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<document id="9EAAD3D9B5FD8B7E467A11237FE34BAD" ID-CLB-Dataset="3942" ID-DOI="10.5281/zenodo.1414213" ID-GBIF-Dataset="2f3fb2f9-a5a8-4ca1-b481-13c0c2e949da" ID-ISSN="0035-418" ID-Zenodo-Dep="1414213" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634784160167" checkinUser="plazi" docAuthor="Yin, Zi-Wei &amp; Cuccodoro, Giulio" docDate="2018" docId="038E8212FFA4E548C188F98CFA63C0F9" docLanguage="en" docName="6_Yin_Cuccodoro.pdf" docOrigin="Revue suisse de Zoologie 125 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Smetanabatrus loebli Yin &amp; Cuccodoro 2018, sp. nov." docType="treatment" docVersion="6" lastPageNumber="247" masterDocId="FFB7FA6AFFA4E54AC243FFEFFFE2C476" masterDocTitle="Notes on Smetanabatrus from Southeast Asia (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="248" masterPageNumber="245" pageNumber="245" updateTime="1699464062487" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="954781438AA8359B14F5EF3475439C78" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A28DD0241538BEAFCE03BDC300A8CF38">
<mods:title id="9386000E27F47DBB87AA247213C7014B">Notes on Smetanabatrus from Southeast Asia (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="5B956D354360EBF635C3ABD2EEFC8E1E" type="personal">
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<mods:roleTerm id="7FDC4963599E199496EBC202425DF309">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D954DB082E56F7AF791A0DC7B3DBDFA">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="61904D66215C3C843BB50894C90F9797">Department of Biology, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="B2D1AFDCA1BD9C763F8315C36516D52D">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="600360C5ED0F4EC43D80B90C570F14AC">Muséum d'histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland. E-mail: gulio. cuccodoro @ ville-ge. ch</mods:affiliation>
</mods:name>
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<mods:title id="26F87721D5D252AD757F853F3091AE31">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="8DC54AE2DEC8EDC0C8A489F845CF955B">
<mods:date id="5F56C7922430ED78A964E64662BAEC6C">2018</mods:date>
<mods:detail id="AF63C3008F4BB377D97152D226D73885" type="pubDate">
<mods:number id="5D53ABFC6D6CAD6E2D37F71BE26316C0">2018-09-28</mods:number>
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<mods:detail id="E36B210F8DE1F21D837FBF35DE905E0F" type="volume">
<mods:number id="CC3978248436B65549E2479858D34748">125</mods:number>
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<mods:classification id="FE81159EAD321D07236B2C6B892EB1AC">journal article</mods:classification>
<mods:identifier id="0A0C1690ED1080DE729E97C3869DD3E0" type="CLB-Dataset">3942</mods:identifier>
<mods:identifier id="F9D07851526CD986AA3E5752221309D3" type="DOI">10.5281/zenodo.1414213</mods:identifier>
<mods:identifier id="0C2100B6053B2069C887CB21DF02CCDC" type="GBIF-Dataset">2f3fb2f9-a5a8-4ca1-b481-13c0c2e949da</mods:identifier>
<mods:identifier id="92D9961FB18DB3CE8EB06AD836A71EA3" type="ISSN">0035-418</mods:identifier>
<mods:identifier id="8FED01C1BE7545A9C165DC09D08AF02A" type="Zenodo-Dep">1414213</mods:identifier>
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<treatment id="038E8212FFA4E548C188F98CFA63C0F9" ID-DOI="http://doi.org/10.5281/zenodo.5590701" ID-GBIF-Taxon="189153342" ID-Zenodo-Dep="5590701" LSID="urn:lsid:plazi:treatment:038E8212FFA4E548C188F98CFA63C0F9" httpUri="http://treatment.plazi.org/id/038E8212FFA4E548C188F98CFA63C0F9" lastPageId="2" lastPageNumber="247" pageId="0" pageNumber="245">
<subSubSection id="C33D608FFFA4E54AC188F98CFB78C2EE" pageId="0" pageNumber="245" type="nomenclature">
<paragraph id="8B983304FFA4E54AC188F98CFAF3C20D" blockId="0.[832,1297,1591,1688]" box="[971,1297,1635,1659]" pageId="0" pageNumber="245">
<emphasis id="B953EF16FFA4E54AC188F98CFAF3C20D" bold="true" box="[971,1297,1635,1659]" pageId="0" pageNumber="245">
