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<document id="D15B9C9CAB9608471D9A93DC09921C02" ID-DOI="10.11646/zootaxa.4044.2.2" ID-GBIF-Dataset="8b09a6a1-35c3-407c-a4b5-076c4af030db" ID-ISSN="1175-5326" ID-Zenodo-Dep="241241" ID-ZooBank="7000893A-A8BC-4196-B76D-385C97B96FBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461338858631" checkinUser="plazi" docAuthor="Kanao, Taisuke &amp; Maruyama, Munetoshi" docDate="2015" docId="0395F739FFA6BF21FF021B6CFA5DBF70" docLanguage="en" docName="zt04044p223.pdf" docOrigin="Zootaxa 4044 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discoxenus" docType="key" docVersion="11" lastPageNumber="205" masterDocId="FFAC8F41FFA5BF25FF951C5CFFC1BC65" masterDocTitle="Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="223" masterPageNumber="201" pageNumber="204" updateTime="1698619795159" updateUser="plazi">
<mods:mods id="5C026D17F1717AF97584EB573251EF61" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="42B4B9299A4D0AD2E9F33103FD38FEE5">
<mods:title id="7FA5B9988238255D7AE0FBC5CE444B19">Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="FE31ABEE6559CCAAD8A779585D0866FF">Kanao, Taisuke</mods:namePart>
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<mods:name id="C09B2147CE0F4E9C9FB777E9BF0552CE" type="personal">
<mods:role id="AA2AB00F1FFC2E1ACF64421A65716B1A">
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<mods:namePart id="23CF0FBA0AD46C8CF145FB4094900FF3">Maruyama, Munetoshi</mods:namePart>
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<mods:title id="521FF4F5F118F0C44AD9A042F1097AB4">Zootaxa</mods:title>
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<mods:part id="1716EC71EBE6CF80D518B8AA9063C91A">
<mods:date id="10D60D0EB5CB32A1995C68C446368AA8">2015</mods:date>
<mods:detail id="06BEECBB42A881A3A07C9180CFF5EC20" type="volume">
<mods:number id="61373EB6651B7A3C04545CE93B73A5D6">4044</mods:number>
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<mods:number id="37C579060CC2C5B73E6EDE0A0A90B497">2</mods:number>
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<mods:start id="EE0E424FCFD8F7F5607E2AC3F4B94FD6">201</mods:start>
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<mods:classification id="B3F4922A3B0532AC6407296815ED7CE8">journal article</mods:classification>
<mods:identifier id="6286992B8A8D0138ED818BA6FCCE9A69" type="DOI">10.11646/zootaxa.4044.2.2</mods:identifier>
<mods:identifier id="25EB9F7748C820FE9E0400A555D15E30" type="GBIF-Dataset">8b09a6a1-35c3-407c-a4b5-076c4af030db</mods:identifier>
<mods:identifier id="A85B568497BBB5E747258A3E872A37E3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2D28633B42CC5B9CAEDA4F3D72DD9188" type="Zenodo-Dep">241241</mods:identifier>
<mods:identifier id="16EE6D718CD443174CA00CEAA69809A0" type="ZooBank">7000893A-A8BC-4196-B76D-385C97B96FBD</mods:identifier>
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<treatment id="0395F739FFA6BF21FF021B6CFA5DBF70" ID-DOI="http://doi.org/10.5281/zenodo.6111771" ID-GBIF-Taxon="119644718" ID-Zenodo-Dep="6111771" LSID="urn:lsid:plazi:treatment:0395F739FFA6BF21FF021B6CFA5DBF70" httpUri="http://treatment.plazi.org/id/0395F739FFA6BF21FF021B6CFA5DBF70" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
<subSubSection id="C32615A4FFA6BF26FF021B6CFD39BB2C" box="[151,760,1839,1866]" pageId="3" pageNumber="204" type="nomenclature">
<paragraph id="8B83462FFFA6BF26FF021B6CFD39BB2C" blockId="3.[151,760,1839,1866]" box="[151,760,1839,1866]" pageId="3" pageNumber="204">
<heading id="D0CBF143FFA6BF26FF021B6CFD39BB2C" bold="true" box="[151,760,1839,1866]" fontSize="11" level="1" pageId="3" pageNumber="204" reason="1">
