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<document id="EC0B9D6F034EB511A8F251883A7F265A" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A3787F6AFF67FA2A8A76DEAB" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) siklesensis Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="324" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="322" updateTime="1698468866275" updateUser="ExternalLinkService">
<mods:mods id="3015C68D979C0AEAD95EBE621944A26F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0623E152D52B9DF20FEA36E562A35500">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<mods:namePart id="2B9B99B0A3E97F9B4A60947AA68AB03F">Schmidt, Joachim</mods:namePart>
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<mods:part id="02400DD4605DEFA0F22E20BC75103ED6">
<mods:date id="532548ACEF9ABA374C71B68AA913AAF7">2017</mods:date>
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<mods:number id="0CCB1CE1C341B31AFE0898E0F40BBA58">2017-09-22</mods:number>
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<mods:identifier id="32799F22A3845A77F61F516D2D7C3F00" type="DOI">10.11646/zootaxa.4323.3.1</mods:identifier>
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<treatment id="03956A03A3787F6AFF67FA2A8A76DEAB" ID-DOI="http://doi.org/10.5281/zenodo.6015753" ID-GBIF-Taxon="134839414" ID-Zenodo-Dep="6015753" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB" httpUri="http://treatment.plazi.org/id/03956A03A3787F6AFF67FA2A8A76DEAB" lastPageId="23" lastPageNumber="324" pageId="21" pageNumber="322">
<subSubSection id="C326889EA3787F68FF67FA2A8AF8DC8F" pageId="21" pageNumber="322" type="nomenclature">
<paragraph id="8B83DB15A3787F68FF67FA2A89FEDCAE" blockId="21.[151,674,1412,1471]" box="[151,674,1412,1438]" pageId="21" pageNumber="322">
<heading id="D0CB6C79A3787F68FF67FA2A89FEDCAE" bold="true" box="[151,674,1412,1438]" fontSize="11" level="1" pageId="21" pageNumber="322" reason="1">
<emphasis id="B9480707A3787F68FF67FA2A89FEDCAE" bold="true" box="[151,674,1412,1438]" pageId="21" pageNumber="322">
<taxonomicName id="4C3CA096A3787F68FF67FA2A8905DCAE" ID-CoL="58LHZ" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,601,1412,1438]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="siklesensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FF67FA2A8A1DDCAE" bold="true" box="[151,321,1412,1438]" italics="true" pageId="21" pageNumber="322">Trichotichnus</emphasis>
(
<emphasis id="B9480707A3787F68FEA2FA2A8A95DCAE" bold="true" box="[338,457,1412,1438]" italics="true" pageId="21" pageNumber="322">Bottchrus</emphasis>
)
<emphasis id="B9480707A3787F68FE2AFA2A8905DCAE" bold="true" box="[474,601,1412,1438]" italics="true" pageId="21" pageNumber="322">siklesensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FD90FA2A89FEDCAE" box="[608,674,1412,1438]" pageId="21" pageNumber="322" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B83DB15A3787F68FF67FA088AF8DC8F" blockId="21.[151,674,1412,1471]" box="[151,420,1446,1471]" pageId="21" pageNumber="322">
(
<figureCitation id="1307C790A3787F68FF6FFA088BA9DC8F" box="[159,245,1446,1471]" captionStart="FIGURES 38 – 41" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[273,1314,196,1918]" captionTargetId="figure@15.[271,1315,193,1921]" captionTargetPageId="15" captionText="FIGURES 38 – 41. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 38, T. schawalleri sp. n., holotype; 39, T. obliquebasalis sp. n., paratype, male; 40, T. siklesensis sp. n., paratype, male; 41, T. panchhaseensis sp. n., paratype, male." httpUri="https://zenodo.org/record/919620/files/figure.png" pageId="21" pageNumber="322">Figs 40</figureCitation>
,
<figureCitation id="1307C790A3787F68FEF1FA088A17DC8F" box="[257,331,1446,1471]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="21" pageNumber="322">60–66</figureCitation>
,
<figureCitation id="1307C790A3787F68FEA5FA088ADDDC8F" box="[341,385,1446,1471]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="21" pageNumber="322">151</figureCitation>
,
<figureCitation id="1307C790A3787F68FE7DFA098AC7DC8F" box="[397,411,1447,1471]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="21" pageNumber="322">
<emphasis id="B9480707A3787F68FE7DFA098AC7DC8F" box="[397,411,1447,1471]" italics="true" pageId="21" pageNumber="322">8</emphasis>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C326889EA3787F68FF67FA408831DE7B" pageId="21" pageNumber="322" type="materials_examined">
<paragraph id="8B83DB15A3787F68FF67FA408933DF1B" blockId="21.[151,1437,1518,2011]" pageId="21" pageNumber="322">
<materialsCitation id="3B54D148A3787F68FF67FA408933DF1B" ID-GBIF-Occurrence="1655986850" collectingDate="20. IV." collectionCode="HIMAL" collectorName="N Pokhara &amp; Garlang" country="Nepal" county="Jager" elevation="2300" location="Beifang" municipality="Schmidt" pageId="21" pageNumber="322" specimenCode="HIMAL 1996" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9480707A3787F68FF67FA408A1FDF37" bold="true" box="[151,323,1518,1543]" pageId="21" pageNumber="322">
<typeStatus id="548765B7A3787F68FF67FA408B8EDF37" box="[151,210,1518,1543]" pageId="21" pageNumber="322">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548765B7A3787F68FEA6FA408A9FDF37" box="[342,451,1518,1543]" pageId="21" pageNumber="322" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D3A109CA3787F68FE23FA418AB6DF36" box="[467,490,1519,1542]" pageId="21" pageNumber="322" type="male">♂</specimenCount>
, “
<collectorName id="26C9BEC3A3787F68FDF6FA4189DBDF37" box="[518,647,1518,1543]" pageId="21" pageNumber="322">N Pokhara</collectorName>
, oberh.