<taxonomicName id="4C274887FFA4E54AC188F98CFB57C20D" ID-CoL="4XVLL" authority="Yin &amp; Cuccodoro, 2018" authorityName="Yin &amp; Cuccodoro" authorityYear="2018" box="[971,1205,1635,1659]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="245" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B953EF16FFA4E54AC188F98CFB57C20D" bold="true" box="[971,1205,1635,1659]" italics="true" pageId="0" pageNumber="245">Smetanabatrus loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260526DFFA4E54AC6F8F98BFAF3C20D" box="[1211,1297,1636,1659]" pageId="0" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B983304FFA4E54AC601F96EFB78C2EE" blockId="0.[832,1297,1591,1688]" box="[1090,1178,1665,1688]" pageId="0" pageNumber="245">
<figureCitation id="131C2F81FFA4E54AC601F96EFB78C2EE" box="[1090,1178,1665,1688]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[151,186,1339,1360]" captionStartId-1="2.[831,866,953,974]" captionTargetBox-0="[155,1436,257,1277]" captionTargetBox-1="[831,1436,257,918]" captionTargetId-0="figure-235@1.[151,1436,257,1277]" captionTargetId-1="figure-611@2.[831,1439,257,918]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." captionText-1="Fig. 2. Distribution of Smetanabatrus species. 1 – S. kinabalu, 2 – S. ghecu, 3 – S. loebli sp. nov. Circles are published records, squares represent new data." figureDoi-0="http://doi.org/10.5281/zenodo.5590714" figureDoi-1="http://doi.org/10.5281/zenodo.5590716" httpUri-0="https://zenodo.org/record/5590714/files/figure.png" httpUri-1="https://zenodo.org/record/5590716/files/figure.png" pageId="0" pageNumber="245">Figs 1-2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D608FFFA4E54BC103F95EFE46C2E1" lastPageId="1" lastPageNumber="246" pageId="0" pageNumber="245" type="materials_examined">
<paragraph id="8B983304FFA4E54AC103F95EFC4BC29E" blockId="0.[832,1436,1713,1768]" pageId="0" pageNumber="245">
<materialsCitation id="3B4F3959FFA4E54AC103F95EFC47C29E" ID-GBIF-Occurrence="3391055302" country="Malaysia" location="Genting Highlands" municipality="Peninsular" pageId="0" pageNumber="245" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="B953EF16FFA4E54AC103F95EFC3DC2BE" bold="true" box="[832,991,1713,1736]" pageId="0" pageNumber="245">
<typeStatus id="549C8DA6FFA4E54AC103F95EFC9AC2BE" box="[832,888,1713,1736]" pageId="0" pageNumber="245">Type</typeStatus>
locality:
</emphasis>
<location id="8EF865DFFFA4E54AC1A9F95EFB5EC2BE" LSID="urn:lsid:plazi:treatment:038E8212FFA4E548C188F98CFA63C0F9:8EF865DFFFA4E54AC1A9F95EFB5EC2BE" box="[1002,1212,1713,1736]" country="Malaysia" municipality="Peninsular" name="Genting Highlands" pageId="0" pageNumber="245" stateProvince="Pahang">Genting Highlands</location>
,
<collectingRegion id="49E3FDE6FFA4E54AC689F95EFAFEC2BE" box="[1226,1308,1713,1736]" country="Malaysia" name="Pahang" pageId="0" pageNumber="245">Pahang</collectingRegion>
,
<collectingMunicipality id="6BFCA97EFFA4E54AC768F95EFA7EC2BE" box="[1323,1436,1713,1736]" pageId="0" pageNumber="245">Peninsular</collectingMunicipality>
<collectingCountry id="F3307394FFA4E54AC103F93EFC47C29E" box="[832,933,1745,1768]" name="Malaysia" pageId="0" pageNumber="245">Malaysia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B983304FFA4E54BC103F8EFFE46C2E1" blockId="0.[832,1437,1792,1944]" lastBlockId="1.[151,756,1562,1687]" lastPageId="1" lastPageNumber="246" pageId="0" pageNumber="245">
<emphasis id="B953EF16FFA4E54AC103F8EFFC1DC361" bold="true" box="[832,1023,1792,1815]" pageId="0" pageNumber="245">Type material</emphasis>
(
<specimenCount id="9D21F88DFFA4E54AC669F8EFFB9DC361" box="[1066,1151,1792,1815]" pageId="0" pageNumber="245" type="generic">2 ex.</specimenCount>
):
<materialsCitation id="3B4F3959FFA4E54AC6ECF8EFFB09C3EE" ID-GBIF-Occurrence="3391055305" collectorName="Lobl" country="Malaysia" elevation="950" location="Awana" municipality="Genting Highlands" pageId="0" pageNumber="245" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
<typeStatus id="549C8DA6FFA4E54AC6ECF8EFFAFBC361" box="[1199,1305,1792,1815]" pageId="0" pageNumber="245" type="holotype">Holotype</typeStatus>
, MHNG- ENTO-00013301;
<specimenCount id="9D21F88DFFA4E54AC65BF8CFFBCEC341" box="[1048,1068,1824,1847]" pageId="0" pageNumber="245" type="male">♂</specimenCount>
; W.