<emphasis id="B9489A3DFFA6BF26FF021B6CFD39BB2C" bold="true" box="[151,760,1839,1866]" pageId="3" pageNumber="204">
Key to the Southeast Asian species of
<taxonomicName id="4C3C3DACFFA6BF26FDF91B73FD39BB2C" ID-CoL="8XKYL" box="[620,760,1839,1865]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B9489A3DFFA6BF26FDF91B73FD39BB2C" bold="true" box="[620,760,1839,1865]" italics="true" pageId="3" pageNumber="204">Discoxenus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFA6BF21FF021B29FA5DBF70" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204" type="key">
<keyStep id="30C85E8AFFA6BF26FF021B29FA5ABBCC" pageId="3" pageNumber="204">
<paragraph id="8B83462FFFA6BF26FF021B29FA5ABBEE" blockId="3.[151,1436,1909,2019]" box="[151,1435,1909,1931]" pageId="3" pageNumber="204">
<keyLead id="30CDE51AFFA6BF26FF021B29FA5ABBEE" box="[151,1435,1909,1931]" pageId="3" pageNumber="204">1. Macrochaetotaxy of abdominal tergites III–VIII = 6, 6, 6, 6, 6, 6............................................... 2</keyLead>
</paragraph>
<paragraph id="8B83462FFFA6BF26FF021BC8FA5ABBCC" blockId="3.[151,1436,1909,2019]" box="[151,1435,1939,1961]" pageId="3" pageNumber="204">
<keyLead id="30CDE51AFFA6BF26FF021BC8FA5ABBCC" box="[151,1435,1939,1961]" pageId="3" pageNumber="204">- Macrochaetotaxy of abdominal tergites III–VIII variable...................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA6BF21FF021BECFA5ABCAF" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
<paragraph id="8B83462FFFA6BF26FF021BECFA5DBB86" blockId="3.[151,1436,1909,2019]" pageId="3" pageNumber="204">
<keyLead id="30CDE51AFFA6BF26FF021BECFA5DBB86" pageId="3" pageNumber="204">
2. Antennal segment XI approximately 1.4 times longer than wide (
<figureCitation id="13075AAAFFA6BF26FCCC1BECFC6FBBA3" box="[857,942,1968,1990]" captionStart="FIGURES 149 – 164" captionStartId="20.[151,264,1496,1519]" captionTargetBox="[157,1431,367,1462]" captionTargetId="figure@20.[151,1436,357,1491]" captionTargetPageId="20" captionText="FIGURES 149 – 164. Discoxenus katayamai: 149. Head, 150. Antenna, 151. Labrum (left side = dorsal view, right side = ventral view), 152. Mandible, left, 153. Mandible, right, 154. Mentum, 155. Pronotum, 156. Elytron, right, 157. Tergite VIII, 158. Sternite VIII, male, 159. Sternite VIII, female, 160. Tergites IX and X, 161. Median lobe of aedeagus, ventral view, 162. Median lobe of aedeagus, lateral view, 163. Paramere, 164. Spermatheca." httpUri="https://zenodo.org/record/241253/files/figure.png" pageId="3" pageNumber="204">Fig. 150</figureCitation>
); elytra sparsely covered with relatively long setae (
<figureCitation id="13075AAAFFA6BF26FF7D1B91FEFBBB86" box="[232,314,1997,2019]" captionStart="FIGURES 149 – 164" captionStartId="20.[151,264,1496,1519]" captionTargetBox="[157,1431,367,1462]" captionTargetId="figure@20.[151,1436,357,1491]" captionTargetPageId="20" captionText="FIGURES 149 – 164. Discoxenus katayamai: 149. Head, 150. Antenna, 151. Labrum (left side = dorsal view, right side = ventral view), 152. Mandible, left, 153. Mandible, right, 154. Mentum, 155. Pronotum, 156. Elytron, right, 157. Tergite VIII, 158. Sternite VIII, male, 159. Sternite VIII, female, 160. Tergites IX and X, 161. Median lobe of aedeagus, ventral view, 162. Median lobe of aedeagus, lateral view, 163. Paramere, 164. Spermatheca." httpUri="https://zenodo.org/record/241253/files/figure.png" pageId="3" pageNumber="204">Fig. 156</figureCitation>
)..................................................................................