<collectorName id="26C9BEC3A3787F68FD1BFA408814DF37" box="[747,840,1518,1543]" pageId="21" pageNumber="322">Garlang</collectorName>
,
<elevation id="00113C26A3787F68FCA9FA4188F6DF37" box="[857,938,1519,1543]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="322" unit="m" value="2300.0">
<quantity id="4CC476F0A3787F68FCA9FA4188F6DF37" box="[857,938,1519,1543]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="322" unit="m" value="2300.0">2300m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FC4AFA418F5DDF37" box="[954,1025,1519,1543]" pageId="21" pageNumber="322">
<date id="FF82FDD5A3787F68FC4AFA418F5DDF37" box="[954,1025,1519,1543]" pageId="21" pageNumber="322">20.IV.</date>
</collectingDate>
,
<location id="8EE38DCEA3787F68FBE1FA408F33DF37" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FBE1FA408F33DF37" box="[1041,1135,1518,1543]" country="Nepal" county="Jager" municipality="Schmidt" name="Beifang" pageId="21" pageNumber="322">Beifang</location>
”, “
<collectingCountry id="F32B9B85A3787F68FB63FA418FB2DF37" box="[1171,1262,1519,1543]" name="Nepal" pageId="21" pageNumber="322">NEPAL</collectingCountry>
–
<specimenCode id="DB9A736EA3787F68FB0CFA418EC0DF37" box="[1276,1436,1518,1543]" collectionCode="HIMAL" pageId="21" pageNumber="322">HIMAL 1996</specimenCode>
<location id="8EE38DCEA3787F68FF67F9BD8A16DF1A" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FF67F9BD8A16DF1A" box="[151,330,1555,1579]" country="Nepal" county="Jager" municipality="Schmidt" name="Annapurna mts." pageId="21" pageNumber="322">Annapurna mts.</location>
lg.
<collectingMunicipality id="6BE7416FA3787F68FE85F9BC8A89DF1B" box="[373,469,1554,1579]" pageId="21" pageNumber="322">Schmidt</collectingMunicipality>
,
<collectingCounty id="62E2A399A3787F68FE10F9BD8942DF1B" box="[480,542,1555,1579]" pageId="21" pageNumber="322">Jäger</collectingCounty>
” (cJS).
</materialsCitation>
</paragraph>
<paragraph id="8B83DB15A3787F68FF37F9998831DE7B" blockId="21.[151,1437,1518,2011]" pageId="21" pageNumber="322">
<materialsCitation id="3B54D148A3787F68FF37F9998A4EDF43" ID-GBIF-Occurrence="1655986847" collectingDate="1996-04-27" collectionCode="ZIN" collectorName="J. Schmidt" elevation="2150" location="Dhara Kharka" municipality="Sikles" pageId="21" pageNumber="322" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="548765B7A3787F68FF37F9998A64DF7F" box="[199,312,1591,1615]" pageId="21" pageNumber="322" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3A109CA3787F68FEB4F9998A2CDF7E" box="[324,368,1591,1615]" pageId="21" pageNumber="322" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D3A109CA3787F68FE8FF9998AF7DF7E" box="[383,427,1591,1615]" pageId="21" pageNumber="322" type="female">2 ♀</specimenCount>
, Annapurna, Sikles range,
<location id="8EE38DCEA3787F68FD11F99888DFDF7F" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FD11F99888DFDF7F" box="[737,899,1590,1615]" municipality="Sikles" name="Dhara Kharka" pageId="21" pageNumber="322">Dhara Kharka</location>
, N
<collectingMunicipality id="6BE7416FA3787F68FC5DF99888A8DF7F" box="[941,1012,1590,1615]" pageId="21" pageNumber="322">Sikles</collectingMunicipality>
,
<elevation id="00113C26A3787F68FBF1F9988F05DF7E" box="[1025,1113,1590,1615]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="21" pageNumber="322" unit="m" value="2150.0">
<quantity id="4CC476F0A3787F68FBF1F9988F05DF7E" box="[1025,1113,1590,1615]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="21" pageNumber="322" unit="m" value="2150.0">2150 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FB97F9988FB9DF7F" box="[1127,1253,1590,1615]" pageId="21" pageNumber="322" value="1996-04-27">
<date id="FF82FDD5A3787F68FB97F9988FB9DF7F" box="[1127,1253,1590,1615]" pageId="21" pageNumber="322" value="1996-04-27">27.IV.1996</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FB02F9998E36DF7F" box="[1266,1386,1590,1615]" pageId="21" pageNumber="322">J. Schmidt</collectorName>
leg. (cJS,
<collectionCode id="ED2D43D0A3787F68FF27F9F58A56DF43" box="[215,266,1627,1651]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="21" pageNumber="322">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B54D148A3787F68FED0F9F58A57DFA7" ID-GBIF-Occurrence="1655986854" collectingDate="1996-04-28" collectorName="J. Schmidt" county="Pokhara" elevation="1850" location="Kyojo Kharka" municipality="Sikles" pageId="21" pageNumber="322" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D3A109CA3787F68FED0F9F58A10DF42" box="[288,332,1627,1651]" pageId="21" pageNumber="322" type="female">1 ♀</specimenCount>
, Annapurna, Sikles range,
<location id="8EE38DCEA3787F68FD74F9F48875DF43" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FD74F9F48875DF43" box="[644,809,1626,1651]" county="Pokhara" municipality="Sikles" name="Kyojo Kharka" pageId="21" pageNumber="322">Kyojo Kharka</location>
, N
<collectingMunicipality id="6BE7416FA3787F68FCA3F9F488C6DF43" box="[851,922,1626,1651]" pageId="21" pageNumber="322">Sikles</collectingMunicipality>
, N
<collectingCounty id="62E2A399A3787F68FC35F9F48F78DF43" box="[965,1060,1626,1651]" pageId="21" pageNumber="322">Pokhara</collectingCounty>
,
<elevation id="00113C26A3787F68FBC1F9F48FD6DF42" box="[1073,1162,1626,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="21" pageNumber="322" unit="m" value="1850.0">
<quantity id="4CC476F0A3787F68FBC1F9F48FD6DF42" box="[1073,1162,1626,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="21" pageNumber="322" unit="m" value="1850.0">1850 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FB68F9F48E4BDF43" box="[1176,1303,1626,1651]" pageId="21" pageNumber="322" value="1996-04-28">
