524 lines
67 KiB
XML
524 lines
67 KiB
XML
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<mods:title id="A04351B2426500695CBFDCE8FE05A664">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<mods:namePart id="B243FA67BB708A37C5B5800193B64DB6">Schmidt, Joachim</mods:namePart>
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<mods:title id="133C995C867F888BEBC078F390F42FE6">Zootaxa</mods:title>
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<mods:part id="AECD9EF57252C22007472076A4376EDE">
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<mods:date id="4822B6EF10EFD402242F95BE2C18E442">2017</mods:date>
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<mods:number id="90B25197CB8A1FF49CAA9C4CC81F949E">2017-09-22</mods:number>
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<mods:number id="3B002C842E71EF9A3A49D849D83CC470">4323</mods:number>
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<treatment id="03956A03A36E7F7AFF67FA378AEEDA25" ID-DOI="http://doi.org/10.5281/zenodo.6015738" ID-GBIF-Taxon="134839420" ID-Zenodo-Dep="6015738" LSID="urn:lsid:plazi:treatment:03956A03A36E7F7AFF67FA378AEEDA25" httpUri="http://treatment.plazi.org/id/03956A03A36E7F7AFF67FA378AEEDA25" lastPageId="7" lastPageNumber="308" pageId="3" pageNumber="304">
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<subSubSection id="C326889EA36E7F7EFF67FA378AD4DCE4" pageId="3" pageNumber="304" type="nomenclature">
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<paragraph id="8B83DB15A36E7F7EFF67FA37883DDC83" blockId="3.[151,865,1433,1493]" box="[151,865,1433,1459]" pageId="3" pageNumber="304">
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<heading id="D0CB6C79A36E7F7EFF67FA37883DDC83" bold="true" box="[151,865,1433,1459]" fontSize="11" level="1" pageId="3" pageNumber="304" reason="1">
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<taxonomicName id="4C3CA096A36E7F7EFF67FA37883DDC83" ID-CoL="58LGV" authority="Kirschenhofer, 1992" authorityName="Kirschenhofer" authorityYear="1992" box="[151,865,1433,1459]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
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<emphasis id="B9480707A36E7F7EFF67FA37883DDC83" bold="true" box="[151,865,1433,1459]" pageId="3" pageNumber="304">
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<emphasis id="B9480707A36E7F7EFF67FA378A1DDC83" bold="true" box="[151,321,1433,1459]" italics="true" pageId="3" pageNumber="304">Trichotichnus</emphasis>
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||
(
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<emphasis id="B9480707A36E7F7EFEA2FA378A95DC83" bold="true" box="[338,457,1433,1459]" italics="true" pageId="3" pageNumber="304">Bottchrus</emphasis>
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)
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<emphasis id="B9480707A36E7F7EFE2AFA378902DC83" bold="true" box="[474,606,1433,1459]" italics="true" pageId="3" pageNumber="304">holzschuhi</emphasis>
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<bibRefCitation id="EFADA6E4A36E7F7EFD95FA37883DDC83" author="Kirschenhofer" box="[613,865,1433,1459]" pageId="3" pageNumber="304" refString="Kirschenhofer, E. (1992) Beitrag zur Kenntnis der Harpalinen des Himalaya: Harpalus Latreille und Trichotichnus Morawitz (Coleoptera: Carabidae). Koleopterologische Rundschau, 62, 21 - 41." type="journal article" year="1992">Kirschenhofer, 1992</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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<paragraph id="8B83DB15A36E7F7EFF67FA128AD4DCE4" blockId="3.[151,865,1433,1493]" box="[151,392,1468,1493]" pageId="3" pageNumber="304">
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(Figs 1, 5–13, 151,
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<emphasis id="B9480707A36E7F7EFE81FA128A23DCE4" box="[369,383,1468,1492]" italics="true" pageId="3" pageNumber="304">1</emphasis>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C326889EA36E7F7EFF67F9AA89F4DF08" pageId="3" pageNumber="304" type="reference_group">
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||
<paragraph id="8B83DB15A36E7F7EFF67F9AA89F4DF08" blockId="3.[151,1435,1540,1592]" pageId="3" pageNumber="304">
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||
<treatmentCitationGroup id="AB2CFC3BA36E7F7EFF67F9AA89F4DF08" pageId="3" pageNumber="304">
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||
<materialsCitation id="3B54D148A36E7F7EFF67F9AA895CDF08" ID-GBIF-Occurrence="1655986842" county="Sindhupalchok District" location="Malaphu" municipality="Tatopani" pageId="3" pageNumber="304" specimenCount="1" stateProvince="Koshi" typeStatus="holotype">
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<taxonomicName id="4C3CA096A36E7F7EFF67F9AA887EDF2A" ID-CoL="58LGV" authority="Kirschenhofer, 1992: 35" authorityName="Kirschenhofer" authorityPageNumber="35" authorityYear="1992" box="[151,802,1540,1562]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bellogenus">
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<emphasis id="B9480707A36E7F7EFF67F9AA8A7FDF2A" box="[151,291,1540,1562]" italics="true" pageId="3" pageNumber="304">Trichotichnus</emphasis>
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(
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<emphasis id="B9480707A36E7F7EFEC2F9AA8AFEDF2A" box="[306,418,1540,1562]" italics="true" pageId="3" pageNumber="304">Bellogenus</emphasis>
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||
)
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<emphasis id="B9480707A36E7F7EFE43F9AA8943DF2A" box="[435,543,1540,1562]" italics="true" pageId="3" pageNumber="304">holzschuhi</emphasis>
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<treatmentCitation id="0A9DFD04A36E7F7EFDD7F9AA887EDF2A" author="Kirschenhofer" box="[551,802,1540,1562]" page="35" pageId="3" pageNumber="304" year="1992">
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<bibRefCitation id="EFADA6E4A36E7F7EFDD7F9AA89A6DF2A" author="Kirschenhofer" box="[551,762,1540,1562]" pageId="3" pageNumber="304" refString="Kirschenhofer, E. (1992) Beitrag zur Kenntnis der Harpalinen des Himalaya: Harpalus Latreille und Trichotichnus Morawitz (Coleoptera: Carabidae). Koleopterologische Rundschau, 62, 21 - 41." type="journal article" year="1992">Kirschenhofer, 1992</bibRefCitation>
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||
: 35
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||
</treatmentCitation>
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||
</taxonomicName>
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||
.