<collectingCountry id="F3307394FFA4E54AC63BF8CFFB39C341" box="[1144,1243,1824,1847]" name="Malaysia" pageId="0" pageNumber="245">Malaysia</collectingCountry>
# 28,
<collectingRegion id="49E3FDE6FFA4E54AC705F8CFFA7AC341" box="[1350,1432,1824,1847]" country="Malaysia" name="Pahang" pageId="0" pageNumber="245">Pahang</collectingRegion>
,
<collectingMunicipality id="6BFCA97EFFA4E54AC103F8AFFBF3C321" box="[832,1041,1856,1879]" pageId="0" pageNumber="245">Genting Highlands</collectingMunicipality>
,
<location id="8EF865DFFFA4E54AC65EF8AFFB8BC321" LSID="urn:lsid:plazi:treatment:038E8212FFA4E548C188F98CFA63C0F9:8EF865DFFFA4E54AC65EF8AFFB8BC321" box="[1053,1129,1856,1879]" country="Malaysia" municipality="Genting Highlands" name="Awana" pageId="0" pageNumber="245" stateProvince="Pahang">Awana</location>
,
<quantity id="4CDF9EE1FFA4E54AC634F8AFFB23C321" box="[1143,1217,1856,1879]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="0" pageNumber="245" unit="m" value="950.0">
<elevation id="000AD437FFA4E54AC634F8AFFB23C321" box="[1143,1217,1856,1879]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="0" pageNumber="245" unit="m" value="950.0">950 m</elevation>
</quantity>
; 4.iv.93; coll.
<collectorName id="26D256D2FFA4E54AC721F8AFFA7AC321" box="[1378,1432,1856,1879]" pageId="0" pageNumber="245">Löbl</collectorName>
; veg. debris; “
<typeStatus id="549C8DA6FFA4E54AC18FF88EFBB9C30E" box="[972,1115,1889,1912]" pageId="0" pageNumber="245" type="holotype">HOLOTYPE</typeStatus>
(red, printed),
<taxonomicName id="4C274887FFA4E54AC6BFF88FFB02C3EE" authority="Yin &amp; Cuccodoro 2018" authorityName="Yin &amp; Cuccodoro" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="245" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="B953EF16FFA4E54AC6BFF88FFC9DC3EE" italics="true" pageId="0" pageNumber="245">Smetanabatrus loebli</emphasis>
Yin &amp; Cuccodoro 2018
</taxonomicName>
”
</materialsCitation>
. –
<materialsCitation id="3B4F3959FFA4E54BC777F86EFE42C2E1" ID-GBIF-Occurrence="3391055304" collectorName="Rougemont" country="Malaysia" elevation="1500" lastPageId="1" lastPageNumber="246" location="Fraser's Hill" municipality="Malaya" pageId="0" pageNumber="245" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="549C8DA6FFA4E54AC777F86EFA7AC3EE" box="[1332,1432,1921,1944]" pageId="0" pageNumber="245" type="paratype">Paratype</typeStatus>
: MHNG-ENTO-00013318;
<specimenCount id="9D21F88DFFA5E54BC39BF9F5FE0EC247" box="[472,492,1562,1585]" pageId="1" pageNumber="246" type="female">♀</specimenCount>
;
<collectingCountry id="F3307394FFA5E54BC052F9F5FD9BC247" box="[529,633,1562,1585]" name="Malaysia" pageId="1" pageNumber="246">Malaysia</collectingCountry>
,
<collectingMunicipality id="6BFCA97EFFA5E54BC0D8F9F5FD12C247" box="[667,752,1562,1585]" pageId="1" pageNumber="246">Malaya</collectingMunicipality>
,
<location id="8EF865DFFFA5E54BC2D4F9D3FEC6C225" LSID="urn:lsid:plazi:treatment:038E8212FFA4E548C188F98CFA63C0F9:8EF865DFFFA5E54BC2D4F9D3FEC6C225" box="[151,292,1596,1619]" country="Malaysia" municipality="Malaya" name="Fraser's Hill" pageId="1" pageNumber="246">Fraser’s Hill</location>
,
<quantity id="4CDF9EE1FFA5E54BC377F9D3FE6FC225" box="[308,397,1596,1619]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="246" unit="m" value="1500.0">
<elevation id="000AD437FFA5E54BC377F9D3FE6FC225" box="[308,397,1596,1619]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="246" unit="m" value="1500.0">1500 m</elevation>
</quantity>
; 11.iii.(19)90; coll.