<taxonomicName id="4C3C3DACFFA6BF26FA8E1B92FA5DBB86" ID-CoL="6CZCM" box="[1307,1436,1998,2019]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="katayamai">
<emphasis id="B9489A3DFFA6BF26FA8E1B92FA5DBB86" box="[1307,1436,1998,2019]" italics="true" pageId="3" pageNumber="204">D. katayamai</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021CCBFA5ABCAF" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021CCBFA5ABCAF" pageId="4" pageNumber="205">
- Antennal segment IX more than 2 times longer than wide (
<figureCitation id="13075AAAFFA1BF21FC871CCBFCA3BCC8" box="[786,866,151,173]" captionStart="FIGURES 54 – 72" captionStartId="9.[151,264,1536,1559]" captionTargetBox="[157,1426,198,1505]" captionTargetId="figure@9.[151,1436,193,1515]" captionTargetPageId="9" captionText="FIGURES 54 – 72. Discoxenus phourini n. sp.: 54. Head, 55. Antenna, 56. Labrum (left side = dorsal view, right side = ventral view), 57. Mandible, left, 58. Mandible, right, 59. Maxilla, 60. Mentum, 61. Labium, 62. Pronotum, 63. Elytron, right, 64. Meso- and metaventrites, 65. Tergite VIII, 66. Sternite VIII, male, 67. Sternite VIII, female, 68. Tergites IX and X, 69. Median lobe of aedeagus, ventral view, 70. Median lobe of aedeagus, lateral view, 71. Paramere, 72. Spermatheca." httpUri="https://zenodo.org/record/241247/files/figure.png" pageId="4" pageNumber="205">Figs. 55</figureCitation>
,
<figureCitation id="13075AAAFFA1BF21FCF91CCBFC47BCC8" box="[876,902,151,173]" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="4" pageNumber="205">74</figureCitation>
,
<figureCitation id="13075AAAFFA1BF21FC051CCBFC6BBCC8" box="[912,938,151,173]" captionStart="FIGURES 89 – 104" captionStartId="13.[151,264,1283,1306]" captionTargetBox="[162,1429,201,1252]" captionTargetId="figure@13.[151,1436,193,1262]" captionTargetPageId="13" captionText="FIGURES 89 – 104. Discoxenus hirsutus n. sp.: 89. Head, 90. Antenna, 91. Labrum (left side = dorsal view, right side = ventral view), 92. Mandible, left, 93. Mandible, right, 94. Mentum, 95. Pronotum, 96. Elytron, right, 97. Tergite VIII, 98. Sternite VIII, male, 99. Sternite VIII, female, 100. Tergites IX and X. 101. Median lobe of aedeagus, ventral view, 102. Median lobe of aedeagus, lateral view, 103. Paramere, 104. Spermatheca." httpUri="https://zenodo.org/record/241249/files/figure.png" pageId="4" pageNumber="205">90</figureCitation>
,
<figureCitation id="13075AAAFFA1BF21FC211CCBFC1BBCC8" box="[948,986,151,173]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="4" pageNumber="205">106</figureCitation>
,
<figureCitation id="13075AAAFFA1BF21FC711CCBFBCBBCC8" box="[996,1034,151,173]" captionStart="FIGURES 121 – 136" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[168,1426,489,1599]" captionTargetId="figure@16.[151,1436,481,1609]" captionTargetPageId="16" captionText="FIGURES 121 – 136. Discoxenus lucidus n. sp.: 121. Head, 122. Antenna, 123. Labrum (left side = dorsal view, right side = ventral view), 124. Mandible, left, 125. Mandible, right, 126. Mentum, 127. Pronotum, 128. Elytron, right, 129. Tergite VIII, 130. Sternite VIII, male, 131. Sternite VIII, female, 132. Tergites IX and X, 133. Median lobe of aedeagus, ventral view, 134. Median lobe of aedeagus, lateral view, 135. Paramere, 136. Spermatheca." httpUri="https://zenodo.org/record/241251/files/figure.png" pageId="4" pageNumber="205">122</figureCitation>
,
<figureCitation id="13075AAAFFA1BF21FB861CCBFBFBBCC8" box="[1043,1082,151,173]" captionStart="FIGURES 137 – 148" captionStartId="18.[151,264,1794,1817]" captionTargetBox="[176,1420,892,1760]" captionTargetId="figure@18.[151,1436,879,1772]" captionTargetPageId="18" captionText="FIGURES 137 – 148. Discoxenus kakizoei n. sp.: 137. Head, 138. Antenna, 139. Labrum (left side = dorsal view, right side = ventral view), 140. Mandible, left, 141. Mandible, right, 142. Mentum, 143. Pronotum, 144. Elytron, right, 145. Tergite VIII, 146. Sternite VIII, 147. Tergites IX and X, 148. Spermatheca." httpUri="https://zenodo.org/record/241252/files/figure.png" pageId="4" pageNumber="205">138</figureCitation>