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||
<typeStatus id="548765B7A36E7F7EFCDDF9AB8803DF2A" box="[813,863,1541,1562]" pageId="3" pageNumber="304">Type</typeStatus>
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||
locality: “
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<location id="8EE38DCEA36E7F7EFC3CF9AA8F76DF2A" LSID="urn:lsid:plazi:treatment:03956A03A36E7F7AFF67FA378AEEDA25:8EE38DCEA36E7F7EFC3CF9AA8F76DF2A" box="[972,1066,1540,1562]" county="Sindhupalchok District" municipality="Tatopani" name="Malaphu" pageId="3" pageNumber="304" stateProvince="Koshi">Malaphu</location>
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, near village
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<collectingMunicipality id="6BE7416FA36E7F7EFB45F9AA8E4CDF2A" box="[1205,1296,1540,1562]" pageId="3" pageNumber="304">Tatopani</collectingMunicipality>
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, Bhote
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<collectingRegion id="49F815F7A36E7F7EFAAFF9AA8EC7DF2A" box="[1375,1435,1540,1562]" country="Nepal" name="Kosi" pageId="3" pageNumber="304">Koshi</collectingRegion>
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Valley,
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<collectingCounty id="62E2A399A36E7F7EFEE3F98C895CDF08" box="[275,512,1570,1592]" pageId="3" pageNumber="304">Sindhupalchok District</collectingCounty>
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</materialsCitation>
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,
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<materialsCitation id="3B54D148A36E7F7EFDF9F98C89F4DF08" ID-GBIF-Occurrence="1655986848" box="[521,680,1570,1592]" country="Nepal" location="Central" pageId="3" pageNumber="304" specimenCount="1" stateProvince="Central" typeStatus="holotype">
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||
<collectingRegion id="49F815F7A36E7F7EFDF9F98C8908DF08" box="[521,596,1570,1592]" country="Sri Lanka" name="Central" pageId="3" pageNumber="304">Central</collectingRegion>
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<collectingCountry id="F32B9B85A36E7F7EFDAAF98C89C7DF08" box="[602,667,1570,1592]" name="Nepal" pageId="3" pageNumber="304">Nepal</collectingCountry>
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.”
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||
</materialsCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C326889EA36E7F7EFF67F9CB8900DE3E" pageId="3" pageNumber="304" type="materials_examined">
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||
<paragraph id="8B83DB15A36E7F7EFF67F9CB8E70DFF6" blockId="3.[151,1436,1637,2022]" pageId="3" pageNumber="304">
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||
<emphasis id="B9480707A36E7F7EFF67F9CB8A9BDF4E" bold="true" box="[151,455,1637,1662]" pageId="3" pageNumber="304">
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||
<typeStatus id="548765B7A36E7F7EFF67F9CB8B8EDF4E" box="[151,210,1637,1662]" pageId="3" pageNumber="304">Type</typeStatus>
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||
material examined
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||
</emphasis>
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||
.
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||
<typeStatus id="548765B7A36E7F7EFE2DF9CB8916DF4E" box="[477,586,1637,1662]" pageId="3" pageNumber="304" type="holotype">Holotype</typeStatus>
|
||
:
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||
<specimenCount id="9D3A109CA36E7F7EFDACF9C8892FDF4D" box="[604,627,1638,1661]" pageId="3" pageNumber="304" type="male">♂</specimenCount>
|
||
, “C-Nepal,
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||
<collectingRegion id="49F815F7A36E7F7EFCF8F9CB883BDF4E" box="[776,871,1637,1662]" country="Nepal" name="Bagmati" pageId="3" pageNumber="304">Bagmati</collectingRegion>
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||
Bhote
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||
<collectingRegion id="49F815F7A36E7F7EFC38F9CB8F57DF4E" box="[968,1035,1637,1662]" country="Nepal" name="Kosi" pageId="3" pageNumber="304">Koshi</collectingRegion>
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||
khol. [sic], Tatophani [Tatopani] Malaphu,
|
||
<quantity id="4CC476F0A36E7F7EFEE4F9248A34DF91" box="[276,360,1674,1698]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="304" unit="m" value="700.0">700 m</quantity>
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,
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<date id="FF82FDD5A36E7F7EFE70F9248ABADF92" box="[384,486,1674,1698]" pageId="3" pageNumber="304" value="1987-06-30">30.VI.87</date>
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, lg. Holzschuh”, “
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<typeStatus id="548765B7A36E7F7EFD24F9278812DF92" box="[724,846,1673,1698]" pageId="3" pageNumber="304" type="holotype">Holotypus</typeStatus>
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||
,
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<taxonomicName id="4C3CA096A36E7F7EFC94F9278E76DF92" authorityName="Kirschenhofer" authorityYear="1992" box="[868,1322,1673,1698]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bellogenus">
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||
<emphasis id="B9480707A36E7F7EFC94F92788A1DF92" box="[868,1021,1673,1698]" italics="true" pageId="3" pageNumber="304">Trichotichnus</emphasis>
|
||
(
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||
<emphasis id="B9480707A36E7F7EFBEAF9278FCADF92" box="[1050,1174,1673,1698]" italics="true" pageId="3" pageNumber="304">Bellogenus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A36E7F7EFB42F9278E76DF92" box="[1202,1322,1673,1698]" italics="true" pageId="3" pageNumber="304">holzschuhi</emphasis>
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||
</taxonomicName>
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||
m., det. Kirschenhofer, 91” (
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||
<collectionCode id="ED2D43D0A36E7F7EFE8BF9008A82DFF6" box="[379,478,1710,1734]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="304">NHMW</collectionCode>
|
||
).
|
||
<typeStatus id="548765B7A36E7F7EFE00F900890ADFF6" box="[496,598,1710,1734]" pageId="3" pageNumber="304" type="paratype">Paratype</typeStatus>
|
||
:
|
||
<specimenCount id="9D3A109CA36E7F7EFD90F90089D1DFF5" box="[608,653,1710,1734]" pageId="3" pageNumber="304" type="male">1 ♂</specimenCount>
|
||
, same data as
|
||
<typeStatus id="548765B7A36E7F7EFCDDF90388CEDFF6" box="[813,914,1709,1734]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
|
||
, but labelled “
|
||
<typeStatus id="548765B7A36E7F7EFBC2F9008FF9DFF6" box="[1074,1189,1710,1734]" pageId="3" pageNumber="304" type="paratype">Paratypus</typeStatus>
|
||
” (
|
||
<collectionCode id="ED2D43D0A36E7F7EFB4DF9008E7CDFF6" box="[1213,1312,1710,1734]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="304">NHMW</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A36E7F7EFF37F97F8900DE3E" blockId="3.[151,1436,1637,2022]" pageId="3" pageNumber="304">
|
||
<materialsCitation id="3B54D148A36E7F7EFF37F97F8900DE3E" ID-GBIF-Occurrence="1655986831" collectorName="S Khumbu-Himal" country="Nepal" elevation="2800" location="NEPAL" pageId="3" pageNumber="304" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
|
||
<emphasis id="B9480707A36E7F7EFF37F97F8960DFDA" bold="true" box="[199,572,1745,1770]" pageId="3" pageNumber="304">Additional material examined.</emphasis>
|
||
<specimenCount id="9D3A109CA36E7F7EFDB7F97C892BDFD9" box="[583,631,1746,1770]" pageId="3" pageNumber="304" type="male">3 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA36E7F7EFD78F97C89EADFD9" box="[648,694,1746,1770]" pageId="3" pageNumber="304" type="female">2 ♀</specimenCount>
|
||
, “Sete, Umgeb.