<collectorName id="26D256D2FFA5E54BC02CF9D3FD12C225" box="[623,752,1596,1619]" pageId="1" pageNumber="246">Rougemont</collectorName>
; “
<typeStatus id="549C8DA6FFA5E54BC2E3F9B1FECAC203" box="[160,296,1630,1653]" pageId="1" pageNumber="246" type="paratype">PARATYPE</typeStatus>
(yellow, printed),
<taxonomicName id="4C274887FFA5E54BC044F9B2FE77C2E1" authority="Yin &amp; Cuccodoro 2018" authorityName="Yin &amp; Cuccodoro" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="B953EF16FFA5E54BC044F9B2FD11C203" box="[519,755,1629,1653]" italics="true" pageId="1" pageNumber="246">Smetanabatrus loebli</emphasis>
Yin &amp; Cuccodoro 2018
</taxonomicName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF58638CFFA5E54BC2D4FAD4FB7AC1B5" ID-DOI="http://doi.org/10.5281/zenodo.5590714" ID-Zenodo-Dep="5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="1" pageNumber="246" startId="1.[151,186,1339,1360]" targetBox="[155,1436,257,1277]" targetPageId="1">
<paragraph id="8B983304FFA5E54BC2D4FAD4FB7AC1B5" blockId="1.[151,1437,1338,1475]" pageId="1" pageNumber="246">
Fig. 1.
<taxonomicName id="4C274887FFA5E54BC2A0FAD5FE52C139" authorityName="Yin &amp; Cuccodoro" authorityYear="2018" box="[227,432,1338,1359]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="B953EF16FFA5E54BC2A0FAD5FE52C139" box="[227,432,1338,1359]" italics="true" pageId="1" pageNumber="246">Smetanabatrus loebli</emphasis>
</taxonomicName>
male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection.
</paragraph>
</caption>
<subSubSection id="C33D608FFFA5E54BC2D4F95EFE90CC6B" pageId="1" pageNumber="246" type="diagnosis">
<paragraph id="8B983304FFA5E54BC2D4F95EFE90CC6B" blockId="1.[151,757,1713,2077]" pageId="1" pageNumber="246">
<emphasis id="B953EF16FFA5E54BC2D4F95EFEEDC2BE" bold="true" box="[151,271,1713,1736]" pageId="1" pageNumber="246">Diagnosis:</emphasis>
Body medium-sized, length
<quantity id="4CDF9EE1FFA5E54BC010F95EFD13C2BE" box="[595,753,1713,1736]" metricMagnitude="-3" metricUnit="m" metricValue="2.725" metricValueMax="2.79" metricValueMin="2.66" pageId="1" pageNumber="246" unit="mm" value="2.725" valueMax="2.79" valueMin="2.66">2.66-2.79 mm</quantity>
. Head and pronotum regularly punctate, anterior margin of clypeus angularly projecting anteriorly at middle. Pronotum with thin discal carinae, with distinct median and lateral sulci. Maxillary palpomere IV greatly lobed at inner margin. Distinct ocular canthus present. Males have abdominal tergite 1 (IV), and sternite 3 (V) modified; protrochanter protuberant at ventral margin, metatibia strongly dilated at apical portion; aedeagus with large and transverse basal capsule, median lobe slightly asymmetric.
