@3.[265,1323,205,1598]" captionTargetPageId="3" captionText="Fig. 2. Dichodactylus shinshuensis sp. nov., male (A, B: holotype, NSMT-Ar 13339), female (C, D: paratype, NSMT-Ar 13341). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CD, copulatory duct; CO, conductor; CY, cymbial furrow; EM, embolus; LAM, lamellas; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583722" httpUri="https://zenodo.org/record/4583722/files/figure.png" pageId="2" pageNumber="31">Fig. 2A</figureCitation>
), the presence of a pair of lamellae laterally on the epigynum (
<figureCitation id="130EB7C11C5EA558FB4D7D5BC80F4166" box="[1241,1327,328,352]" captionStart="Fig" captionStartId="3.[156,187,1637,1659]" captionTargetBox="[278,1319,206,1597]" captionTargetId="figure-161@3.[265,1323,205,1598]" captionTargetPageId="3" captionText="Fig. 2. Dichodactylus shinshuensis sp. nov., male (A, B: holotype, NSMT-Ar 13339), female (C, D: paratype, NSMT-Ar 13341). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CD, copulatory duct; CO, conductor; CY, cymbial furrow; EM, embolus; LAM, lamellas; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583722" httpUri="https://zenodo.org/record/4583722/files/figure.png" pageId="2" pageNumber="31">Figs 2C</figureCitation>
,
<figureCitation id="130EB7C11C5EA558FAAA7D5BC87E4166" box="[1342,1374,328,352]" captionStart="Fig" captionStartId="5.[156,187,1156,1178]" captionTargetBox="[258,1330,205,1117]" captionTargetId="figure-449@5.[258,1330,205,1117]" captionTargetPageId="5" captionText="Fig. 5. Comparison of genitalic characters of three species of Dichodactylus gen. nov. from Japan: A, D, D. shinshuensis sp. nov., male (A: NSMT-Ar 13339), and female (D: NSMT-Ar 13341); B, E, D. tarumii (Arita, 1976), male (B: NSMT-Ar 14732), and female (E: NSMT-Ar 14733); C, F, D. satoi (Nishikawa, 2003), male (C: NSMT-Ar 14735), and female (F: NSMT-Ar 14734). AC, retrolateral views of male palp; DF, ventral views of epigynum. Abbreviations: CO, conductor; ET, epigynal teeth; LAM, lamellas; PA, patellar apophyses; SW, swollen portion of tegulum. Black circle on tibia shows the vestigial LTA." figureDoi="http://doi.org/10.5281/zenodo.4583728" httpUri="https://zenodo.org/record/4583728/files/figure.png" pageId="2" pageNumber="31">5D</figureCitation>
) and the prominent copulatory ducts visible in ventral view of cleared vulva (
<figureCitation id="130EB7C11C5EA558FC527D9AC93441A7" box="[966,1044,393,417]" captionStart="Fig" captionStartId="3.[156,187,1637,1659]" captionTargetBox="[278,1319,206,1597]" captionTargetId="figure-161@3.[265,1323,205,1598]" captionTargetPageId="3" captionText="Fig. 2. Dichodactylus shinshuensis sp. nov., male (A, B: holotype, NSMT-Ar 13339), female (C, D: paratype, NSMT-Ar 13341). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CD, copulatory duct; CO, conductor; CY, cymbial furrow; EM, embolus; LAM, lamellas; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583722" httpUri="https://zenodo.org/record/4583722/files/figure.png" pageId="2" pageNumber="31">Fig. 2D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32FF8CF1C5EA558FCDA7DB9C83045D5" pageId="2" pageNumber="31" type="description">
<paragraph id="8B8AAB441C5EA558FCDA7DB9C9AB42C0" blockId="2.[820,1432,198,1653]" pageId="2" pageNumber="31">
<emphasis id="B94177561C5EA558FCDA7DB9C9E141C7" bold="true" box="[846,1217,425,450]" pageId="2" pageNumber="31">
Description of male (
<typeStatus id="548E15E61C5EA558FBC67DBAC99541C7" box="[1106,1205,425,449]" pageId="2" pageNumber="31" type="holotype">holotype</typeStatus>
).
</emphasis>
Total length 5.6, carapace 2.9 long, 2.0 wide; abdomen 2.7 long, 1.6 wide; sternum 1.6 long, 1.3 wide. Eye sizes; AME 0.07, ALE 0.15, PME 0.12, PLE 0.14. Distances between eyes; AME-AME 0.07, AME-ALE 0.04, PME-PME 0.09, PME-PLE 0.12, AME-PME 0.10, ALE-PLE 0.04. MOA; anterior width 0.21, posterior width 0.33, length 0.29. Leg measurements: I: 7.9 (2.1, 2.7, 1.9, 1.2); II: 7.1 (2.1, 2.3, 1.7, 1.0); III: 6.6 (1.7, 2.1, 1.9, 0.9); IV: 9.4 (2.5, 3.0, 2.7, 1.2).