|
||
<elevation id="00113C26A36E7F7EFC8CF97F8F47DFD9" box="[892,1051,1745,1770]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="3" pageNumber="304" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">
|
||
<quantity id="4CC476F0A36E7F7EFC8CF97F8F47DFD9" box="[892,1051,1745,1770]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="3" pageNumber="304" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">2600–3000 m</quantity>
|
||
</elevation>
|
||
27.4.”, “
|
||
<collectingCountry id="F32B9B85A36E7F7EFB76F97C8F83DFDA" box="[1158,1247,1746,1770]" name="Nepal" pageId="3" pageNumber="304">NEPAL</collectingCountry>
|
||
–HIMALAYA
|
||
<collectorName id="26C9BEC3A36E7F7EFA7CF97C8A14DE3E" pageId="3" pageNumber="304">S Khumbu-Himal</collectorName>
|
||
lg. Kleeberg ' 93” (cJS).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA36E7F7BFF37F8B78979DED6" lastPageId="6" lastPageNumber="307" pageId="3" pageNumber="304" type="description">
|
||
<paragraph id="8B83DB15A36E7F7EFF37F8B78ED9DE65" blockId="3.[151,1436,1637,2022]" pageId="3" pageNumber="304">
|
||
<emphasis id="B9480707A36E7F7EFF37F8B78A0DDE02" bold="true" box="[199,337,1817,1842]" pageId="3" pageNumber="304">Description</emphasis>
|
||
. Habitus as in Fig. 1. Body length in
|
||
<typeStatus id="548765B7A36E7F7EFCF7F8B78835DE02" box="[775,873,1817,1842]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CC476F0A36E7F7EFC82F8B78890DE01" box="[882,972,1817,1842]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="304" unit="mm" value="5.9">5.9 mm</quantity>
|
||
, in
|
||
<typeStatus id="548765B7A36E7F7EFC09F8B48F05DE02" box="[1017,1113,1818,1842]" pageId="3" pageNumber="304" type="paratype">paratype</typeStatus>
|
||
<quantity id="4CC476F0A36E7F7EFB92F8B78FE0DE01" box="[1122,1212,1817,1842]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="304" unit="mm" value="6.2">6.2 mm</quantity>
|
||
, in specimens from Sete
|
||
<quantity id="4CC476F0A36E7F7EFF3EF8938A04DE65" box="[206,344,1853,1878]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.1" metricValueMin="5.6" pageId="3" pageNumber="304" unit="mm" value="5.85" valueMax="6.1" valueMin="5.6">5.6–6.1 mm</quantity>
|
||
; body width in
|
||
<typeStatus id="548765B7A36E7F7EFDF6F8938934DE66" box="[518,616,1853,1878]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CC476F0A36E7F7EFD9FF893899BDE65" box="[623,711,1853,1878]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="304" unit="mm" value="2.6">2.6 mm</quantity>
|
||
, in
|
||
<typeStatus id="548765B7A36E7F7EFD1FF890880CDE66" box="[751,848,1854,1878]" pageId="3" pageNumber="304" type="paratype">paratype</typeStatus>
|
||
<quantity id="4CC476F0A36E7F7EFCA7F89088F3DE65" box="[855,943,1854,1878]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="304" unit="mm" value="2.7">2.7 mm</quantity>
|
||
, and in specimens from Sete
|
||
<quantity id="4CC476F0A36E7F7EFB07F8938EDDDE65" box="[1271,1409,1853,1878]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="3" pageNumber="304" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A36E7F7EFF37F8CF895FDED6" blockId="3.[151,1436,1637,2022]" pageId="3" pageNumber="304">Colour: Body piceous, shiny on dorsum, with light bluish tinge on elytra and pronotum; labrum, in most specimens also base of mandibles, very narrow margins of pronotum, and elytra paler; epipleurae of elytra usually also much paler than elytral disc, reddish brown; palpi, antennae and legs brownish yellow, with femora usually more or less distinctly infuscate.</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37FF398EC1DBB5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
Head: Large, with wide neck (in males, HWmax/PWmax = 0.64–0.71, HWmin/PWmax = 0.56–0.59; in females, these indices 0.64–0.68 and 0.56–0.58, respectively). Eyes moderately convex (in males, HWmax/ HWmin = 1.18–1.26, and in females, 1.18–1.22), in lateral view oval, slightly elongate. Tempora comparatively long, about half as long as eye, more or less flat, somewhat abrupt to neck. Genae wide, much wider than width of antennomere 1, with few short setae. Lateral margin of head between antenna and eye more or less markedly protruded laterally, forming a small obtuse angle near eyes, distinctly bordered. Area between supraorbital furrow and upper margin of eye comparatively wide, approximately as wide as width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow at least by width of antennomere 3 apically. Labrum slightly or moderately concave anteriorly. Clypeus slightly emarginate along anterior margin, moderately convex basally, flatly sloped to apex. Frons and vertex convex.
|
||
<collectingRegion id="49F815F7A3697F79FF05FDAE8A08DB27" box="[245,340,512,535]" country="Myanmar" name="Chin" pageId="4" pageNumber="305">Mentum</collectingRegion>
|
||
(as in Fig. 14) and submentum completely fused, submentum on each side with one long seta and occasionally one shorter seta laterally of the long seta. Left mandible not truncate, at most only slightly blunted at apex. Dorsal microsculpture visible only laterally above and behind eyes, consisting of transverse meshes. Antennae short, slightly not extending to the pronotal basal edge, with antennomeres 5–8 slightly longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37FD3E8ECDDADD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">Pronotum: Slightly transverse (PWmax/PL = 1.34–1.39), widest slightly before the end of anterior third, narrowed posteriad (PWmax/PWmin = 1.17–1.25), with one lateral seta inserted before widest point. Sides almost straight or slightly sinuate in posterior third. Apical margin very slightly emarginate, with apical border interrupted at middle. Apical angles narrowly rounded, slightly protruded anteriad. Basal margin very nearly straight or slightly oblique laterally, vaguely bordered almost throughout, somewhat longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles obtuse, sharp at apex, usually without distinct denticle at tip. Pronotal disc convex, strongly sloped to apical angles, not depressed basally, slightly flattened at basal angles. Lateral gutter slightly widened in apical third. Median line distinct, fine, ended before apical and basal margins. Basal foveae oval, poorly delimited, very shallow. Surface sparsely and irregularly punctate only along base and in latero-basal area till lateral seta. Microsculpture evident throughout, consisting of fine transverse meshes, more distinct latero-basally.