); elytra covered with no or few short setae except for anterolateral outer corner................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021C8DFA5ABD60" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021C8DFA5ABC82" blockId="4.[151,1436,151,789]" box="[151,1435,209,231]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021C8DFA5ABC82" box="[151,1435,209,231]" pageId="4" pageNumber="205">3. Abdominal sternites IV–VII with 2 rows of macrosetae....................................................... 4</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021CB3FA5ABD60" blockId="4.[151,1436,151,789]" box="[151,1435,239,261]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021CB3FA5ABD60" box="[151,1435,239,261]" pageId="4" pageNumber="205">- Abdominal sternites IV–VII with 1 row of macrosetae at posterior margin......................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021D50FA5DBD25" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021D50FA5DBD47" blockId="4.[151,1436,151,789]" box="[151,1436,268,290]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021D50FA5DBD47" box="[151,1436,268,290]" pageId="4" pageNumber="205">
4. Abdominal sternites IV–VII with 1 pair of macrosetae subapically.................................
<taxonomicName id="4C3C3DACFFA1BF21FB651D51FA9EBD47" ID-CoL="6CZC8" box="[1264,1375,269,290]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="phourini" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB651D51FA9EBD47" box="[1264,1375,269,290]" italics="true" pageId="4" pageNumber="205">D. phourini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFA1BF21FAF31D50FA5DBD47" box="[1382,1436,268,290]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B9489A3DFFA1BF21FAF31D50FA5DBD47" bold="true" box="[1382,1436,268,290]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021D76FA5DBD25" blockId="4.[151,1436,151,789]" box="[151,1436,297,320]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021D76FA5DBD25" box="[151,1436,297,320]" pageId="4" pageNumber="205">
- Abdominal sternites IV–VII with more than 5 pairs of macrosetae (10–11 macrosetae) subapically.......
<taxonomicName id="4C3C3DACFFA1BF21FB601D76FAA1BD5A" ID-CoL="6CZCJ" box="[1269,1376,298,319]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB601D76FAA1BD5A" box="[1269,1376,298,319]" italics="true" pageId="4" pageNumber="205">D. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFA1BF21FAF31D76FA5DBD25" box="[1382,1436,298,320]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B9489A3DFFA1BF21FAF31D76FA5DBD25" bold="true" box="[1382,1436,298,320]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021D1BFA5ABD1F" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021D1BFA5DBD38" blockId="4.[151,1436,151,789]" box="[151,1436,327,349]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021D1BFA5DBD38" box="[151,1436,327,349]" pageId="4" pageNumber="205">
5. Spermatheca with apical part longer than basal part (
<figureCitation id="13075AAAFFA1BF21FD4C1D1BFCEABD38" box="[729,811,327,349]" captionStart="FIGURES 137 – 148" captionStartId="18.[151,264,1794,1817]" captionTargetBox="[176,1420,892,1760]" captionTargetId="figure@18.[151,1436,879,1772]" captionTargetPageId="18" captionText="FIGURES 137 – 148. Discoxenus kakizoei n. sp.: 137. Head, 138. Antenna, 139. Labrum (left side = dorsal view, right side = ventral view), 140. Mandible, left, 141. Mandible, right, 142. Mentum, 143. Pronotum, 144. Elytron, right, 145. Tergite VIII, 146. Sternite VIII, 147. Tergites IX and X, 148. Spermatheca." httpUri="https://zenodo.org/record/241252/files/figure.png" pageId="4" pageNumber="205">Fig. 148</figureCitation>
).....................................