</paragraph>
</subSubSection>
<subSubSection id="C33D608FFFA5E548C17CF9F5FEA3C1AF" lastPageId="2" lastPageNumber="247" pageId="1" pageNumber="246" type="description">
<paragraph id="8B983304FFA5E548C17CF9F5FF33C0CF" blockId="1.[831,1437,1562,2077]" lastBlockId="2.[151,756,258,1497]" lastPageId="2" lastPageNumber="247" pageId="1" pageNumber="246">
<emphasis id="B953EF16FFA5E54BC17CF9F5FC2FC247" bold="true" box="[831,973,1562,1585]" pageId="1" pageNumber="246">Description:</emphasis>
Male (
<figureCitation id="131C2F81FFA5E54BC660F9F5FB9FC247" box="[1059,1149,1562,1585]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="1" pageNumber="246">Fig. 1A</figureCitation>
). Body length (combined length of head, pronotum, elytra, and abdomen)
<quantity id="4CDF9EE1FFA5E54BC17CF9B4FC4AC204" box="[831,936,1627,1650]" metricMagnitude="-3" metricUnit="m" metricValue="2.66" pageId="1" pageNumber="246" unit="mm" value="2.66">2.66 mm</quantity>
. Body reddish brown, tarsi and mouthparts lighter in color. Head distinctly transverse, length from clypeal anterior margin to head base
<quantity id="4CDF9EE1FFA5E54BC6AAF972FAADC2C2" box="[1257,1359,1693,1716]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="246" unit="mm" value="0.55">0.55 mm</quantity>
, width across eyes
<quantity id="4CDF9EE1FFA5E54BC181F951FBC4C2A3" box="[962,1062,1726,1749]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="246" unit="mm" value="0.71">0.71 mm</quantity>
; surface regularly punctate; vertex convex, foveae far below level of posterior margins of eyes, lacking reverse U-shaped sulcus connecting foveae, thin median carina present from base toward apex of frons, anterior margin of clypeus angularly protuberant (
<figureCitation id="131C2F81FFA5E54BC193F88DFBCAC30F" box="[976,1064,1890,1913]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="1" pageNumber="246">Fig. 1B</figureCitation>
;
<emphasis id="B953EF16FFA5E54BC679F88DFBB7C30F" box="[1082,1109,1890,1913]" italics="true" pageId="1" pageNumber="246">ap</emphasis>
) at middle; eyes moderately prominent, longitudinal length
<quantity id="4CDF9EE1FFA5E54BC6F2F86DFAFDC3EF" box="[1201,1311,1922,1945]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="246" unit="mm" value="0.15">0.15 mm</quantity>
, each eye composed of about 45 facets; with short ocular canthus (
<figureCitation id="131C2F81FFA5E54BC10BF82BFC46C3AD" box="[840,932,1988,2011]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="1" pageNumber="246">Fig. 1C</figureCitation>
;
<emphasis id="B953EF16FFA5E54BC1FAF82BFC30C3AD" box="[953,978,1988,2011]" italics="true" pageId="1" pageNumber="246">oc</emphasis>
); maxillary palpus with palpomere IV greatly lobed on inner margin (
<figureCitation id="131C2F81FFA5E54BC6E6F80AFB1CC38A" box="[1189,1278,2021,2044]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="1" pageNumber="246">Fig. 1C</figureCitation>
;
<emphasis id="B953EF16FFA5E54BC74CF80BFADFC38A" box="[1295,1341,2020,2044]" italics="true" pageId="1" pageNumber="246">mp4</emphasis>
); length of antenna
<quantity id="4CDF9EE1FFA5E54BC18FF7E9FBD8CC6B" box="[972,1082,2054,2077]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="246" unit="mm" value="1.85">1.85 mm</quantity>
; antennomeres each elongate, club indistinct (
<figureCitation id="131C2F81FFA6E548C30AFEEDFE46C56F" box="[329,420,258,281]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1A</figureCitation>
). Pronotum nearly cordiform, about as long as wide, length along midline
<quantity id="4CDF9EE1FFA6E548C0C8FECDFD12C54F" box="[651,752,290,313]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="247" unit="mm" value="0.6">0.60 mm</quantity>
, maximum width
<quantity id="4CDF9EE1FFA6E548C322FEADFE2EC52F" box="[353,460,322,345]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="247" unit="mm" value="0.64">0.64 mm</quantity>
, regularly punctate; with triangular antebasal spines, deep median and thinner lateral sulci, and slightly curved discal carinae on disc; lateral margins rounded on apical half, straight and nearly parallel from middle toward base. Elytra slightly wider than long, length along suture
<quantity id="4CDF9EE1FFA6E548C0C8FE0DFD12C58F" box="[651,752,482,505]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="247" unit="mm" value="0.88">0.88 mm</quantity>
, maximum width 1.00 mm; surface finely punctate; with three large, deep basal foveae, thin and shallow discal sulcus extending just past half elytral length. Legs moderately elongate; protrochanter with large flatten projection (
<figureCitation id="131C2F81FFA6E548C358FD6DFE9BC6EF" box="[283,377,642,665]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1D</figureCitation>