<date id="FF82FDD5A3787F68FB68F9F48E4BDF43" box="[1176,1303,1626,1651]" pageId="21" pageNumber="322" value="1996-04-28">28.IV.1996</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FAD4F9F58EC0DF43" box="[1316,1436,1626,1651]" pageId="21" pageNumber="322">J. Schmidt</collectorName>
leg. (cJS)
</materialsCitation>
;
<materialsCitation id="3B54D148A3787F68FEEDF9D189DFDF8B" ID-GBIF-Occurrence="1655986859" collectingDate="1996-04-26" collectorName="J. Schmidt &amp; Jaeger" county="Dhara-Kharka Plateau" elevation="2200" location="Annapurna Mts." municipality="Siklis" pageId="21" pageNumber="322" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3A109CA3787F68FEEDF9D18A12DFA6" box="[285,334,1663,1687]" pageId="21" pageNumber="322" type="male">1 ♂</specimenCount>
,
<location id="8EE38DCEA3787F68FE92F9D1897DDFA6" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FE92F9D1897DDFA6" box="[354,545,1663,1687]" county="Dhara-Kharka Plateau" municipality="Siklis" name="Annapurna Mts." pageId="21" pageNumber="322">Annapurna Mts.</location>
,
<location id="8EE38DCEA3787F68FDC4F9D089A9DFA7" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FDC4F9D089A9DFA7" box="[564,757,1662,1687]" county="Dhara-Kharka Plateau" municipality="Siklis" name="Madi Khola Tal" pageId="21" pageNumber="322">Madi Khola Tal</location>
, N
<collectingMunicipality id="6BE7416FA3787F68FCD9F9D08837DFA7" box="[809,875,1662,1687]" pageId="21" pageNumber="322">Siklis</collectingMunicipality>
,
<collectingCounty id="62E2A399A3787F68FC8EF9D08FDDDFA7" box="[894,1153,1662,1687]" pageId="21" pageNumber="322">Dhara-Kharka Plateau</collectingCounty>
,
<elevation id="00113C26A3787F68FB63F9D18FB2DFA6" box="[1171,1262,1663,1687]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="322" unit="m" value="2200.0">
<quantity id="4CC476F0A3787F68FB63F9D18FB2DFA6" box="[1171,1262,1663,1687]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="322" unit="m" value="2200.0">2200 m</quantity>
</elevation>
NN,
<elevation id="00113C26A3787F68FAC9F9D18EC4DFA6" box="[1337,1432,1663,1687]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="322" unit="m" value="2200.0">
<quantity id="4CC476F0A3787F68FAC9F9D18EC4DFA6" box="[1337,1432,1663,1687]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="322" unit="m" value="2200.0">2200 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FF67F90C8A49DF8B" box="[151,277,1698,1723]" pageId="21" pageNumber="322" value="1996-04-26">
<date id="FF82FDD5A3787F68FF67F90C8A49DF8B" box="[151,277,1698,1723]" pageId="21" pageNumber="322" value="1996-04-26">26.IV.1996</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FED3F90D8AC7DF8B" box="[291,411,1698,1723]" pageId="21" pageNumber="322">J. Schmidt</collectorName>
&amp;
<collectorName id="26C9BEC3A3787F68FE33F90D8957DF8B" box="[451,523,1699,1723]" pageId="21" pageNumber="322">Jaeger</collectorName>
leg. (cJS)
</materialsCitation>
;
<materialsCitation id="3B54D148A3787F68FD60F90D8928DFEF" ID-GBIF-Occurrence="1655986856" collectingDate="2007-05-29" collectorName="Tamang &amp; Rai" county="Pokhara" elevation="2550" location="Annapurna Mts." municipality="Sikles" pageId="21" pageNumber="322" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D3A109CA3787F68FD60F90D89E0DF8A" box="[656,700,1699,1723]" pageId="21" pageNumber="322" type="female">1 ♀</specimenCount>
,
<location id="8EE38DCEA3787F68FD3CF90D88DBDF8A" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FD3CF90D88DBDF8A" box="[716,903,1699,1723]" county="Pokhara" municipality="Sikles" name="Annapurna Mts." pageId="21" pageNumber="322">Annapurna Mts.</location>
,
<location id="8EE38DCEA3787F68FC65F90C8F61DF8B" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FC65F90C8F61DF8B" box="[917,1085,1698,1723]" county="Pokhara" municipality="Sikles" name="Nyauli Kharka" pageId="21" pageNumber="322">Nyauli Kharka</location>
SW
<collectingMunicipality id="6BE7416FA3787F68FB88F90C8FE3DF8B" box="[1144,1215,1698,1723]" pageId="21" pageNumber="322">Sikles</collectingMunicipality>
, N
<collectingCounty id="62E2A399A3787F68FB1AF90C8E15DF8B" box="[1258,1353,1698,1723]" pageId="21" pageNumber="322">Pokhara</collectingCounty>
,
<elevation id="00113C26A3787F68FAA6F90D8BB1DFEE" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="21" pageNumber="322" unit="m" value="2550.0" valueMax="2700.0" valueMin="2400.0">
<quantity id="4CC476F0A3787F68FAA6F90D8BB1DFEE" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="21" pageNumber="322" unit="m" value="2550.0" valueMax="2700.0" valueMin="2400.0">2400– 2700 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FF09F9698A32DFEF" box="[249,366,1735,1759]" pageId="21" pageNumber="322" value="2007-05-29">
<date id="FF82FDD5A3787F68FF09F9698A32DFEF" box="[249,366,1735,1759]" pageId="21" pageNumber="322" value="2007-05-29">29.V.2007</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FE89F9688A8FDFEF" box="[377,467,1734,1759]" pageId="21" pageNumber="322">Tamang</collectorName>
/
<collectorName id="26C9BEC3A3787F68FE2BF968895DDFEF" box="[475,513,1734,1759]" pageId="21" pageNumber="322">Rai</collectorName>
leg. (cJS)
</materialsCitation>
;
<materialsCitation id="3B54D148A3787F68FD8FF9698AD4DE33" ID-GBIF-Occurrence="1655986835" collectingDate="1996-05-14" collectorName="J. Schmidt" county="Khilang" elevation="1900" location="Annapurna Mts." municipality="Sikles" pageId="21" pageNumber="322" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3A109CA3787F68FD8FF96989F5DFEE" box="[639,681,1735,1759]" pageId="21" pageNumber="322" type="male">1 ♂</specimenCount>
,