</paragraph>
<paragraph id="8B8AAB441C5EA558FCDA7EDDCE8D43CC" blockId="2.[820,1432,198,1653]" pageId="2" pageNumber="31">
Palp (
<figureCitation id="130EB7C11C5EA558FC1C7EDDCED442E0" box="[904,1012,718,743]" captionStart="Fig" captionStartId="3.[156,187,1637,1659]" captionTargetBox="[278,1319,206,1597]" captionTargetId="figure-161@3.[265,1323,205,1598]" captionTargetPageId="3" captionText="Fig. 2. Dichodactylus shinshuensis sp. nov., male (A, B: holotype, NSMT-Ar 13339), female (C, D: paratype, NSMT-Ar 13341). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CD, copulatory duct; CO, conductor; CY, cymbial furrow; EM, embolus; LAM, lamellas; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583722" httpUri="https://zenodo.org/record/4583722/files/figure.png" pageId="2" pageNumber="31">Figs 2A, B</figureCitation>
,
<figureCitation id="130EB7C11C5EA558FC6B7EDDC90042E0" box="[1023,1056,718,742]" captionStart="Fig" captionStartId="5.[156,187,1156,1178]" captionTargetBox="[258,1330,205,1117]" captionTargetId="figure-449@5.[258,1330,205,1117]" captionTargetPageId="5" captionText="Fig. 5. Comparison of genitalic characters of three species of Dichodactylus gen. nov. from Japan: A, D, D. shinshuensis sp. nov., male (A: NSMT-Ar 13339), and female (D: NSMT-Ar 13341); B, E, D. tarumii (Arita, 1976), male (B: NSMT-Ar 14732), and female (E: NSMT-Ar 14733); C, F, D. satoi (Nishikawa, 2003), male (C: NSMT-Ar 14735), and female (F: NSMT-Ar 14734). AC, retrolateral views of male palp; DF, ventral views of epigynum. Abbreviations: CO, conductor; ET, epigynal teeth; LAM, lamellas; PA, patellar apophyses; SW, swollen portion of tegulum. Black circle on tibia shows the vestigial LTA." figureDoi="http://doi.org/10.5281/zenodo.4583728" httpUri="https://zenodo.org/record/4583728/files/figure.png" pageId="2" pageNumber="31">5A</figureCitation>
): two patellar apophyses present, situated at a right angle from each other in retrolateral view, LTA reduced, RTA large and broad; cymbial furrow short and indistinct about one-sixth of cymbial length; embolus flagelliform and short, conductor spatulate and tip twisted proximally at a right angle; median apophysis small, crescent-shaped; tegulum with swollen portion visible in retrolateral view.
</paragraph>
<paragraph id="8B8AAB441C5EA558FCDA7FC1C9AB44E9" blockId="2.[820,1432,198,1653]" pageId="2" pageNumber="31">
<emphasis id="B94177561C5EA558FCDA7FC1C9E843EC" bold="true" box="[846,1224,978,1002]" pageId="2" pageNumber="31">
Description of female (
<typeStatus id="548E15E61C5EA558FBCE7FC1C99D43EC" box="[1114,1213,978,1002]" pageId="2" pageNumber="31" type="paratype">paratype</typeStatus>
).
</emphasis>
Total length 5.0, carapace 2.7 long, 1.5 wide; abdomen 2.3 long, 1.4 wide; sternum 3.2 long, 2.4 wide. Eye sizes; AME 0.06, ALE 0.14, PME 0.12, PLE 0.10. Distances between eyes; AME-AME 0.04, AME-ALE 0.03, PME-PME 0.07, PME-PLE 0.10, AME-PME 0.09, ALE-PLE 0.03. MOA; anterior width 0.16, posterior width 0.31, length 0.27. Leg measurements: I: 6.3 (1.9, 2.1, 1.4, 0.9); II: 5.6 (1.7, 1.8, 1.3, 0-8); III: 5.3 (1.5, 1.8, 1.3, 0.7); IV: 7.2 (2.0, 2.4, 1.9, 0.9).