;
<emphasis id="B953EF16FFA6E548C3CFFD6DFE47C6EF" box="[396,421,642,665]" italics="true" pageId="2" pageNumber="247">vp</emphasis>
) on ventral margin, protibia (
<figureCitation id="131C2F81FFA6E548C2DCFD4DFF1AC6CF" box="[159,248,674,697]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1A</figureCitation>
) with lateral margin slightly expanded near middle; metatibia (
<figureCitation id="131C2F81FFA6E548C32AFD2DFE5EC6AF" box="[361,444,706,729]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1E</figureCitation>
) greatly dilated laterally on apical third, expansion broadly concaved on ventral surface. Abdomen much wider than long, length of dorsally visible part along midline
<quantity id="4CDF9EE1FFA6E548C054FCCDFD99C74F" box="[535,635,802,825]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="247" unit="mm" value="0.63">0.63 mm</quantity>
, maximum width
<quantity id="4CDF9EE1FFA6E548C2A1FCADFEA9C72F" box="[226,331,834,857]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="247" unit="mm" value="0.9">0.90 mm</quantity>
; tergite 1 (IV) longest, at basal half with transverse oval patch (
<figureCitation id="131C2F81FFA6E548C386FC8DFDFEC70F" box="[453,540,866,889]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1G</figureCitation>
;
<emphasis id="B953EF16FFA6E548C06AFC8DFDA6C70F" box="[553,580,866,889]" italics="true" pageId="2" pageNumber="247">pa</emphasis>
) slightly raised, maximum diameter of patch
<quantity id="4CDF9EE1FFA6E548C3A2FC6DFDAAC7EF" box="[481,584,898,921]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="247" unit="mm" value="0.14">0.14 mm</quantity>
, area lateral to midline moderately raised (
<figureCitation id="131C2F81FFA6E548C392FC4DFDCAC7CF" box="[465,552,930,953]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1F</figureCitation>
; indicated by red arrows); tergites 2-4 (V-VII) unmodified, with tergite 4 longer than tergites 2 and 3; sternite 3 (V) with reniform (in ventral view) median protuberance (
<figureCitation id="131C2F81FFA6E548C002FBEDFD5DC06F" box="[577,703,1026,1049]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1F, 1H</figureCitation>
;
<emphasis id="B953EF16FFA6E548C089FBEDFD09C06F" box="[714,747,1026,1049]" italics="true" pageId="2" pageNumber="247">mp</emphasis>
) strongly projecting ventrally. Aedeagus (
<figureCitation id="131C2F81FFA6E548C008FBCDFD71C04F" box="[587,659,1058,1081]" captionStart="Fig" captionStartId="1.[151,186,1339,1360]" captionTargetBox="[155,1436,257,1277]" captionTargetId="figure-235@1.[151,1436,257,1277]" captionTargetPageId="1" captionText="Fig. 1. Smetanabatrus loebli male. (A) Dorsal habitus. (B) Head dorsum. (C) Anterolateral view of head, showing form of maxillary palpomere IV and ocular canthus. (D) Protrochanter and profemur. (E) Metatibia. (F) Lateral view of abdomen, showing projections on tergite 1 (IV) and sternite 3 (V). (G) Tergite 1. (H) Median projection of sternite 3, in ventral view. (I) Aedeagus, in ventral view. Scale bars: 1 mm in A; 0.2 mm in B-H, 0.1 mm in I. Abbreviations: ap – apical protuberance, mp – median projection, mp4 – maxillary palpomere IV, oc – ocular canthus, pa – patch, vp – ventral projection." figureDoi="http://doi.org/10.5281/zenodo.5590714" httpUri="https://zenodo.org/record/5590714/files/figure.png" pageId="2" pageNumber="247">Fig. 1I</figureCitation>
) slightly asymmetric, length
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; basal bulb transverse, with large foramen; ventral lobe elongate, broad at base, narrowing at middle, and then roundly expanded toward apex.
</paragraph>
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<paragraph id="8B983304FFA6E548C17CFC56FB05C07E" blockId="2.[831,1436,953,1032]" pageId="2" pageNumber="247">
Fig. 2. Distribution of
<taxonomicName id="4C274887FFA6E548C65CFC56FAF0C7B8" box="[1055,1298,953,974]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B953EF16FFA6E548C65CFC56FB4DC7B8" box="[1055,1199,953,974]" italics="true" pageId="2" pageNumber="247">Smetanabatrus</emphasis>
species. 1
</taxonomicName>
–
<taxonomicName id="4C274887FFA6E548C769FC56FA74C7B8" box="[1322,1430,953,974]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B953EF16FFA6E548C769FC56FAD4C7B8" box="[1322,1334,953,974]" italics="true" pageId="2" pageNumber="247">S</emphasis>
.
<emphasis id="B953EF16FFA6E548C701FC56FA74C7B8" box="[1346,1430,953,974]" italics="true" pageId="2" pageNumber="247">kinabalu</emphasis>
</taxonomicName>
, 2 –
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<emphasis id="B953EF16FFA6E548C1ECFC39FC59C79D" box="[943,955,982,1003]" italics="true" pageId="2" pageNumber="247">S</emphasis>
.