<location id="8EE38DCEA3787F68FD46F9698832DFEE" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FD46F9698832DFEE" box="[694,878,1735,1759]" county="Khilang" municipality="Sikles" name="Annapurna Mts." pageId="21" pageNumber="322">Annapurna Mts.</location>
,
<collectingMunicipality id="6BE7416FA3787F68FC89F96888E2DFEF" box="[889,958,1734,1759]" pageId="21" pageNumber="322">Sikles</collectingMunicipality>
range, above
<collectingCounty id="62E2A399A3787F68FBA9F9688FEADFEF" box="[1113,1206,1734,1759]" pageId="21" pageNumber="322">Khilang</collectingCounty>
,
<elevation id="00113C26A3787F68FB30F9698E4BDFEE" box="[1216,1303,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="322" unit="m" value="1900.0">
<quantity id="4CC476F0A3787F68FB30F9698E4BDFEE" box="[1216,1303,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="322" unit="m" value="1900.0">1900 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FAD2F9688ECBDFEF" box="[1314,1431,1734,1759]" pageId="21" pageNumber="322" value="1996-05-14">
<date id="FF82FDD5A3787F68FAD2F9688ECBDFEF" box="[1314,1431,1734,1759]" pageId="21" pageNumber="322" value="1996-05-14">14.V.1996</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FF67F9458A53DE33" box="[151,271,1770,1795]" pageId="21" pageNumber="322">J. Schmidt</collectorName>
leg. (cJS)
</materialsCitation>
;
<materialsCitation id="3B54D148A3787F68FE66F9458E42DE33" ID-GBIF-Occurrence="1655986843" box="[406,1310,1770,1795]" collectingDate="1997-05-30" collectorName="J. Schmidt" elevation="2500" location="Annapurna Mts." municipality="Krapa Danda" pageId="21" pageNumber="322" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D3A109CA3787F68FE66F9458A9EDE32" box="[406,450,1771,1795]" pageId="21" pageNumber="322" type="female">1 ♀</specimenCount>
,
<location id="8EE38DCEA3787F68FE23F94589D2DE32" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FE23F94589D2DE32" box="[467,654,1771,1795]" municipality="Krapa Danda" name="Annapurna Mts." pageId="21" pageNumber="322">Annapurna Mts.</location>
,
<collectingMunicipality id="6BE7416FA3787F68FD6BF9458869DE33" box="[667,821,1770,1795]" pageId="21" pageNumber="322">Krapa Danda</collectingMunicipality>
,
<elevation id="00113C26A3787F68FCB2F94488C0DE32" box="[834,924,1770,1795]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="322" unit="m" value="2500.0">
<quantity id="4CC476F0A3787F68FCB2F94488C0DE32" box="[834,924,1770,1795]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="322" unit="m" value="2500.0">2500 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FC5AF9458F7CDE33" box="[938,1056,1771,1795]" pageId="21" pageNumber="322" value="1997-05-30">
<date id="FF82FDD5A3787F68FC5AF9458F7CDE33" box="[938,1056,1771,1795]" pageId="21" pageNumber="322" value="1997-05-30">30.V.1997</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FBDDF9458FF9DE33" box="[1069,1189,1770,1795]" pageId="21" pageNumber="322">J. Schmidt</collectorName>
leg. (cJS)
</materialsCitation>
;
<materialsCitation id="3B54D148A3787F68FADCF9458F40DE17" ID-GBIF-Occurrence="1655986860" collectingDate="1997-05-27" collectionCode="ZIN" collectorName="J. Schmidt" elevation="1900" location="Annapurna Mts." municipality="Krapa Danda" pageId="21" pageNumber="322" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D3A109CA3787F68FADCF9458E06DE32" box="[1324,1370,1771,1795]" pageId="21" pageNumber="322" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D3A109CA3787F68FA99F9458EC9DE32" box="[1385,1429,1771,1795]" pageId="21" pageNumber="322" type="female">3 ♀</specimenCount>
, same data as preceding, but
<elevation id="00113C26A3787F68FE20F8A18930DE16" box="[464,620,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="21" pageNumber="322" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<quantity id="4CC476F0A3787F68FE20F8A18930DE16" box="[464,620,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="21" pageNumber="322" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FD88F8A189B1DE17" box="[632,749,1807,1831]" pageId="21" pageNumber="322" value="1997-05-27">
<date id="FF82FDD5A3787F68FD88F8A189B1DE17" box="[632,749,1807,1831]" pageId="21" pageNumber="322" value="1997-05-27">27.V.1997</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FD08F8A18831DE17" box="[760,877,1806,1831]" pageId="21" pageNumber="322">J. Schmidt</collectorName>
leg. (cJS,
<collectionCode id="ED2D43D0A3787F68FC10F8A18F4FDE17" box="[992,1043,1807,1831]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="21" pageNumber="322">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B54D148A3787F68FBD6F8A18831DE7B" ID-GBIF-Occurrence="1655986863" collectingDate="1996-05-11" collectorName="J. Schmidt" county="Pokhara" elevation="1950" location="Annapurna Mts." municipality="Sikles" pageId="21" pageNumber="322" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3A109CA3787F68FBD6F8A18F0CDE16" box="[1062,1104,1807,1831]" pageId="21" pageNumber="322" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D3A109CA3787F68FBAEF8A18FDBDE16" box="[1118,1159,1807,1831]" pageId="21" pageNumber="322" type="female">1 ♀</specimenCount>
,
<location id="8EE38DCEA3787F68FB64F8A18E11DE16" LSID="urn:lsid:plazi:treatment:03956A03A3787F6AFF67FA2A8A76DEAB:8EE38DCEA3787F68FB64F8A18E11DE16" box="[1172,1357,1807,1831]" county="Pokhara" municipality="Sikles" name="Annapurna Mts." pageId="21" pageNumber="322">Annapurna Mts.</location>
,
<collectingMunicipality id="6BE7416FA3787F68FAA7F8A08EC7DE17" box="[1367,1435,1806,1831]" pageId="21" pageNumber="322">Sikles</collectingMunicipality>
vill., N