</paragraph>
<paragraph id="8B8AAB441C5EA558FCDA78E4C83045D5" blockId="2.[820,1432,198,1653]" pageId="2" pageNumber="31">
Epigynum and internal genitalia (
<figureCitation id="130EB7C11C5EA558FB3378E4C8374509" box="[1191,1303,1271,1296]" captionStart="Fig" captionStartId="3.[156,187,1637,1659]" captionTargetBox="[278,1319,206,1597]" captionTargetId="figure-161@3.[265,1323,205,1598]" captionTargetPageId="3" captionText="Fig. 2. Dichodactylus shinshuensis sp. nov., male (A, B: holotype, NSMT-Ar 13339), female (C, D: paratype, NSMT-Ar 13341). A, ventral view of male palp; B, retrolateral view of male palp; C, ventral view of epigynum; D, dorsal view of internal female genitalia. Abbreviations: CD, copulatory duct; CO, conductor; CY, cymbial furrow; EM, embolus; LAM, lamellas; MA, median apophysis; PA, patellar apophyses; RTA, retrolateral tibial apophysis; SP, spermatheca. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4583722" httpUri="https://zenodo.org/record/4583722/files/figure.png" pageId="2" pageNumber="31">Figs 2C, D</figureCitation>
,
<figureCitation id="130EB7C11C5EA558FAB778E4C8654509" box="[1315,1349,1271,1295]" captionStart="Fig" captionStartId="5.[156,187,1156,1178]" captionTargetBox="[258,1330,205,1117]" captionTargetId="figure-449@5.[258,1330,205,1117]" captionTargetPageId="5" captionText="Fig. 5. Comparison of genitalic characters of three species of Dichodactylus gen. nov. from Japan: A, D, D. shinshuensis sp. nov., male (A: NSMT-Ar 13339), and female (D: NSMT-Ar 13341); B, E, D. tarumii (Arita, 1976), male (B: NSMT-Ar 14732), and female (E: NSMT-Ar 14733); C, F, D. satoi (Nishikawa, 2003), male (C: NSMT-Ar 14735), and female (F: NSMT-Ar 14734). AC, retrolateral views of male palp; DF, ventral views of epigynum. Abbreviations: CO, conductor; ET, epigynal teeth; LAM, lamellas; PA, patellar apophyses; SW, swollen portion of tegulum. Black circle on tibia shows the vestigial LTA." figureDoi="http://doi.org/10.5281/zenodo.4583728" httpUri="https://zenodo.org/record/4583728/files/figure.png" pageId="2" pageNumber="31">5D</figureCitation>
): epigynum hourglass-shaped, atrium large, almost rectangular and somewhat longer than wide, lamellae situated laterally to atrium, epigynal teeth absent, spermathecae elongated anteroposteriorly and well-separated on lateral edges of vulva, copulatory ducts prominent, large, irregularly convoluted and very visible in dorsal view of cleared vulva.
</paragraph>
</subSubSection>
<subSubSection id="C32FF8CF1C5EA558FCDA79C8C90F4612" pageId="2" pageNumber="31" type="distribution">
<paragraph id="8B8AAB441C5EA558FCDA79C8C90F4612" blockId="2.[820,1432,198,1653]" pageId="2" pageNumber="31">
<emphasis id="B94177561C5EA558FCDA79C8CEF945F5" bold="true" box="[846,985,1499,1523]" pageId="2" pageNumber="31">Distribution.</emphasis>
Chubu District, central
<collectingCountry id="F322EBD41C5EA558FB6E79CFC81245F5" box="[1274,1330,1500,1523]" name="Japan" pageId="2" pageNumber="31">Japan</collectingCountry>
(
<collectingRegion id="49F165A61C5EA558FAD379C8CE474612" country="Japan" name="Nagano" pageId="2" pageNumber="31">Nagano Pref.</collectingRegion>
,
<collectingRegion id="49F165A61C5EA558FCFB79EFCEF44612" box="[879,980,1532,1556]" country="Japan" name="Gifu" pageId="2" pageNumber="31">Gifu Pref.</collectingRegion>
) (
<figureCitation id="130EB7C11C5EA558FC7379EFC9024612" box="[999,1058,1532,1556]" captionStart="Fig" captionStartId="6.[156,187,941,963]" captionTargetBox="[340,1247,205,902]" captionTargetId="figure-443@6.[340,1247,205,902]" captionTargetPageId="6" captionText="Fig. 6. Distribution of three species of the genus Dichodactylus gen. nov. from Japan. Closed circles denote D. shinshuensis sp. nov.; closed inverted triangles, D. tarumii (Arita, 1976); crosses, D. satoi (Nishikawa, 2003)." figureDoi="http://doi.org/10.5281/zenodo.4583730" httpUri="https://zenodo.org/record/4583730/files/figure.png" pageId="2" pageNumber="31">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32FF8CF1C5EA558FCDA7A0FCEA34673" pageId="2" pageNumber="31" type="etymology">
<paragraph id="8B8AAB441C5EA558FCDA7A0FCEA34673" blockId="2.[820,1432,198,1653]" pageId="2" pageNumber="31">
<emphasis id="B94177561C5EA558FCDA7A0FCEE54632" bold="true" box="[846,965,1564,1588]" pageId="2" pageNumber="31">Etymology.</emphasis>
Specific name is an adjective derived from the provincial name of
<collectingRegion id="49F165A61C5EA558FB8A7A2EC9FA4653" box="[1054,1242,1597,1621]" country="Japan" name="Nagano" pageId="2" pageNumber="31">Nagano Prefecture</collectingRegion>
including the
<typeStatus id="548E15E61C5EA558FAF87A2EC8B84653" box="[1388,1432,1597,1621]" pageId="2" pageNumber="31">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>