</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37FC598889DC48" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">Elytra: In lateral and caudal view convex, in dorsal view oval, relatively wide and short (in males, EL/EW = 1.33–1.38; EL/PL = 2.29–2.42; EW/PWmax = 1.25–1.31; in females, these indices 1.32–1.39, 2.34–2.41, 1.25– 1.29, respectively), widest slightly behind middle, with somewhat acute apex. Humeri prominent, angularly rounded at tip, without denticle. Subapical sinuation very shallow. Sutural angle acutangular, somewhat sharp or slightly blunted at tip. Basal edge sinuate laterally, forming an obtuse angle with lateral margin. Striae slightly impressed along entire length or superficial. Parascutellar setigerous pore present, large. Parascutellar striole short, as long as or slightly longer than distance from parascutellar pore to suture, with apex free or occasionally connected with stria 1. Intervals slightly convex, slightly narrowed posteriad. Interval 3 with or without a discal setigerous pore at stria 2 behind middle. Marginal umbilicate series widely interrupted at middle, with 7–8 pores in anterior group and 8–9 pores in posterior group. Microsculpture hardly visible, particularly on inner intervals, more distinct on two lateral intervals, consisting of very fine transverse meshes.</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37FA2A890FDCAD" blockId="4.[151,1437,151,2013]" box="[199,595,1412,1437]" pageId="4" pageNumber="305">Hindwings: Reduced to tiny scales.</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37FA0989DCDF38" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
|
||
<figureCitation id="1307C790A3697F79FB90FA098FE8DCF0" box="[1120,1204,1447,1472]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="4" pageNumber="305">Fig. 10</figureCitation>
|
||
) slightly wider than long, strongly narrowed posteriad. Apex of last visible (VII) abdominal sternite in male somewhat widely (
|
||
<figureCitation id="1307C790A3697F79FACCFA628ED2DCD4" box="[1340,1422,1484,1509]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="4" pageNumber="305">Fig. 12</figureCitation>
|
||
), in female more narrowly (
|
||
<figureCitation id="1307C790A3697F79FE4CFA418951DF38" box="[444,525,1519,1544]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="4" pageNumber="305">Fig. 13</figureCitation>
|
||
) rounded.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37F9BB880CDFA8" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">Legs: Protibia without longitudinal sulcus on upper surface. Profemur with three or four setae on anterioventral margin. Tarsi short, metatarsus noticeably shorter than HWmin, with metatarsomeres markedly widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two pairs of ventrolateral setae. In male, pro- and mesotarsi moderately widened.</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37F90A89EBDE35" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">Female genitalia: Laterotergite (hemisternite) somewhat symmetrical, with two thick setae apically. Basal stylomere with one preapical spine on external margin. Apical stylomere moderately curved, with a peg-like spine at both ventral and dorsal edges of outer margin.</paragraph>
|
||
<paragraph id="8B83DB15A3697F79FF37F8A18E34DEED" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
Male genitalia: Median lobe of aedeagus (
|
||
<figureCitation id="1307C790A3697F79FD57F8A1886ADE18" box="[679,822,1807,1832]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="4" pageNumber="305">Figs 6–8, 11</figureCitation>
|
||
) in lateral view rather evenly arcuate, C-shaped, with apical portion slightly curved ventrad, in dorsal view almost straight, strongly narrowed to apex in distal third. Terminal lamella (
|
||
<figureCitation id="1307C790A3697F79FE99F8F98AF1DE40" box="[361,429,1879,1904]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="4" pageNumber="305">Fig. 5</figureCitation>
|
||
) moderately long, narrowed slightly to apex, at most only very slightly sinuate at sides before apical capitulum. The latter in lateral view (
|
||
<figureCitation id="1307C790A3697F79FD3EF8D28861DEA4" box="[718,829,1916,1941]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="4" pageNumber="305">Figs 6, 11</figureCitation>
|
||
) oblique, with acute dorsal flange located less distally than rounded ventral flange; in caudal view (
|
||
<figureCitation id="1307C790A3697F79FD65F80E8985DE88" box="[661,729,1952,1977]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="4" pageNumber="305">Fig. 9</figureCitation>
|
||
) semi-oval, rounded at tip. Internal sac with two small groups (sometimes connected forming one narrow group) of very small spines just behind the middle of median lobe.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3687F78FF67F80F8AB2DEC5" pageId="5" pageNumber="306">
|
||
<table id="F93C29B5A3688082FF67F80F8EC1DEC5" box="[151,1437,1953,2037]" gridcols="4" gridrows="3" pageId="5" pageNumber="306">
|
||
<tr id="350CD957A3688082FF67F80F8EC1DE88" box="[151,1437,1953,1976]" gridrow="0" pageId="5" pageNumber="306">
|
||
<th id="76DDB02BA3688082FF67F80F8943DE88" box="[151,543,1953,1976]" gridcol="0" gridrow="0" pageId="5" pageNumber="306">
|
||
<emphasis id="B9480707A3687F78FF67F80F8A18DE87" bold="true" box="[151,324,1953,1976]" pageId="5" pageNumber="306">FIGURES 1–4.</emphasis>
|
||
Himalayan species
|
||
</th>
|
||
<th id="76DDB02BA3688082FDD8F80F89BADE88" box="[552,742,1953,1976]" gridcol="1" gridrow="0" pageId="5" pageNumber="306">
|
||
of
|
||
<taxonomicName id="4C3CA096A3687F78FDA3F80C8982DE88" authorityName="Morawitz" authorityYear="1863" box="[595,734,1954,1976]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A3687F78FDA3F80C8982DE88" box="[595,734,1954,1976]" italics="true" pageId="5" pageNumber="306">Trichotichnus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76DDB02BA3688082FD1EF80F8E7FDE88" box="[750,1315,1953,1976]" gridcol="2" gridrow="0" pageId="5" pageNumber="306">
|
||
subgenus
|
||
<taxonomicName id="4C3CA096A3687F78FCADF80C889CDE88" authorityName="Jedlicka" authorityYear="1935" box="[861,960,1954,1976]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A3687F78FCADF80C889CDE88" box="[861,960,1954,1976]" italics="true" pageId="5" pageNumber="306">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
, photographs of habitus. 1,
|
||
<emphasis id="B9480707A3687F78FAFCF80D8E7CDE88" box="[1292,1312,1955,1976]" italics="true" pageId="5" pageNumber="306">T.</emphasis>
|
||
</th>
|
||
<th id="76DDB02BA3688082FADCF80F8EC1DE88" box="[1324,1437,1953,1976]" gridcol="3" gridrow="0" pageId="5" pageNumber="306">
|
||
<taxonomicName id="4C3CA096A3687F78FADFF80C8EC7DE88" authority="Kirschenhofer, 1992" authorityName="Kirschenhofer" authorityYear="1992" box="[1327,1435,1954,1976]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="holzschuhi">
|
||