<collectingCounty id="62E2A399A3787F68FF1DF89C8A10DE7B" box="[237,332,1842,1867]" pageId="21" pageNumber="322">Pokhara</collectingCounty>
,
<elevation id="00113C26A3787F68FEA6F89D8AAFDE7A" box="[342,499,1843,1867]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="21" pageNumber="322" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">
<quantity id="4CC476F0A3787F68FEA6F89D8AAFDE7A" box="[342,499,1843,1867]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="21" pageNumber="322" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900–2000 m</quantity>
</elevation>
,
<collectingDate id="EFC6043DA3787F68FE0FF89C892FDE7B" box="[511,627,1842,1867]" pageId="21" pageNumber="322" value="1996-05-11">
<date id="FF82FDD5A3787F68FE0FF89C892FDE7B" box="[511,627,1842,1867]" pageId="21" pageNumber="322" value="1996-05-11">11.V.1996</date>
</collectingDate>
,
<collectorName id="26C9BEC3A3787F68FD8FF89D89A9DE7B" box="[639,757,1842,1867]" pageId="21" pageNumber="322">J. Schmidt</collectorName>
leg. (cJS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C326889EA3787F6AFF37F8F888FBD855" lastPageId="23" lastPageNumber="324" pageId="21" pageNumber="322" type="description">
<paragraph id="8B83DB15A3787F68FF37F8F88F0CDE5E" blockId="21.[151,1437,1518,2011]" box="[199,1104,1878,1903]" pageId="21" pageNumber="322">
<emphasis id="B9480707A3787F68FF37F8F88A0DDE5F" bold="true" box="[199,337,1878,1903]" pageId="21" pageNumber="322">Description</emphasis>
. Habitus as in
<figureCitation id="1307C790A3787F68FE08F8F88917DE5F" box="[504,587,1878,1903]" captionStart="FIGURES 38 – 41" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[273,1314,196,1918]" captionTargetId="figure@15.[271,1315,193,1921]" captionTargetPageId="15" captionText="FIGURES 38 – 41. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 38, T. schawalleri sp. n., holotype; 39, T. obliquebasalis sp. n., paratype, male; 40, T. siklesensis sp. n., paratype, male; 41, T. panchhaseensis sp. n., paratype, male." httpUri="https://zenodo.org/record/919620/files/figure.png" pageId="21" pageNumber="322">Fig. 40</figureCitation>
. Body length
<quantity id="4CC476F0A3787F68FD17F8F88833DE5E" box="[743,879,1878,1903]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.2" metricValueMin="5.5" pageId="21" pageNumber="322" unit="mm" value="5.85" valueMax="6.2" valueMin="5.5">5.5–6.2 mm</quantity>
, width
<quantity id="4CC476F0A3787F68FC32F8F88F10DE5E" box="[962,1100,1878,1903]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="21" pageNumber="322" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 mm</quantity>
.
</paragraph>
<paragraph id="8B83DB15A3787F68FF37F8D48EDEDEA3" blockId="21.[151,1437,1518,2011]" box="[199,1410,1914,1939]" pageId="21" pageNumber="322">
Colour: Similar to that of
<taxonomicName id="4C3CA096A3787F68FE19F8D289BADEA3" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[489,742,1914,1939]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3787F68FE19F8D28AA2DEA3" box="[489,510,1916,1939]" italics="true" pageId="21" pageNumber="322">T.</emphasis>
(
<emphasis id="B9480707A3787F68FDFEF8D28943DEA3" box="[526,543,1916,1939]" italics="true" pageId="21" pageNumber="322">B</emphasis>
.)
<emphasis id="B9480707A3787F68FDC6F8D489BADEA3" box="[566,742,1914,1939]" italics="true" pageId="21" pageNumber="322">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3787F68FD1DF8D48870DEA3" box="[749,812,1914,1939]" pageId="21" pageNumber="322" rank="species">
<emphasis id="B9480707A3787F68FD1DF8D48870DEA3" bold="true" box="[749,812,1914,1939]" pageId="21" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
, femora and metatrochanteres more or less infuscate.
</paragraph>
<paragraph id="8B83DB15A3787F6BFF37F8308851DBB4" blockId="21.[151,1437,1518,2011]" lastBlockId="22.[151,1437,151,2013]" lastPageId="22" lastPageNumber="323" pageId="21" pageNumber="322">
Head: Comparatively large, with wide neck (in males, HWmax/PWmax = 0.65–0.69, HWmin/PWmax = 0.52– 0.55; in females, these indices 0.68–0.72 and 0.56–0.59, respectively). Eyes moderately or markedly convex (in males, HWmax/HWmin = 1.23–1.27, and in females, 1.18–1.25), in lateral view wide oval, occasionally almost round or slightly elongate. Tempora about one third to half as long as eye, slightly convex, sloped more or less abruptly to neck. Genae noticeably wider than width of antennomere 1, with few short setae. Lateral margin of head between antenna and eye, like in all the preceding species, slightly protruded laterally, forming an obtuse angle near eyes, distinctly bordered. Area between supraorbital furrow and upper margin of eye approximately as wide as or slightly shorter than width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow approximately by width of antennomere 3 apically. Labrum slightly concave anteriorly. Clypeus almost straight, only slightly emarginate along anterior margin, slightly convex basally, flatly sloped to apex, with a moderately large setigerous pore at each apical angle. Frons and vertex convex.
<collectingRegion id="49F815F7A37B7F6BFE4DFE738940D8C4" box="[445,540,477,500]" country="Myanmar" name="Chin" pageId="22" pageNumber="323">Mentum</collectingRegion>
and submentum completely fused, submentum on each side with one long seta and additionally with one short seta laterally of long seta. Left mandible not truncate, slightly blunted at apex. Dorsal surface smooth, highly obliterate transverse meshes recognizable only laterally behind eyes. Antennae comparatively short, with antennomeres 4–8 slightly longer than wide, in male, in female not extending to the basal edge of pronotum, with relatively shorter antennomeres.