<emphasis id="B953EF16FFA6E548C185FC39FC1DC79D" box="[966,1023,982,1003]" italics="true" pageId="2" pageNumber="247">ghecu</emphasis>
</taxonomicName>
, 3 –
<taxonomicName id="4C274887FFA6E548C66CFC39FB9CC79D" authority="Yin &amp; Cuccodoro, 2018" authorityName="Yin &amp; Cuccodoro" authorityYear="2018" box="[1071,1150,982,1003]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B953EF16FFA6E548C66CFC39FBD9C79D" box="[1071,1083,982,1003]" italics="true" pageId="2" pageNumber="247">S</emphasis>
.
<emphasis id="B953EF16FFA6E548C604FC39FB9CC79D" box="[1095,1150,982,1003]" italics="true" pageId="2" pageNumber="247">loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260526DFFA6E548C6C0FC39FB2CC79D" box="[1155,1230,982,1003]" pageId="2" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
Circles are published records, squares represent new data.
</paragraph>
</caption>
<paragraph id="8B983304FFA6E548C2D4FB2DFEA3C1AF" blockId="2.[151,756,258,1497]" pageId="2" pageNumber="247">
Female. Similar to male in external morphology, except having unmodified legs and abdominal segments, slightly shorter antennae, and smaller eyes each composed of about 25 facets. Measurements (as for male): Body length
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; head length
<quantity id="4CDF9EE1FFA6E548C3CEFAADFE12C12F" box="[397,496,1346,1369]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="247" unit="mm" value="0.55">0.55 mm</quantity>
, width
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; length of eye
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; length of antenna
<quantity id="4CDF9EE1FFA6E548C05CFA8DFD66C10F" box="[543,644,1378,1401]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" pageId="2" pageNumber="247" unit="mm" value="1.59">1.59 mm</quantity>
; length of pronotum
<quantity id="4CDF9EE1FFA6E548C348FA6DFE91C1EF" box="[267,371,1410,1433]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="2" pageNumber="247" unit="mm" value="0.59">0.59 mm</quantity>
, width
<quantity id="4CDF9EE1FFA6E548C38EFA6DFDD7C1EF" box="[461,565,1410,1433]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="247" unit="mm" value="0.66">0.66 mm</quantity>
; length of elytra
<quantity id="4CDF9EE1FFA6E548C2D4FA4DFF18C1CF" box="[151,250,1442,1465]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="247" unit="mm" value="0.8">0.80 mm</quantity>
, width
<quantity id="4CDF9EE1FFA6E548C308FA4DFE4CC1CF" box="[331,430,1442,1465]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="247" unit="mm" value="0.95">0.95 mm</quantity>
; length of abdomen
<quantity id="4CDF9EE1FFA6E548C0CEFA4DFD12C1CF" box="[653,752,1442,1465]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="247" unit="mm" value="0.85">0.85 mm</quantity>
, width
<quantity id="4CDF9EE1FFA6E548C298FA2DFEDFC1AF" box="[219,317,1474,1497]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="247" unit="mm" value="0.89">0.89 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D608FFFA6E548C2D4FA1EFDE3C35E" pageId="2" pageNumber="247" type="discussion">
<paragraph id="8B983304FFA6E548C2D4FA1EFDE3C35E" blockId="2.[151,756,1521,1832]" pageId="2" pageNumber="247">
<emphasis id="B953EF16FFA6E548C2D4FA1EFEFEC27E" bold="true" box="[151,284,1521,1544]" pageId="2" pageNumber="247">Comments:</emphasis>
Several shared features,
<emphasis id="B953EF16FFA6E548C002FA1EFD80C27E" box="[577,610,1521,1544]" italics="true" pageId="2" pageNumber="247">i.e.</emphasis>
, the slightly expanded outer margin of the protibia, modified male tergite 1 (IV), and strongly dilated male metatibia suggest that the new species may be more closely related to
<taxonomicName id="4C274887FFA6E548C34CF99EFE6FC2FE" box="[271,397,1648,1672]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="B953EF16FFA6E548C34CF99EFEFEC2FE" box="[271,284,1649,1672]" italics="true" pageId="2" pageNumber="247">S</emphasis>
.
<emphasis id="B953EF16FFA6E548C36CF99FFE6FC2FE" box="[303,397,1648,1672]" italics="true" pageId="2" pageNumber="247">kinabalu</emphasis>
</taxonomicName>
from Borneo than to
<taxonomicName id="4C274887FFA6E548C0D0F99EFD16C2FE" box="[659,756,1648,1672]" class="Insecta" family="Staphylinidae" genus="Smetanabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="ghecu">
<emphasis id="B953EF16FFA6E548C0D0F99EFD42C2FE" box="[659,672,1649,1672]" italics="true" pageId="2" pageNumber="247">S</emphasis>
.
<emphasis id="B953EF16FFA6E548C0F7F99FFD16C2FE" box="[692,756,1648,1672]" italics="true" pageId="2" pageNumber="247">ghecu</emphasis>
</taxonomicName>
from
<collectingCountry id="F3307394FFA6E548C29AF97EFEA6C2DE" box="[217,324,1681,1704]" name="Myanmar" pageId="2" pageNumber="247">Myanmar</collectingCountry>
. Males of the latter species have the protibia slightly broadened on inner margin, and have unmodified tergite 1 (IV) and metatibia. Separation of the males of these three species may be accomplished by using the key presented below.
</paragraph>
</subSubSection>
<subSubSection id="C33D608FFFA6E548C2D4F8AFFDA6C3C1" pageId="2" pageNumber="247" type="biology_ecology">
<paragraph id="8B983304FFA6E548C2D4F8AFFDA6C3C1" blockId="2.[151,756,1856,1975]" pageId="2" pageNumber="247">
<emphasis id="B953EF16FFA6E548C2D4F8AFFF14C321" bold="true" box="[151,246,1856,1879]" pageId="2" pageNumber="247">Biology:</emphasis>
According to unpublished MHNG field notes, #28 on the label of the
<typeStatus id="549C8DA6FFA6E548C383F88FFDFCC301" box="[448,542,1888,1911]" pageId="2" pageNumber="247" type="holotype">holotype</typeStatus>
indicates that the individual was sifted from an accumulation of branches and bamboo leaves in a rather dry forest
</paragraph>
</subSubSection>
<subSubSection id="C33D608FFFA6E548C2D4F820FA63C0F9" pageId="2" pageNumber="247" type="distribution">
<paragraph id="8B983304FFA6E548C2D4F820FD6BC390" blockId="2.[151,649,1999,2022]" box="[151,649,1999,2022]" pageId="2" pageNumber="247">
<heading id="D0D08468FFA6E548C2D4F820FD6BC390" box="[151,649,1999,2022]" fontSize="10" level="3" pageId="2" pageNumber="247" reason="3">
<emphasis id="B953EF16FFA6E548C2D4F820FECEC390" bold="true" box="[151,300,1999,2022]" pageId="2" pageNumber="247">Distribution:</emphasis>
West
<collectingCountry id="F3307394FFA6E548C32EF820FE2DC390" box="[365,463,1999,2022]" name="Malaysia" pageId="2" pageNumber="247">Malaysia</collectingCountry>
(
<collectingRegion id="49E3FDE6FFA6E548C39EF820FDD2C390" box="[477,560,1999,2022]" country="Malaysia" name="Pahang" pageId="2" pageNumber="247">Pahang</collectingRegion>
;
<figureCitation id="131C2F81FFA6E548C078F820FD9EC390" box="[571,636,1999,2022]" captionStart="Fig" captionStartId="2.[831,866,953,974]" captionTargetBox="[831,1436,257,918]" captionTargetId="figure-611@2.[831,1439,257,918]" captionTargetPageId="2" captionText="Fig. 2. Distribution of Smetanabatrus species. 1 – S. kinabalu, 2 – S. ghecu, 3 – S. loebli sp. nov. Circles are published records, squares represent new data." figureDoi="http://doi.org/10.5281/zenodo.5590716" httpUri="https://zenodo.org/record/5590716/files/figure.png" pageId="2" pageNumber="247">Fig. 2</figureCitation>
).
</heading>
</paragraph>
<paragraph id="8B983304FFA6E548C17CFBB7FA63C0F9" blockId="2.[831,1436,1112,1167]" pageId="2" pageNumber="247">
<emphasis id="B953EF16FFA6E548C17CFBB7FC26C019" bold="true" box="[831,964,1112,1135]" pageId="2" pageNumber="247">Etymology:</emphasis>
The new species is named after Ivan Löbl (
<collectingRegion id="49E3FDE6FFA6E548C104FB97FC7FC0F9" box="[839,925,1144,1167]" country="Switzerland" name="Geneve" pageId="2" pageNumber="247">Geneva</collectingRegion>
,
<collectingCountry id="F3307394FFA6E548C1E4FB97FBC9C0F9" box="[935,1067,1144,1167]" name="Switzerland" pageId="2" pageNumber="247">Switzerland</collectingCountry>
), who collected the single male.
</paragraph>
</subSubSection>
</treatment>
</document>
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