<emphasis id="B9480707A3687F78FADFF80C8EC7DE88" box="[1327,1435,1954,1976]" italics="true" pageId="5" pageNumber="306">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="350CD957A3688082FF67F86F8EC1DEE7" box="[151,1437,1985,2007]" gridrow="1" pageId="5" pageNumber="306">
|
||
<th id="76DDB02BA3688082FF67F86F8943DEE7" box="[151,543,1985,2007]" gridcol="0" gridrow="1" pageId="5" pageNumber="306">Kirschenhofer, 1992, male from Sete;</th>
|
||
<td id="76DDB02BA3688082FDD8F86F89BADEE7" box="[552,742,1985,2007]" gridcol="1" gridrow="1" pageId="5" pageNumber="306">
|
||
2,
|
||
<taxonomicName id="4C3CA096A3687F78FDB4F86C89BADEE7" authorityName="Schmidt" authorityYear="2017" box="[580,742,1985,2007]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="baglungensis">
|
||
<emphasis id="B9480707A3687F78FDB4F86C89BADEE7" box="[580,742,1985,2007]" italics="true" pageId="5" pageNumber="306">T. baglungensis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76DDB02BA3688082FD1EF86F8E7FDEE7" box="[750,1315,1985,2007]" gridcol="2" gridrow="1" pageId="5" pageNumber="306">
|
||
<emphasis id="B9480707A3687F78FD1EF86F8876DEE6" bold="true" box="[750,810,1985,2006]" pageId="5" pageNumber="306">sp. n.</emphasis>
|
||
, paratype, male; 3,
|
||
<taxonomicName id="4C3CA096A3687F78FC0DF86C8F2DDEE7" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1021,1137,1985,2007]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
|
||
<emphasis id="B9480707A3687F78FC0DF86C8F2DDEE7" box="[1021,1137,1985,2007]" italics="true" pageId="5" pageNumber="306">T. parvulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3687F78FB8AF86F8FE9DEE6" box="[1146,1205,1985,2006]" pageId="5" pageNumber="306" rank="species">
|
||
<emphasis id="B9480707A3687F78FB8AF86F8FE9DEE6" bold="true" box="[1146,1205,1985,2006]" pageId="5" pageNumber="306">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype,
|
||
</td>
|
||
<td id="76DDB02BA3688082FADCF86F8EC1DEE7" box="[1324,1437,1985,2007]" gridcol="3" gridrow="1" pageId="5" pageNumber="306">
|
||
male; 4,
|
||
<emphasis id="B9480707A3687F78FA79F86C8EC1DEE7" box="[1417,1437,1986,2007]" italics="true" pageId="5" pageNumber="306">T.</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="350CD957A3688082FF67F8718EC1DEC5" box="[151,1437,2015,2037]" gridrow="2" pageId="5" pageNumber="306" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76DDB02BA3688082FF67F8718943DEC5" box="[151,543,2015,2037]" gridcol="0" gridrow="2" pageId="5" pageNumber="306">
|
||
<taxonomicName id="4C3CA096A3687F78FF67F8718A5ADEC5" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,262,2015,2037]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="bubsaensis" status="sp. nov.">
|
||
<emphasis id="B9480707A3687F78FF67F8718A5ADEC5" box="[151,262,2015,2037]" italics="true" pageId="5" pageNumber="306">bubsaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3687F78FEFCF84E8A19DEC5" box="[268,325,2016,2037]" pageId="5" pageNumber="306" rank="species">
|
||
<emphasis id="B9480707A3687F78FEFCF84E8A19DEC5" bold="true" box="[268,325,2016,2037]" pageId="5" pageNumber="306">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male.
|
||
</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF438B9DA36B7F7BFF67FB0289ABDC2E" httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="6" pageNumber="307" targetBox="[152,1425,180,1152]" targetPageId="6">
|
||
<paragraph id="8B83DB15A36B7F7BFF67FB0289ABDC2E" blockId="6.[151,1436,1196,1310]" pageId="6" pageNumber="307">
|
||
<emphasis id="B9480707A36B7F7BFF67FB028A15DDF2" bold="true" box="[151,329,1196,1219]" pageId="6" pageNumber="307">FIGURES 5–13.</emphasis>
|
||
<taxonomicName id="4C3CA096A36B7F7BFEA2FB0288F6DDF2" authority="Kirschenhofer, 1992" authorityName="Kirschenhofer" authorityYear="1992" box="[338,938,1196,1218]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFEA2FB028A81DDF2" box="[338,477,1196,1218]" italics="true" pageId="6" pageNumber="307">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFE1FFB02890DDDF2" box="[495,593,1196,1218]" italics="true" pageId="6" pageNumber="307">Bottchrus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A36B7F7BFD93FB028993DDF2" box="[611,719,1196,1218]" italics="true" pageId="6" pageNumber="307">holzschuhi</emphasis>
|
||
Kirschenhofer, 1992
|
||
</taxonomicName>
|
||
(5–7, paratype; 8–13, specimens from Sete). 5, terminal lamella of aedeagus; 6–8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5–8, 10–13; B: Fig. 9).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B83DB15A36B7F7BFF37FAEB8979DED6" blockId="6.[151,1437,1349,2022]" pageId="6" pageNumber="307">
|
||
<emphasis id="B9480707A36B7F7BFF37FAEB8A06DC6E" bold="true" box="[199,346,1349,1374]" pageId="6" pageNumber="307">Comparison</emphasis>
|
||
. This species is distinctly recognized from other brachypterous species of the subgenus
|
||
<taxonomicName id="4C3CA096A36B7F7BFADFFAEB8EC0DC6E" authorityName="Jedlicka" authorityYear="1935" box="[1327,1436,1349,1374]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A36B7F7BFADFFAEB8EC0DC6E" box="[1327,1436,1349,1374]" italics="true" pageId="6" pageNumber="307">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
of the Himalayan region by having small size (
|
||
<quantity id="4CC476F0A36B7F7BFD30FAC4880DDCB2" box="[704,849,1386,1411]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.2" metricValueMin="5.6" pageId="6" pageNumber="307" unit="mm" value="5.9" valueMax="6.2" valueMin="5.6">5.6–6.2 mm</quantity>
|
||
), completely fused mentum and submentum, short metepisternum (slightly wider than long), not sulcate protibia, fine microsculpture on pronotum and elytra, and evenly arcuate, C-shaped median lobe of aedeagus with an oblique apical capitulum, and with two groups of very small spines in the internal sac. Among the species previously described,
|
||
<taxonomicName id="4C3CA096A36B7F7BFC21FA798FCADCDE" authorityName="Kirschenhofer" authorityYear="1992" box="[977,1174,1493,1518]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFC21FA7988B9DCDE" box="[977,997,1495,1518]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFC06FA798F5BDCDE" box="[1014,1031,1495,1518]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFBEFFA7B8FCADCDE" box="[1055,1174,1493,1518]" italics="true" pageId="6" pageNumber="307">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
is somewhat similar in appearance to
|
||
<taxonomicName id="4C3CA096A36B7F7BFEB0FA55894DDF22" authorityName="Ito" authorityYear="1998" box="[320,529,1530,1554]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFEB0FA558A09DF22" box="[320,341,1531,1554]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFE98FA558A25DF22" box="[360,377,1531,1554]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFE64FA55894DDF22" box="[404,529,1531,1554]" italics="true" pageId="6" pageNumber="307">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