<taxonomicName id="4C3C3DACFFA1BF21FB661D1BFA9EBD39" ID-CoL="6CZCL" box="[1267,1375,327,348]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="kakizoei" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB661D1BFA9EBD39" box="[1267,1375,327,348]" italics="true" pageId="4" pageNumber="205">D. kakizoei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFA1BF21FAF31D1BFA5DBD38" box="[1382,1436,327,349]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B9489A3DFFA1BF21FAF31D1BFA5DBD38" bold="true" box="[1382,1436,327,349]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021D39FA5ABD1F" blockId="4.[151,1436,151,789]" box="[151,1435,356,378]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021D39FA5ABD1F" box="[151,1435,356,378]" pageId="4" pageNumber="205">
- Spermatheca with apical part shorter than basal part (
<figureCitation id="13075AAAFFA1BF21FD4A1D38FCEEBD1F" box="[735,815,356,378]" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="4" pageNumber="205">Figs. 88</figureCitation>
,
<figureCitation id="13075AAAFFA1BF21FCAC1D38FC9FBD1F" box="[825,862,356,378]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="4" pageNumber="205">120</figureCitation>
)............................................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021DDDFA5DBD95" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021DDDFA5DBDD0" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021DDDFA5DBDD0" pageId="4" pageNumber="205">
6. Median lobe of male aedeagus with distal crest moderately pointed at apex (
<figureCitation id="13075AAAFFA1BF21FC521DDDFBD1BDF2" box="[967,1040,385,407]" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="4" pageNumber="205">Fig. 86</figureCitation>
); paramere with paramerite narrow (
<figureCitation id="13075AAAFFA1BF21FAE11DDDFF3BBDD0" captionStart="FIGURES 73 – 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 – 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="4" pageNumber="205">Fig. 87</figureCitation>
)...............................................................................
<taxonomicName id="4C3C3DACFFA1BF21FB541DC3FAA1BDD1" ID-CoL="6CZCB" box="[1217,1376,415,436]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="kohkongensis" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB541DC3FAA1BDD1" box="[1217,1376,415,436]" italics="true" pageId="4" pageNumber="205">D. kohkongensis</emphasis>
</taxonomicName>
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<emphasis id="B9489A3DFFA1BF21FAF31DC3FA5DBDD0" bold="true" box="[1382,1436,415,437]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021DE1FA5DBD95" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021DE1FA5DBD95" pageId="4" pageNumber="205">
- Median lobe of male aedeagus with distal crest rounded apically (
<figureCitation id="13075AAAFFA1BF21FCC21DE0FC6BBDB7" box="[855,938,444,466]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="4" pageNumber="205">Fig. 118</figureCitation>
); paramere with paramerite broad (
<figureCitation id="13075AAAFFA1BF21FB6E1DE0FA8DBDB7" box="[1275,1356,444,466]" captionStart="FIGURES 105 – 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 – 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="4" pageNumber="205">Fig. 119</figureCitation>
)............................................................................................
<taxonomicName id="4C3C3DACFFA1BF21FB631D86FAA1BD8A" ID-CoL="6CZBW" box="[1270,1376,474,495]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB631D86FAA1BD8A" box="[1270,1376,474,495]" italics="true" pageId="4" pageNumber="205">D. minutus</emphasis>
</taxonomicName>
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<emphasis id="B9489A3DFFA1BF21FAF31D86FA5DBD95" bold="true" box="[1382,1436,474,496]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021DABFA5ABE4F" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021DABFA5ABE68" blockId="4.[151,1436,151,789]" box="[151,1435,503,525]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021DABFA5ABE68" box="[151,1435,503,525]" pageId="4" pageNumber="205">
7. Body long, length more than
<quantity id="4CC4EBCAFFA1BF21FE6F1DABFD82BE69" box="[506,579,503,525]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="205" unit="mm" value="2.3">2.3 mm</quantity>
...................................................................... 8
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021E49FA5ABE4F" blockId="4.[151,1436,151,789]" box="[151,1435,532,554]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021E49FA5ABE4F" box="[151,1435,532,554]" pageId="4" pageNumber="205">
- Body short, around
<quantity id="4CC4EBCAFFA1BF21FE0A1E48FE17BE4F" box="[415,470,532,554]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="205" unit="mm" value="2.0">2 mm</quantity>