</paragraph>
<paragraph id="8B83DB15A37B7F6BFF37FD3E8A96DD4D" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="323">Pronotum: Very moderately wide (PWmax/PL = 1.28–1.36), widest at the end of anterior third, slightly or moderately narrowed posteriad (PWmax/PWmin = 1.19–1.26), with one lateral seta inserted slightly before widest point. Sides rounded anteriorly, almost straight or more widely rounded in basal third, and slightly sinuate in very short distance just before basal angles. Apical margin slightly emarginate, bordered almost throughout or only laterally. Apical angles slightly, but clearly protruded ahead, comparatively narrowly rounded at tip. Basal margin nearly straight, vaguely bordered laterally or almost throughout, markedly longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles subdenticulate, slightly more than rectangular or almost rectangular, in some specimens with acute apex projecting laterad. Pronotal disc moderately convex, strongly sloped to apical angles and moderately so to basal ones, not depressed basally, but slightly flattened just at basal angles. Lateral gutter very narrow, slightly widened in apical third. Median line very fine, not deepened, ended markedly before apical and basal margins. Basal foveae small, oval, very shallow, usually poorly delimited. Surface sparsely, irregularly punctate in latero-basal areas before lateral seta, with few indistinct punctures or almost impunctate at anterior margin and in middle basal portion. Pronotal disc with indistinct, very obliterate microsculpture along sides.</paragraph>
<paragraph id="8B83DB15A37B7F6BFF37FB29892CDF1D" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="323">
Elytra: In lateral and caudal view convex, in dorsal view oval, comparatively elongate (in males, EL/EW = 1.43–1.47; EL/PL = 2.39–2.48; EW/PWmax = 1.21–1.29; in females, these indices 1.39–1.44, 2.35–2.45, 1.26– 1.27, respectively), widest slightly behind middle, with somewhat acute apex. Humeri prominent, angularly rounded at tip, usually with a tiny acute denticle visible from behind, in some specimens denticle almost indistinct, occasionally absent. Subapical sinuation very shallow. Sutural angle acutangular in general, almost right at apex and somewhat sharp at tip. Basal edge nearly straight, forming a sharp, obtuse angle with lateral margin. Elytral striae thin, usually finer than in
<taxonomicName id="4C3CA096A37B7F6BFE09FACF89AADC48" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[505,758,1375,1400]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37B7F6BFE09FACF895BDC48" box="[505,519,1377,1400]" italics="true" pageId="22" pageNumber="323">T</emphasis>
. (
<emphasis id="B9480707A37B7F6BFDEEFACF8973DC48" box="[542,559,1377,1400]" italics="true" pageId="22" pageNumber="323">B</emphasis>
.)
<emphasis id="B9480707A37B7F6BFDB6FAF189AADC48" box="[582,758,1375,1400]" italics="true" pageId="22" pageNumber="323">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37B7F6BFD0CFAF18867DC48" box="[764,827,1375,1400]" pageId="22" pageNumber="323" rank="species">
<emphasis id="B9480707A37B7F6BFD0CFAF18867DC48" bold="true" box="[764,827,1375,1400]" pageId="22" pageNumber="323">sp. n.</emphasis>
</taxonomicNameLabel>
, very shallow or very slightly impressed; intervals flat or slightly convex throughout, weakly narrowed posteriad. Parascutellar setigerous pore present, large. Parascutellar striole short, at most as long as distance from parascutellar pore to suture, with apex free or connected with stria 1. Interval 3 with a small discal setigerous pore at stria 2 behind the middle. Marginal umbilicate series widely interrupted at middle, with 6–8 pores in anterior group and 8–10 pores in posterior group. Microsculpture strongly reduced, without distinct meshes.
</paragraph>
<paragraph id="8B83DB15A37B7F6BFF37F999890FDF60" blockId="22.[151,1437,151,2013]" box="[199,595,1591,1616]" pageId="22" pageNumber="323">Hindwings: Reduced to tiny scales.</paragraph>
<paragraph id="8B83DB15A37B7F6BFF37F9F289D5DF8D" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="323">
Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
<figureCitation id="1307C790A37B7F6BFBB7F9F28FC4DF45" box="[1095,1176,1628,1653]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="22" pageNumber="323">Fig. 60</figureCitation>
) about as long as wide, moderately narrowed posteriad. Apex of last visible (VII) abdominal sternite in male widely rounded (
<figureCitation id="1307C790A37B7F6BFAEDF9D18E2CDFA8" box="[1309,1392,1663,1688]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="22" pageNumber="323">Fig. 61</figureCitation>
), in female somewhat angulate at apex (
<figureCitation id="1307C790A37B7F6BFDD8F90A8920DF8D" box="[552,636,1700,1725]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="22" pageNumber="323">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="8B83DB15A37B7F6BFF37F9668F3ADE7D" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="323">Legs: Protibia very finely, more or less distinctly longitudinally sulcate. Profemur with five to eight setae on anterio-ventral margin. Tarsi short, metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) markedly widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two pairs of ventro-lateral setae. In male, pro- and mesotarsi moderately widened.</paragraph>
<paragraph id="8B83DB15A37B7F6BFF37F8F98952DE89" blockId="22.[151,1437,151,2013]" pageId="22" pageNumber="323">Female genitalia: Laterotergite (hemisternite) symmetrical, with two thick setae apically. Basal stylomere with one preapical spine on external margin. Apical stylomere moderately curved, with a peg-like spine at both ventral and dorsal edges of outer margin.</paragraph>
<paragraph id="8B83DB15A37B7F6AFF37F86A88FBD855" blockId="22.[151,1437,151,2013]" lastBlockId="23.[151,1436,151,357]" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323">
Male genitalia: Median lobe of aedeagus (
<figureCitation id="1307C790A37B7F6BFD5AF86A8870DEED" box="[682,812,1988,2013]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="22" pageNumber="323">Figs 63–66</figureCitation>
) in lateral view arcuate, C-shaped, more strongly bent ventrad just after basal bulb, with ventral margin slightly convex in apical half and slightly curved ventrad at apex; in dorsal view median lobe slightly curved to the right, somewhat evenly narrowed to apex in distal half. Terminal lamella (
<figureCitation id="1307C790A37A7F6AFF08FF718A15D9C8" box="[248,329,223,248]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="23" pageNumber="324">Fig. 65</figureCitation>
) short, narrowed to apex. Apical capitulum large, in lateral view (
<figureCitation id="1307C790A37A7F6AFBD7FF718F24D9C8" box="[1063,1144,223,248]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="23" pageNumber="324">Fig. 63</figureCitation>
) more strongly prominent dorsally than ventrally, slightly oblique, with ventral flange situated more proximally than dorsal flange, in caudal view (
<figureCitation id="1307C790A37A7F6AFF2FFE898A6FD870" box="[223,307,295,320]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="23" pageNumber="324">Fig. 66</figureCitation>
) moderately wide oval. Internal sac with a post-medial group of small spines on the right side of median lobe and additionally with a small spine located more distally.