from East
|
||
<collectingCountry id="F32B9B85A36B7F7BFD6AFA5489BCDF23" box="[666,736,1530,1555]" name="Nepal" pageId="6" pageNumber="307">Nepal</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C3CA096A36B7F7BFD1EFA5588DFDF23" authorityName="Ito" authorityYear="1998" box="[750,899,1530,1555]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="loebli" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFD1EFA55885FDF22" box="[750,771,1531,1554]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFCE7FA558874DF22" box="[791,808,1531,1554]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFCB3FA5488DFDF23" box="[835,899,1530,1555]" italics="true" pageId="6" pageNumber="307">loebli</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49F815F7A36B7F7BFC3EFA558F3CDF23" box="[974,1120,1530,1555]" country="India" name="West Bengal" pageId="6" pageNumber="307">West Bengal</collectingRegion>
|
||
and
|
||
<taxonomicName id="4C3CA096A36B7F7BFB6EFA558E07DF23" authorityName="Andrewes" authorityYear="1930" box="[1182,1371,1530,1555]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFB6EFA558FEFDF22" box="[1182,1203,1531,1554]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFB36FA558F8BDF22" box="[1222,1239,1531,1554]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFB03FA548E07DF23" box="[1267,1371,1530,1555]" italics="true" pageId="6" pageNumber="307">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49F815F7A36B7F7BFF67F9B38BB7DF06" box="[151,235,1565,1590]" country="India" name="Sikkim" pageId="6" pageNumber="307">Sikkim</collectingRegion>
|
||
, but all three species are larger (
|
||
<quantity id="4CC476F0A36B7F7BFDABF9B389BADF06" box="[603,742,1565,1590]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.7" metricValueMin="6.3" pageId="6" pageNumber="307" unit="mm" value="7.0" valueMax="7.7" valueMin="6.3">6.3–7.7 mm</quantity>
|
||
) and with protibia more or less clearly sulcate on dorsal side basally. In addition,
|
||
<taxonomicName id="4C3CA096A36B7F7BFE8CF9ED8915DF6A" authorityName="Ito" authorityYear="1998" box="[380,585,1602,1626]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFE8CF9ED8ACEDF6A" box="[380,402,1603,1626]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFE52F9ED8AEFDF6A" box="[418,435,1603,1626]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFE3BF9ED8915DF6A" box="[459,585,1603,1626]" italics="true" pageId="6" pageNumber="307">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3CA096A36B7F7BFD71F9ED8853DF6B" authorityName="Ito" authorityYear="1998" box="[641,783,1602,1627]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="loebli" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFD71F9ED89CADF6A" box="[641,662,1603,1626]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFD56F9ED89EBDF6A" box="[678,695,1603,1626]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFD3FF9EC8853DF6B" box="[719,783,1602,1627]" italics="true" pageId="6" pageNumber="307">loebli</emphasis>
|
||
</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="4C3CA096A36B7F7BFC6AF9ED8F3DDF6B" authorityName="Kirschenhofer" authorityYear="1992" box="[922,1121,1602,1627]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFC6AF9ED88F2DF6A" box="[922,942,1603,1626]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFC30F9ED888DDF6A" box="[960,977,1603,1626]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFC19F9EC8F3DDF6B" box="[1001,1121,1602,1627]" italics="true" pageId="6" pageNumber="307">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
in mentum and submentum separated by suture at least medially and in having S-shaped median lobe of aedeagus (as in
|
||
<figureCitation id="1307C790A36B7F7BFB1BF9CB8ECCDF4E" box="[1259,1424,1637,1662]" captionStart="FIGURES 115 – 119" captionStartId="40.[151,264,1519,1542]" captionTargetBox="[210,1376,415,1485]" captionTargetId="figure@40.[192,1394,411,1498]" captionTargetPageId="40" captionText="FIGURES 115 – 119. Trichotichnus (Bottchrus) cyanescens Ito, 1998 (specimen from Dhabeli). 115, left metepisternum; 116, apex of last (VII) abdominal sternite, male; 117, 119, median lobe of aedeagus; 118, apical capitulum. 115 – 116, ventral view; 117, lateral view; 118, caudal view; 119, dorsal view. Scale bars = 0.25 mm (A: Figs 115 – 117, 119; B: Fig. 118)." httpUri="https://zenodo.org/record/919668/files/figure.png" pageId="6" pageNumber="307">Figs 117, 119</figureCitation>
|
||
).
|
||
<taxonomicName id="4C3CA096A36B7F7BFF67F9248A86DF93" authorityName="Andrewes" authorityYear="1930" box="[151,474,1674,1699]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFF67F9248A6DDF93" box="[151,305,1674,1699]" italics="true" pageId="6" pageNumber="307">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFEB5F9258A0ADF92" box="[325,342,1675,1698]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFE82F9248A86DF93" box="[370,474,1674,1699]" italics="true" pageId="6" pageNumber="307">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
, in addition to the markedly larger size (
|
||
<quantity id="4CC476F0A36B7F7BFC33F9248F0DDF92" box="[963,1105,1674,1699]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.7" metricValueMin="6.8" pageId="6" pageNumber="307" unit="mm" value="7.25" valueMax="7.7" valueMin="6.8">6.8–7.7 mm</quantity>
|
||
) and sulcate protibia, easily differs from
|
||
<taxonomicName id="4C3CA096A36B7F7BFEDAF9018AA9DFF6" authorityName="Kirschenhofer" authorityYear="1992" box="[298,501,1709,1734]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFEDAF9018A61DFF6" box="[298,317,1711,1734]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFEA1F9018A3EDFF6" box="[337,354,1711,1734]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFE8DF9038AA9DFF6" box="[381,501,1709,1734]" italics="true" pageId="6" pageNumber="307">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
also in metepisternum noticeably longer than wide (
|
||
<figureCitation id="1307C790A36B7F7BFBA8F9038FE1DFF6" box="[1112,1213,1709,1734]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="6" pageNumber="307">Fig. 137</figureCitation>
|
||
), in pronotum with more obtuse basal angles, and in arcuate median lobe of aedeagus bent just behind basal bulb, with thinner apical capitulum and without any spines in the internal sac (
|
||
<figureCitation id="1307C790A36B7F7BFD00F95B88D2DE3E" box="[752,910,1781,1806]" captionStart="FIGURES 135 – 136" captionStartId="46.[151,264,1244,1267]" captionTargetBox="[301,1277,456,1212]" captionTargetId="figure@46.[271,1316,445,1223]" captionTargetPageId="46" captionText="FIGURES 135 – 136. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930, median lobe of aedeagus, lateral view (135, lectotype; 136, holotype of T. curvatus Ito, 1996). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/919684/files/figure.png" pageId="6" pageNumber="307">Figs 135–136</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A36B7F7BFC6AF9588F5CDE3E" box="[922,1024,1782,1806]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="6" pageNumber="307">140, 142</figureCitation>