or less.......................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021E6DFA5DBEC5" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021E6DFA5DBE00" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021E6DFA5DBE00" pageId="4" pageNumber="205">
8. Pronotum sparsely covered with macrosetae (
<figureCitation id="13075AAAFFA1BF21FD321E6DFD33BE22" box="[679,754,561,583]" captionStart="FIGURES 19 – 37" captionStartId="6.[151,264,1852,1875]" captionTargetBox="[154,1431,557,1814]" captionTargetId="figure@6.[151,1436,555,1831]" captionTargetPageId="6" captionText="FIGURES 19 – 37. Discoxenus latiabdominalis n. sp.: 19. Head, 20. Antenna, 21. Labrum (left side = dorsal view, right side = ventral view), 22. Mandible, left, 23. Mandible, right, 24. Maxilla, 25. Mentum, 26. Labium, 27. Pronotum, 28. Elytron, right, 29. Meso- and metaventrites, 30. Tergite VIII, 31. Sternite VIII, male, 32. Sternite VIII, female, 33. Tergites IX and X, 34. Median lobe of aedeagus, ventral view, 35. Median lobe of aedeagus, lateral view, 36. Paramere, 37. Spermatheca (ap: apical part, bp: basal part)." httpUri="https://zenodo.org/record/241245/files/figure.png" pageId="4" pageNumber="205">Fig. 27</figureCitation>
); macrochaetotaxy of abdominal tergites III–VIII = 6, 6, 6, 6, 6, 4; median lobe of male aedeagus with basal capsule with distal crest weakly developed (
<figureCitation id="13075AAAFFA1BF21FB821E13FB9CBE00" box="[1047,1117,591,613]" captionStart="FIGURES 19 – 37" captionStartId="6.[151,264,1852,1875]" captionTargetBox="[154,1431,557,1814]" captionTargetId="figure@6.[151,1436,555,1831]" captionTargetPageId="6" captionText="FIGURES 19 – 37. Discoxenus latiabdominalis n. sp.: 19. Head, 20. Antenna, 21. Labrum (left side = dorsal view, right side = ventral view), 22. Mandible, left, 23. Mandible, right, 24. Maxilla, 25. Mentum, 26. Labium, 27. Pronotum, 28. Elytron, right, 29. Meso- and metaventrites, 30. Tergite VIII, 31. Sternite VIII, male, 32. Sternite VIII, female, 33. Tergites IX and X, 34. Median lobe of aedeagus, ventral view, 35. Median lobe of aedeagus, lateral view, 36. Paramere, 37. Spermatheca (ap: apical part, bp: basal part)." httpUri="https://zenodo.org/record/241245/files/figure.png" pageId="4" pageNumber="205">Fig. 35</figureCitation>
)......
<taxonomicName id="4C3C3DACFFA1BF21FB391E13FA9EBE01" ID-CoL="6CZCD" box="[1196,1375,591,612]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="latiabdominalis" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB391E13FA9EBE01" box="[1196,1375,591,612]" italics="true" pageId="4" pageNumber="205">D. latiabdominalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFA1BF21FAF31E13FA5DBE00" box="[1382,1436,591,613]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B9489A3DFFA1BF21FAF31E13FA5DBE00" bold="true" box="[1382,1436,591,613]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021E31FA5DBEC5" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021E31FA5DBEC5" pageId="4" pageNumber="205">
- Pronotum with macrosetae only at lateral margin (
<figureCitation id="13075AAAFFA1BF21FD501E30FCD8BEE7" box="[709,793,620,642]" captionStart="FIGURES 121 – 136" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[168,1426,489,1599]" captionTargetId="figure@16.[151,1436,481,1609]" captionTargetPageId="16" captionText="FIGURES 121 – 136. Discoxenus lucidus n. sp.: 121. Head, 122. Antenna, 123. Labrum (left side = dorsal view, right side = ventral view), 124. Mandible, left, 125. Mandible, right, 126. Mentum, 127. Pronotum, 128. Elytron, right, 129. Tergite VIII, 130. Sternite VIII, male, 131. Sternite VIII, female, 132. Tergites IX and X, 133. Median lobe of aedeagus, ventral view, 134. Median lobe of aedeagus, lateral view, 135. Paramere, 136. Spermatheca." httpUri="https://zenodo.org/record/241251/files/figure.png" pageId="4" pageNumber="205">Fig. 127</figureCitation>
); macrochaetotaxy of abdominal tergites III–VIII = 4, 4, 4, 4, 4, 6; median lobe of male aedeagus with distal crest strongly developed (
<figureCitation id="13075AAAFFA1BF21FCFE1ED5FC7DBEFA" box="[875,956,649,671]" captionStart="FIGURES 121 – 136" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[168,1426,489,1599]" captionTargetId="figure@16.[151,1436,481,1609]" captionTargetPageId="16" captionText="FIGURES 121 – 136. Discoxenus lucidus n. sp.: 121. Head, 122. Antenna, 123. Labrum (left side = dorsal view, right side = ventral view), 124. Mandible, left, 125. Mandible, right, 126. Mentum, 127. Pronotum, 128. Elytron, right, 129. Tergite VIII, 130. Sternite VIII, male, 131. Sternite VIII, female, 132. Tergites IX and X, 133. Median lobe of aedeagus, ventral view, 134. Median lobe of aedeagus, lateral view, 135. Paramere, 136. Spermatheca." httpUri="https://zenodo.org/record/241251/files/figure.png" pageId="4" pageNumber="205">Fig. 134</figureCitation>
)..........................
<taxonomicName id="4C3C3DACFFA1BF21FB6C1ED6FA9EBEFA" ID-CoL="6CZBV" box="[1273,1375,650,671]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="lucidus" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB6C1ED6FA9EBEFA" box="[1273,1375,650,671]" italics="true" pageId="4" pageNumber="205">D. lucidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFA1BF21FAF31ED6FA5DBEC5" box="[1382,1436,650,672]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B9489A3DFFA1BF21FAF31ED6FA5DBEC5" bold="true" box="[1382,1436,650,672]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021EFBFA5ABEBF" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021EFBFA5DBED8" blockId="4.[151,1436,151,789]" box="[151,1436,679,701]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021EFBFA5DBED8" box="[151,1436,679,701]" pageId="4" pageNumber="205">
9. Macrochaetotaxy of abdominal tergites III–V = 6, 6, 6......................................
<taxonomicName id="4C3C3DACFFA1BF21FB551EFBFAA1BED9" ID-CoL="6CZCG" box="[1216,1376,679,700]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="cambodiensis" status="sp. nov.">
<emphasis id="B9489A3DFFA1BF21FB551EFBFAA1BED9" box="[1216,1376,679,700]" italics="true" pageId="4" pageNumber="205">D. cambodiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFA1BF21FAF31EFBFA5DBED8" box="[1382,1436,679,701]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B9489A3DFFA1BF21FAF31EFBFA5DBED8" bold="true" box="[1382,1436,679,701]" pageId="4" pageNumber="205">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021E99FA5ABEBF" blockId="4.[151,1436,151,789]" box="[151,1435,708,730]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021E99FA5ABEBF" box="[151,1435,708,730]" pageId="4" pageNumber="205">- Macrochaetotaxy of abdominal tergites III–V = 8, 8, 8....................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C85E8AFFA1BF21FF021EBDFA5DBF70" pageId="4" pageNumber="205">
<paragraph id="8B83462FFFA1BF21FF021EBDFA5DBE92" blockId="4.[151,1436,151,789]" box="[151,1436,737,759]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021EBDFA5DBE92" box="[151,1436,737,759]" pageId="4" pageNumber="205">
10. Macrochaetotaxy of abdominal tergites VI–V = 8, 8...........................................
<taxonomicName id="4C3C3DACFFA1BF21FB771EBEFA5DBE92" ID-CoL="6CZCC" authority="Kistner" authorityName="Kistner" box="[1250,1436,737,759]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="krishnai">
<emphasis id="B9489A3DFFA1BF21FB771EBEFA8FBE92" box="[1250,1358,738,759]" italics="true" pageId="4" pageNumber="205">D. krishnai</emphasis>
Kistner
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B83462FFFA1BF21FF021EA3FA5DBF70" blockId="4.[151,1436,151,789]" box="[151,1436,767,789]" pageId="4" pageNumber="205">
<keyLead id="30CDE51AFFA1BF21FF021EA3FA5DBF70" box="[151,1436,767,789]" pageId="4" pageNumber="205">
- Macrochaetotaxy of abdominal tergites VI–V = 6, 6.......................................
<taxonomicName id="4C3C3DACFFA1BF21FB221EA3FA5DBF70" ID-CoL="6CZC7" authority="Kistner" authorityName="Kistner" box="[1207,1436,767,789]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="malaysiensis">
<emphasis id="B9489A3DFFA1BF21FB221EA3FA8EBF71" box="[1207,1359,767,788]" italics="true" pageId="4" pageNumber="205">D. malaysiensis</emphasis>
Kistner
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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