</paragraph>
</subSubSection>
<caption id="DF438B9DA37A7F6AFF67FB458A44DC6E" httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="23" pageNumber="324" targetBox="[197,1372,421,1231]" targetPageId="23">
<paragraph id="8B83DB15A37A7F6AFF67FB458A44DC6E" blockId="23.[151,1436,1259,1374]" pageId="23" pageNumber="324">
<emphasis id="B9480707A37A7F6AFF67FB458A0FDC31" bold="true" box="[151,339,1259,1282]" pageId="23" pageNumber="324">FIGURES 60–66.</emphasis>
<taxonomicName id="4C3CA096A37A7F6AFEA8FB428992DC32" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[344,718,1260,1282]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="siklesensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFEA8FB428ABFDC32" box="[344,483,1260,1282]" italics="true" pageId="23" pageNumber="324">Trichotichnus</emphasis>
(
<emphasis id="B9480707A37A7F6AFE02FB428909DC32" box="[498,597,1260,1282]" italics="true" pageId="23" pageNumber="324">Bottchrus</emphasis>
)
<emphasis id="B9480707A37A7F6AFD93FB428992DC32" box="[611,718,1260,1282]" italics="true" pageId="23" pageNumber="324">siklesensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFD24FB428850DC31" box="[724,780,1260,1281]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFD24FB428850DC31" bold="true" box="[724,780,1260,1281]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
(60–62, paratypes; 63–66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63–64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60–62, ventral view; 63, lateral view; 64–65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60–64; B: Figs 65–66).
</paragraph>
</caption>
<subSubSection id="C326889EA37A7F6AFF37FA248AC8DE3B" pageId="23" pageNumber="324" type="etymology">
<paragraph id="8B83DB15A37A7F6AFF37FA248A63DCF7" blockId="23.[151,1437,1418,1947]" pageId="23" pageNumber="324">
<emphasis id="B9480707A37A7F6AFF37FA248A1ADC93" bold="true" box="[199,326,1418,1443]" pageId="23" pageNumber="324">Etymology</emphasis>
. The specific name refers to the Sikles mounting range of the Annapurna Himal where the new species occurs.
</paragraph>
<paragraph id="8B83DB15A37A7F6AFF37FA7C8AC8DE3B" blockId="23.[151,1437,1418,1947]" pageId="23" pageNumber="324">
<emphasis id="B9480707A37A7F6AFF37FA7C8A06DCDB" bold="true" box="[199,346,1490,1515]" pageId="23" pageNumber="324">Comparison</emphasis>
. Very similar to
<taxonomicName id="4C3CA096A37A7F6AFDC0FA7A8866DCDB" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[560,826,1490,1515]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFDC0FA7A8961DCDB" box="[560,573,1492,1515]" italics="true" pageId="23" pageNumber="324">T</emphasis>
. (
<emphasis id="B9480707A37A7F6AFDABFA7A8930DCDB" box="[603,620,1492,1515]" italics="true" pageId="23" pageNumber="324">B</emphasis>
.)
<emphasis id="B9480707A37A7F6AFD7AFA7C8866DCDB" box="[650,826,1490,1515]" italics="true" pageId="23" pageNumber="324">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFCB7FA7C88D1DCDB" box="[839,909,1490,1515]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFCB7FA7C88D1DCDB" bold="true" box="[839,909,1490,1515]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
in external morphology and male genitalia.
<taxonomicName id="4C3CA096A37A7F6AFF67FA588A83DF3F" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,479,1526,1551]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="siklesensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFF67FA588A6DDF3F" box="[151,305,1526,1551]" italics="true" pageId="23" pageNumber="324">Trichotichnus</emphasis>
(
<emphasis id="B9480707A37A7F6AFEB1FA568A0EDF3F" box="[321,338,1528,1551]" italics="true" pageId="23" pageNumber="324">B</emphasis>
.)
<emphasis id="B9480707A37A7F6AFE99FA588A83DF3F" box="[361,479,1526,1551]" italics="true" pageId="23" pageNumber="324">siklesensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFE17FA58897ADF3F" box="[487,550,1526,1551]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFE17FA58897ADF3F" bold="true" box="[487,550,1526,1551]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
distinctly differs from the latter species in the terminal lamella of the aedeagus distinctly curved ventrad in lateral view and almost not constricted before the apical capitulum in dorsal view (
<figureCitation id="1307C790A37A7F6AFA9AF9B48BBDDF67" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="23" pageNumber="324">Figs 63–65</figureCitation>
); the latter is more transverse in lateral view (
<figureCitation id="1307C790A37A7F6AFD1CF9908863DF67" box="[748,831,1598,1623]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="23" pageNumber="324">Fig. 63</figureCitation>
) and slightly wider in caudal view (
<figureCitation id="1307C790A37A7F6AFB2BF9908E6DDF67" box="[1243,1329,1598,1623]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,264,1259,1282]" captionTargetBox="[197,1372,421,1231]" captionTargetId="figure@23.[174,1412,409,1238]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Trichotichnus (Bottchrus) siklesensis sp. n. (60 – 62, paratypes; 63 – 66, holotype). 60, left metepisternum; 61, apex of last (VII) abdominal sternite, male; 62, same, female; 63 – 64, median lobe of aedeagus; 65, terminal lamella; 66, apical capitulum. 60 – 62, ventral view; 63, lateral view; 64 – 65, dorsal view; 66, caudal view. Scale bars = 0.25 mm (A: Figs 60 – 64; B: Figs 65 – 66)." httpUri="https://zenodo.org/record/919634/files/figure.png" pageId="23" pageNumber="324">Fig. 66</figureCitation>
). Internal sac with post-medial group of small spines on the right side of the median lobe and additionally with a small spine located more distally; in contrast to this, in
<taxonomicName id="4C3CA096A37A7F6AFD72F92688DCDFAF" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[642,896,1670,1695]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFD72F92689D3DFAF" box="[642,655,1672,1695]" italics="true" pageId="23" pageNumber="324">T</emphasis>
. (
<emphasis id="B9480707A37A7F6AFD57F92689E4DFAF" box="[679,696,1672,1695]" italics="true" pageId="23" pageNumber="324">B</emphasis>
.)
<emphasis id="B9480707A37A7F6AFD20F92888DCDFAF" box="[720,896,1670,1695]" italics="true" pageId="23" pageNumber="324">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFC78F9288894DFAF" box="[904,968,1670,1695]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFC78F9288894DFAF" bold="true" box="[904,968,1670,1695]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
, any spines in the internal sac are absent. In addition,
<taxonomicName id="4C3CA096A37A7F6AFEEEF9028ABCDFF3" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[286,480,1706,1731]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="siklesensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFEEEF9028A6FDFF3" box="[286,307,1708,1731]" italics="true" pageId="23" pageNumber="324">T.</emphasis>
(
<emphasis id="B9480707A37A7F6AFEB2F9028A0FDFF3" box="[322,339,1708,1731]" italics="true" pageId="23" pageNumber="324">B</emphasis>
.)
<emphasis id="B9480707A37A7F6AFE9AF9048ABCDFF3" box="[362,480,1706,1731]" italics="true" pageId="23" pageNumber="324">siklesensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFE16F9048979DFF3" box="[486,549,1706,1731]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFE16F9048979DFF3" bold="true" box="[486,549,1706,1731]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
is, on average, slightly larger than
<taxonomicName id="4C3CA096A37A7F6AFC5BF9028FFBDFF3" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[939,1191,1706,1731]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFC5BF90288E4DFF3" box="[939,952,1708,1731]" italics="true" pageId="23" pageNumber="324">T</emphasis>
. (
<emphasis id="B9480707A37A7F6AFC3FF90288BCDFF3" box="[975,992,1708,1731]" italics="true" pageId="23" pageNumber="324">B</emphasis>
.)
<emphasis id="B9480707A37A7F6AFC07F9048FFBDFF3" box="[1015,1191,1706,1731]" italics="true" pageId="23" pageNumber="324">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFB5DF9048FB0DFF3" box="[1197,1260,1706,1731]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFB5DF9048FB0DFF3" bold="true" box="[1197,1260,1706,1731]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
[5.6–6.6 (mean 6.3) mm (n = 10) versus 5.5–6.2 (mean 5.9) mm (n = 10)], and its elytral striae are usually shallower and intervals are accordingly flatter.
</paragraph>
</subSubSection>
<subSubSection id="C326889EA37A7F6AFF37F8B88A76DEAB" pageId="23" pageNumber="324" type="distribution">
<paragraph id="8B83DB15A37A7F6AFF37F8B88A76DEAB" blockId="23.[151,1437,1418,1947]" pageId="23" pageNumber="324">
<emphasis id="B9480707A37A7F6AFF37F8B88A05DE1F" bold="true" box="[199,345,1814,1839]" pageId="23" pageNumber="324">Distribution</emphasis>
(
<figureCitation id="1307C790A37A7F6AFE99F8B88A96DE1F" box="[361,458,1814,1839]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="23" pageNumber="324">Fig. 151</figureCitation>
,
<figureCitation id="1307C790A37A7F6AFE27F8B98AB9DE1F" box="[471,485,1815,1839]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="23" pageNumber="324">
<emphasis id="B9480707A37A7F6AFE27F8B98AB9DE1F" box="[471,485,1815,1839]" italics="true" pageId="23" pageNumber="324">8</emphasis>
</figureCitation>
). Up to now only known from the mountain slopes both sides of Madi Khola valley south of Annapurna Peak II at altitudes of
<quantity id="4CC476F0A37A7F6AFD68F8958867DE62" box="[664,827,1851,1875]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.7" metricValueMin="1.8" pageId="23" pageNumber="324" unit="m" value="2250.0" valueMax="2700.0" valueMin="1800.0">1800–2700 m</quantity>
.
<taxonomicName id="4C3CA096A37A7F6AFCBAF8948FC0DE63" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[842,1180,1850,1875]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="siklesensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFCBAF89488B8DE63" box="[842,996,1850,1875]" italics="true" pageId="23" pageNumber="324">Trichotichnus</emphasis>
(
<emphasis id="B9480707A37A7F6AFC09F8928F56DE63" box="[1017,1034,1852,1875]" italics="true" pageId="23" pageNumber="324">B</emphasis>
.)
<emphasis id="B9480707A37A7F6AFBD6F8948FC0DE63" box="[1062,1180,1850,1875]" italics="true" pageId="23" pageNumber="324">siklesensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFB58F8948FB1DE63" box="[1192,1261,1850,1875]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFB58F8948FB1DE63" bold="true" box="[1192,1261,1850,1875]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
seems to be a geographical vicariant of
<taxonomicName id="4C3CA096A37A7F6AFE4DF8CE899CDE47" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[445,704,1886,1911]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A37A7F6AFE4DF8CE8A96DE47" box="[445,458,1888,1911]" italics="true" pageId="23" pageNumber="324">T</emphasis>
. (
<emphasis id="B9480707A37A7F6AFE15F8CE8AAADE47" box="[485,502,1888,1911]" italics="true" pageId="23" pageNumber="324">B</emphasis>
.)
<emphasis id="B9480707A37A7F6AFDE0F8F0899CDE47" box="[528,704,1886,1911]" italics="true" pageId="23" pageNumber="324">panchhaseensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA37A7F6AFD39F8F08857DE47" box="[713,779,1886,1911]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9480707A37A7F6AFD39F8F08857DE47" bold="true" box="[713,779,1886,1911]" pageId="23" pageNumber="324">sp. n.</emphasis>
</taxonomicNameLabel>
occurring on Mt. Panchhase south-west of Annapurna II (
<figureCitation id="1307C790A37A7F6AFF6FF82C8A5CDEAB" box="[159,256,1922,1947]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="23" pageNumber="324">Fig. 151</figureCitation>
,
<figureCitation id="1307C790A37A7F6AFEFCF82D8A46DEAA" box="[268,282,1923,1946]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="23" pageNumber="324">
<emphasis id="B9480707A37A7F6AFEFCF82D8A46DEAA" box="[268,282,1923,1946]" italics="true" pageId="23" pageNumber="324">7</emphasis>
</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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