|
||
). The body size and the shape of the median lobe of
|
||
<taxonomicName id="4C3CA096A36B7F7BFEB5F8B58957DE03" authorityName="Kirschenhofer" authorityYear="1992" box="[325,523,1818,1843]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFEB5F8B58A06DE02" box="[325,346,1819,1842]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFE9AF8B58A27DE02" box="[362,379,1819,1842]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFE63F8B48957DE03" box="[403,523,1818,1843]" italics="true" pageId="6" pageNumber="307">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="4C3CA096A36B7F7BFD25F8B5882DDE02" authorityName="Ito" authorityYear="1998" box="[725,881,1818,1842]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="opacus" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFD25F8B589B6DE02" box="[725,746,1819,1842]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFD0AF8B58857DE02" box="[762,779,1819,1842]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFCD2F8B5882DDE02" box="[802,881,1819,1842]" italics="true" pageId="6" pageNumber="307">opacus</emphasis>
|
||
</taxonomicName>
|
||
described from Darjeeling District,
|
||
<collectingRegion id="49F815F7A36B7F7BFAF7F8B58EC4DE03" box="[1287,1432,1818,1843]" country="India" name="West Bengal" pageId="6" pageNumber="307">West Bengal</collectingRegion>
|
||
, but, based on the illustrations provided in the original description (
|
||
<bibRefCitation id="EFADA6E4A36B7F7BFC43F8908F2FDE66" author="Ito" box="[947,1139,1853,1878]" pageId="6" pageNumber="307" refString="Ito, N. (1998) Some species of the genus Trichotichnus from Nepal and the adjascent regions (Coleoptera, Carabidae, Harpalini). Japanese Journal of Systematic Entomology, 4 (2), 277 - 288." type="journal article" year="1998">
|
||
Ito 1998:
|
||
<figureCitation id="1307C790A36B7F7BFBDBF8938F2FDE66" box="[1067,1139,1853,1878]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="6" pageNumber="307">Fig. 7</figureCitation>
|
||
</bibRefCitation>
|
||
), the median lobe of the aedeagus of the latter species has more transverse apical capitulum, the terminal lamella in dorsal view is markedly dilated at apex and the internal sac is without groups of small spines. The status of
|
||
<taxonomicName id="4C3CA096A36B7F7BFBB7F8298FB9DEAE" authorityName="Ito" authorityYear="1998" box="[1095,1253,1926,1950]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="opacus" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFBB7F8298F00DEAE" box="[1095,1116,1927,1950]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFB9DF8298F22DEAE" box="[1133,1150,1927,1950]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFB66F8298FB9DEAE" box="[1174,1253,1927,1950]" italics="true" pageId="6" pageNumber="307">opacus</emphasis>
|
||
</taxonomicName>
|
||
requires further study because this species is known to us only from the very short original description where there is no direct comparison with
|
||
<taxonomicName id="4C3CA096A36B7F7BFEA9F8618942DED6" authorityName="Kirschenhofer" authorityYear="1992" box="[345,542,1997,2022]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36B7F7BFEA9F8618A31DED6" box="[345,365,1999,2022]" italics="true" pageId="6" pageNumber="307">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36B7F7BFE8EF8618AD3DED6" box="[382,399,1999,2022]" italics="true" pageId="6" pageNumber="307">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36B7F7BFE56F8638942DED6" box="[422,542,1997,2022]" italics="true" pageId="6" pageNumber="307">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA36A7F7AFF37FF3989A9D82D" pageId="7" pageNumber="308" type="distribution">
|
||
<paragraph id="8B83DB15A36A7F7AFF37FF3989A9D82D" blockId="7.[151,1436,151,789]" pageId="7" pageNumber="308">
|
||
<emphasis id="B9480707A36A7F7AFF37FF398A05D980" bold="true" box="[199,345,151,176]" pageId="7" pageNumber="308">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1307C790A36A7F7AFE99FF398A96D980" box="[361,458,151,176]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="7" pageNumber="308">Fig. 151</figureCitation>
|
||
,
|
||
<emphasis id="B9480707A36A7F7AFE27FF368AB9D980" box="[471,485,152,176]" italics="true" pageId="7" pageNumber="308">1</emphasis>
|
||
).
|
||
<taxonomicName id="4C3CA096A36A7F7AFE0DFF398816D980" authorityName="Kirschenhofer" authorityYear="1992" box="[509,842,151,176]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36A7F7AFE0DFF3989CBD980" box="[509,663,151,176]" italics="true" pageId="7" pageNumber="308">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36A7F7AFD58FF3789E5D980" box="[680,697,153,176]" italics="true" pageId="7" pageNumber="308">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36A7F7AFD22FF398816D980" box="[722,842,151,176]" italics="true" pageId="7" pageNumber="308">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
is known from the
|
||
<typeStatus id="548765B7A36A7F7AFBD8FF368F04D980" box="[1064,1112,152,176]" pageId="7" pageNumber="308">type</typeStatus>
|
||
locality in the middle Bothe
|
||
<collectingRegion id="49F815F7A36A7F7AFF67FF128B86D9E5" box="[151,218,188,213]" country="Nepal" name="Kosi" pageId="7" pageNumber="308">Koshi</collectingRegion>
|
||
Valley in
|
||
<collectingRegion id="49F815F7A36A7F7AFEA4FF128AFBD9E5" box="[340,423,188,213]" country="Sri Lanka" name="Central" pageId="7" pageNumber="308">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A36A7F7AFE40FF128AAAD9E5" box="[432,502,188,213]" name="Nepal" pageId="7" pageNumber="308">Nepal</collectingCountry>
|
||
, and from the environments of Sete Village on the eastern slope of Likhu Khola Valley on the western slope of the Lamjura Danda mountain range (southern slope of the Solu Khumbu massif) in the eastern part of
|
||
<collectingRegion id="49F815F7A36A7F7AFE97FEAA8AE6D82D" box="[359,442,260,285]" country="Sri Lanka" name="Central" pageId="7" pageNumber="308">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A36A7F7AFE31FEAA8958D82D" box="[449,516,260,285]" name="Nepal" pageId="7" pageNumber="308">Nepal</collectingCountry>
|
||
(see remarks below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA36A7F7AFF37FE898AEEDA25" pageId="7" pageNumber="308" type="discussion">
|
||
<paragraph id="8B83DB15A36A7F7AFF37FE898A58DBB5" blockId="7.[151,1436,151,789]" pageId="7" pageNumber="308">
|
||
<emphasis id="B9480707A36A7F7AFF37FE898A68D870" bold="true" box="[199,308,295,320]" pageId="7" pageNumber="308">Remarks</emphasis>
|
||
. The species was described based on two males and one female collected in one locality in
|
||
<collectingRegion id="49F815F7A36A7F7AFABAFE898EC0D870" box="[1354,1436,295,320]" country="Sri Lanka" name="Central" pageId="7" pageNumber="308">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A36A7F7AFF67FEE28B82D855" box="[151,222,332,357]" name="Nepal" pageId="7" pageNumber="308">Nepal</collectingCountry>
|
||
: “
|
||
<collectingRegion id="49F815F7A36A7F7AFF05FEE28A05D855" box="[245,345,332,357]" country="Nepal" name="Bagmati" pageId="7" pageNumber="308">Bagmati</collectingRegion>
|
||
, Bhote
|
||
<collectingRegion id="49F815F7A36A7F7AFE43FEE28AAAD855" box="[435,502,332,357]" country="Nepal" name="Kosi" pageId="7" pageNumber="308">Koshi</collectingRegion>
|
||
Khola, Tatopani Malaphu,
|
||
<quantity id="4CC476F0A36A7F7AFCC4FEE288DDD854" box="[820,897,332,356]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="308" unit="m" value="700.0">700 m</quantity>
|
||
”. The specimens from Sete are similar in their morphology to the
|
||
<typeStatus id="548765B7A36A7F7AFE9CFEDE8AC0D8B8" box="[364,412,368,392]" pageId="7" pageNumber="308">type</typeStatus>
|
||
specimens, but were collected quite far from the
|
||
<typeStatus id="548765B7A36A7F7AFC35FEDE88A9D8B8" box="[965,1013,368,392]" pageId="7" pageNumber="308">type</typeStatus>
|
||
locality. The
|
||
<typeStatus id="548765B7A36A7F7AFB7EFEDE8FE2D8B8" box="[1166,1214,368,392]" pageId="7" pageNumber="308">type</typeStatus>
|
||
specimens are very likely mislabelled at least with respect to the altitude data: the village Tatopani in the Bhote
|
||
<collectingRegion id="49F815F7A36A7F7AFB6AFE3A8F81D89D" box="[1178,1245,404,429]" country="Nepal" name="Kosi" pageId="7" pageNumber="308">Koshi</collectingRegion>
|
||
Valley is located at an altitude of approximately
|
||
<quantity id="4CC476F0A36A7F7AFE05FE198911D8FF" box="[501,589,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="308" unit="m" value="1500.0">1500 m</quantity>
|
||
. The Bothe
|
||
<collectingRegion id="49F815F7A36A7F7AFD24FE19884BD8E0" box="[724,791,439,464]" country="Nepal" name="Kosi" pageId="7" pageNumber="308">Koshi</collectingRegion>
|
||
River only reaches an altitude of
|
||
<quantity id="4CC476F0A36A7F7AFB61FE168F8BD8FF" box="[1169,1239,440,464]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="308" unit="m" value="700.0">700 m</quantity>
|
||
more than
|
||
<quantity id="4CC476F0A36A7F7AFAA6FE168EC0D8E0" box="[1366,1436,439,464]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="308" unit="km" value="30.0">30 km</quantity>
|
||
south of Tatopani in the foothills of the Himalaya where this species probably does not occur. The true
|
||
<typeStatus id="548765B7A36A7F7AFAE2FE738E1ED8C5" box="[1298,1346,477,501]" pageId="7" pageNumber="308">type</typeStatus>
|
||
locality is thus very likely located on one of the slopes of the Bhote
|
||
<collectingRegion id="49F815F7A36A7F7AFC7FFE51888EDB28" box="[911,978,511,536]" country="Nepal" name="Kosi" pageId="7" pageNumber="308">Koshi</collectingRegion>
|
||
Valley near Tatopani, above
|
||
<quantity id="4CC476F0A36A7F7AFACAFE518EC4DB27" box="[1338,1432,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="308" unit="m" value="1500.0">1500 m</quantity>
|
||
. Nevertheless, if the
|
||
<typeStatus id="548765B7A36A7F7AFE8DFD8B8AF1DB0D" box="[381,429,549,573]" pageId="7" pageNumber="308">type</typeStatus>
|
||
specimens were really collected in the Bhote
|
||
<collectingRegion id="49F815F7A36A7F7AFC30FD8A8F5FDB0D" box="[960,1027,548,573]" country="Nepal" name="Kosi" pageId="7" pageNumber="308">Koshi</collectingRegion>
|
||
River, the status of the populations from Sete needs further study on the basis of additional material particularly from the area between these two localities.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A36A7F7AFF37FD3E8AEEDA25" blockId="7.[151,1436,151,789]" pageId="7" pageNumber="308">
|
||
It should be noted that
|
||
<bibRefCitation id="EFADA6E4A36A7F7AFE38FD218844DB98" author="Kirschenhofer" box="[456,792,655,680]" pageId="7" pageNumber="308" refString="Kirschenhofer, E. (1992) Beitrag zur Kenntnis der Harpalinen des Himalaya: Harpalus Latreille und Trichotichnus Morawitz (Coleoptera: Carabidae). Koleopterologische Rundschau, 62, 21 - 41." type="journal article" year="1992">
|
||
Kirschenhofer (1992:
|
||
<figureCitation id="1307C790A36A7F7AFD4FFD218853DB98" box="[703,783,655,680]" captionStart="FIGURES 32 – 37" captionStartId="13.[151,264,1070,1093]" captionTargetBox="[231,1359,320,1038]" captionTargetId="figure@13.[195,1392,301,1050]" captionTargetPageId="13" captionText="FIGURES 32 – 37. Trichotichnus (Bottchrus) bubsaensis sp. n. (32 – 33, 35 – 37, holotype; 34, paratype). left metepisternum; 33, apex of last (VII) abdominal sternite, male; 34, same, female; 35, 37, median lobe of aedeagus; 36, apical capitulum. 32 – 34, ventral view; 35, lateral view; 36, caudal view; 37, dorsal view. Scale bars = 0.25 mm (A: Figs 32 – 35; B: Fig. 36)." httpUri="https://zenodo.org/record/919616/files/figure.png" pageId="7" pageNumber="308">Fig. 32</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
probably incorrectly illustrated the aedeagal median lobe of
|
||
<taxonomicName id="4C3CA096A36A7F7AFF45FD1B8A26DBFD" authorityName="Kirschenhofer" authorityYear="1992" box="[181,378,692,717]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A36A7F7AFF45FD1B8B94DBFC" box="[181,200,693,716]" italics="true" pageId="7" pageNumber="308">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A36A7F7AFF29FD1B8BB6DBFC" box="[217,234,693,716]" italics="true" pageId="7" pageNumber="308">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A36A7F7AFEF2FD1A8A26DBFD" box="[258,378,692,717]" italics="true" pageId="7" pageNumber="308">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
as having nearly transverse apical capitulum. Examination of the
|
||
<typeStatus id="548765B7A36A7F7AFBAAFD1B8FD6DBFD" box="[1114,1162,693,717]" pageId="7" pageNumber="308">type</typeStatus>
|
||
specimens revealed that the median lobe of the
|
||
<typeStatus id="548765B7A36A7F7AFE51FD79895FDBC0" box="[417,515,727,752]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
|
||
was broken; the median lobe of the
|
||
<typeStatus id="548765B7A36A7F7AFC58FD768F55DBC0" box="[936,1033,728,752]" pageId="7" pageNumber="308" type="paratype">paratype</typeStatus>
|
||
from the same locality has oblique apical capitulum (
|
||
<figureCitation id="1307C790A36A7F7AFE90FD528AF8DA25" box="[352,420,764,789]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="7" pageNumber="308">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |