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<document id="AFD30050B559A0C4B48BD110B4987506" ID-DOI="10.11646/zootaxa.3669.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10098002" ID-ZooBank="35D1C69A-0E41-430A-8483-BEB84F45D8D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629862386453" checkinUser="felipe" docAuthor="Wang, Shuxia, Du, Zhaohui &amp; Li, Houhun" docDate="2013" docId="D957879E0A21D65BA099D323FD1721BF" docLanguage="en" docName="zt03669p455.pdf" docOrigin="Zootaxa 3669 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Promalactis fortijuxtalis Wang, Du &amp; Li 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="418" masterDocId="256EFFE60A31D64AA00ED747FFE12350" masterDocTitle="Taxonomic study of the genus Promalactis Meyrick, 1908 from Southeast Asia (Lepidoptera: Oecophoridae)" masterLastPageNumber="455" masterPageNumber="401" pageNumber="417" updateTime="1699589443423" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4DA36F823ABD3E27B6076ABADEA4AC26">Taxonomic study of the genus Promalactis Meyrick, 1908 from Southeast Asia (Lepidoptera: Oecophoridae)</mods:title>
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<mods:name id="5EBD64E9CEFFD862A3F3A22DD2E11932" type="personal">
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<mods:namePart id="BF3CF7158C13141E1CD25776A0FEF3B1">Wang, Shuxia</mods:namePart>
<mods:affiliation id="C2F1E7C474D2E03C4C3C71AF20C45063">These authors contributed equally to this work.</mods:affiliation>
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<mods:namePart id="6857623A52AEC6FD2A2A546C4AA548AE">Du, Zhaohui</mods:namePart>
<mods:affiliation id="476ABD77EF6379381500C9971F7861EF">These authors contributed equally to this work.</mods:affiliation>
</mods:name>
<mods:name id="34BE421FCCE903BCFB5C82120BB1DE1E" type="personal">
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<mods:namePart id="459B68260653ED12E85FB36D90F08346">Li, Houhun</mods:namePart>
<mods:nameIdentifier id="F63856E364EDD6BE80383048F085AB81" type="email">lihouhun@nankai.edu.cn</mods:nameIdentifier>
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<mods:title id="1E746FEED45C652DD705522BD4DB4955">Zootaxa</mods:title>
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<mods:part id="2A808F9B5D0182CD91C0495C0398FCCD">
<mods:date id="C074D153574AED8430283BB050DB3F12">2013</mods:date>
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<mods:number id="FB7B511C7CD4A1B2055A80C3384E423B">2013-06-10</mods:number>
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<mods:number id="171FF573C695789AF1D3EEDF1931402B">3669</mods:number>
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<mods:identifier id="5D63F36CE8B004843B234CA585646938" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="19E465030A21D65AA099D323FD11272D" box="[151,752,1123,1150]" pageId="16" pageNumber="417" type="nomenclature">
<paragraph id="514136880A21D65AA099D323FD11272D" blockId="16.[151,752,1123,1182]" box="[151,752,1123,1150]" pageId="16" pageNumber="417">
<heading id="0A0981E40A21D65AA099D323FD11272D" bold="true" box="[151,752,1123,1150]" fontSize="11" level="1" pageId="16" pageNumber="417" reason="1">
<emphasis id="638AEA9A0A21D65AA099D323FD11272D" bold="true" box="[151,752,1123,1150]" pageId="16" pageNumber="417">
<taxonomicName id="96FE4D0B0A21D65AA099D323FD64272D" authority="Wang, Du &amp; Li" authorityName="Wang, Du &amp; Li" authorityYear="2013" box="[151,645,1123,1150]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="fortijuxtalis" status="sp. nov.">
<emphasis id="638AEA9A0A21D65AA099D323FE5D272E" bold="true" box="[151,444,1124,1150]" italics="true" pageId="16" pageNumber="417">Promalactis fortijuxtalis</emphasis>
Wang, Du &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="78B957E10A21D65AA29CD324FD11272D" box="[658,752,1123,1149]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="19E465030A21D65AA099D3C1FEC427CE" box="[151,293,1158,1182]" pageId="16" pageNumber="417" type="description">
<paragraph id="514136880A21D65AA099D3C1FEC427CE" blockId="16.[151,752,1123,1182]" box="[151,293,1158,1182]" pageId="16" pageNumber="417">
(
<figureCitation id="C9C52A0D0A21D65AA091D3C1FF1527CE" box="[159,244,1158,1182]" captionStart="FIGURES 916" captionStartId="14.[151,263,1904,1927]" captionTargetBox="[218,1375,185,1886]" captionTargetId="figure-93@14.[218,1375,185,1886]" captionTargetPageId="14" captionText="FIGURES 916. Adults of Promalactis spp. 9, P.flagellaris sp. nov., holotype; 10, P.foraminulata sp. nov., holotype; 11, P. fortijuxtalis sp. nov., paratype; 12, P.lancea sp. nov., paratype; 13, P.latuncata sp. nov., holotype; 14, P. longisaccata sp. nov., holotype; 15, P. longisaccata sp. nov., paratype, showing the variation of dorsal spot; 16, P. lophacantha sp. nov., holotype (9 11, 1316: ♂, 12: ♀)." figureDoi="http://doi.org/10.5281/zenodo.10098006" httpUri="https://zenodo.org/record/10098006/files/figure.png" pageId="16" pageNumber="417">Figs 11</figureCitation>
,
<figureCitation id="C9C52A0D0A21D65AA10ED3C1FEFC27CE" box="[256,285,1158,1182]" captionStart="FIGURES 4146" captionStartId="38.[151,263,1874,1897]" captionTargetBox="[255,1341,183,1850]" captionTargetId="figure-109@38.[206,1381,151,1879]" captionTargetPageId="38" captionText="FIGURES 4146. Male genitalia of Promalactis spp. 41, P. extumida sp. nov., holotype, slide No. ZMUC-NK004; 42, P. fascisetacea sp. nov., holotype, 42a, enlarged papillary processes of uncus, slide No. MNHU-NK015; 43, P.flagellaris sp. nov., holotype, slide No. ZMUC-NK017; 44, P. foraminulata sp. nov., holotype, 44a, enlarged basal process of costa and left lateral process of juxta, slide No. BMNH-33560; 45, P. fortijuxtalis sp. nov., holotype, slide No. ZMUC-NK013; 46, P.lancea sp. nov., holotype, slide No. ZMUC-NK011." figureDoi="http://doi.org/10.5281/zenodo.10098017" httpUri="https://zenodo.org/record/10098017/files/figure.png" pageId="16" pageNumber="417">45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="19E465030A21D65AA099D388FE1E267E" pageId="16" pageNumber="417" type="materials_examined">
<paragraph id="514136880A21D65AA099D388FE1E267E" blockId="16.[151,1436,1230,2011]" pageId="16" pageNumber="417">
<emphasis id="638AEA9A0A21D65AA099D388FEA827B7" bold="true" box="[151,329,1230,1255]" pageId="16" pageNumber="417">Type material:</emphasis>
<materialsCitation id="E1963CD50A21D65AA140D389FB10265A" collectingDate="1984-09-29" collectingDateMax="1984-10-06" collectingDateMin="1984-09-29" collectionCode="ZMUC" collectorName="Karsholt &amp; Lomholdt &amp; Nielsen &amp; No." country="Thailand" elevation="700" latitude="12.816667" location="Khao Yai Nat. Park" longLatPrecision="1292" longitude="99.933334" pageId="16" pageNumber="417" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Nayok" typeStatus="holotype">
<typeStatus id="8E45882A0A21D65AA140D389FE5727B6" box="[334,438,1230,1254]" pageId="16" pageNumber="417" type="holotype">Holotype</typeStatus>
<specimenCount id="47F8FD010A21D65AA1B0D388FE3227B6" box="[446,467,1231,1254]" pageId="16" pageNumber="417" type="male"></specimenCount>
,
<emphasis id="638AEA9A0A21D65AA1EFD389FD9927B7" bold="true" box="[481,632,1230,1255]" pageId="16" pageNumber="417">
<collectingCountry id="29E976180A21D65AA1EFD389FD9927B7" box="[481,632,1230,1255]" name="Thailand" pageId="16" pageNumber="417">THAILAND</collectingCountry>
</emphasis>
,
<location id="542160530A21D65AA288D389FC8127B6" LSID="urn:lsid:plazi:treatment:D957879E0A21D65BA099D323FD1721BF:542160530A21D65AA288D389FC8127B6" box="[646,864,1230,1254]" country="Thailand" latitude="12.816667" longLatPrecision="1292" longitude="99.933334" name="Khao Yai Nat. Park" pageId="16" pageNumber="417" stateProvince="Nakhon Nayok">Khao Yai Nat. Park</location>
(
<geoCoordinate id="34CA504F0A21D65AA37ED389FC3027B6" box="[880,977,1230,1255]" degrees="12" direction="north" minutes="49" orientation="latitude" pageId="16" pageNumber="417" precision="925" value="12.816667">12°49'N</geoCoordinate>
,
<geoCoordinate id="34CA504F0A21D65AA3D3D389FBDB27B6" box="[989,1082,1230,1255]" degrees="99" direction="east" minutes="56" orientation="longitude" pageId="16" pageNumber="417" precision="925" value="99.933334">99°56'E</geoCoordinate>
),
<collectingRegion id="933AF86A0A21D65AA440D389FA8727B6" box="[1102,1382,1230,1254]" country="Thailand" name="Nakhon Nayok" pageId="16" pageNumber="417">Nakhon Nayok Province</collectingRegion>
,
<quantity id="96069B6D0A21D65AA57CD389FF4E265A" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="417" unit="m" value="700.0">
<elevation id="DAD3D1BB0A21D65AA57CD389FF4E265A" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="417" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="254010480A21D65AA0B4D3B5FE89265A" box="[186,360,1266,1290]" pageId="16" pageNumber="417" value="1984-09-29" valueMax="1984-10-06" valueMin="1984-09-29">
<collectingDate id="3504E9A00A21D65AA0B4D3B5FE89265A" box="[186,360,1266,1290]" pageId="16" pageNumber="417" value="1984-09-29" valueMax="1984-10-06" valueMin="1984-09-29">29.ix6.x.1984</collectingDate>
</date>
, leg.
<collectorName id="FC0B535E0A21D65AA1ACD3B5FDE4265A" box="[418,517,1266,1290]" pageId="16" pageNumber="417">Karsholt</collectorName>
,
<collectorName id="FC0B535E0A21D65AA21ED3B5FD9F265A" box="[528,638,1266,1290]" pageId="16" pageNumber="417">Lomholdt</collectorName>
&amp;
<collectorName id="FC0B535E0A21D65AA2AFD3B5FD18265A" box="[673,761,1266,1290]" pageId="16" pageNumber="417">Nielsen</collectorName>
, genitalia slide
<collectorName id="FC0B535E0A21D65AA3A9D3B4FC30265A" box="[935,977,1267,1290]" pageId="16" pageNumber="417">No.</collectorName>
ZMUC-NK013 (
<collectionCode id="37EFAE4D0A21D65AA49CD3B5FB08265A" box="[1170,1257,1266,1290]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="16" pageNumber="417">ZMUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E1963CD50A21D65AA4F5D3B4FE1A267E" collectingDate="1984-09-29" collectingDateMax="1984-10-06" collectingDateMin="1984-09-29" collectionCode="ZMUC" collectorName="Karsholt &amp; Lomholdt &amp; Nielsen &amp; No." country="Thailand" elevation="700" latitude="12.816667" location="Khao Yai Nat. Park" longLatPrecision="1292" longitude="99.933334" pageId="16" pageNumber="417" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Nayok" typeStatus="paratype">
<typeStatus id="8E45882A0A21D65AA4F5D3B4FA80265B" box="[1275,1377,1267,1291]" pageId="16" pageNumber="417" type="paratype">Paratype</typeStatus>
:
<specimenCount id="47F8FD010A21D65AA565D3B5FA74265A" box="[1387,1429,1266,1290]" pageId="16" pageNumber="417" type="male">1 ♂</specimenCount>
, same data as holotype (
<collectionCode id="37EFAE4D0A21D65AA192D251FE12267E" box="[412,499,1302,1326]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="16" pageNumber="417">ZMUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19E465030A21D65BA0C9D27DFE7122A4" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417" type="description">
<paragraph id="514136880A21D65AA0C9D27DFE7F243E" blockId="16.[151,1436,1230,2011]" pageId="16" pageNumber="417">
<emphasis id="638AEA9A0A21D65AA0C9D27DFEB92603" bold="true" box="[199,344,1338,1363]" inLineHeading="true" pageId="16" pageNumber="417" reason="1">Description.</emphasis>
Imago (
<figureCitation id="C9C52A0D0A21D65AA1BBD27DFDE52602" box="[437,516,1338,1362]" captionStart="FIGURES 916" captionStartId="14.[151,263,1904,1927]" captionTargetBox="[218,1375,185,1886]" captionTargetId="figure-93@14.[218,1375,185,1886]" captionTargetPageId="14" captionText="FIGURES 916. Adults of Promalactis spp. 9, P.flagellaris sp. nov., holotype; 10, P.foraminulata sp. nov., holotype; 11, P. fortijuxtalis sp. nov., paratype; 12, P.lancea sp. nov., paratype; 13, P.latuncata sp. nov., holotype; 14, P. longisaccata sp. nov., holotype; 15, P. longisaccata sp. nov., paratype, showing the variation of dorsal spot; 16, P. lophacantha sp. nov., holotype (9 11, 1316: ♂, 12: ♀)." figureDoi="http://doi.org/10.5281/zenodo.10098006" httpUri="https://zenodo.org/record/10098006/files/figure.png" pageId="16" pageNumber="417">Fig. 11</figureCitation>
) with wingspan 11.512.0 mm. Head with vertex shining white, frons and occiput dark brown. Labial palpus with basal and second segments yellowish brown on inner surface, dark brown mixed with black on outer surface; third segment black, about same length as second. Antenna with scape white; flagellum with basal several flagellomeres white, remaining ones white and black on dorsal surface, dark brown on ventral surface. Thorax and tegula dark ochreous brown. Forewing dark ochreous yellow; markings white edged with black scales: costal margin with a narrow triangular patch at 3/5, obliquely extending down to midwing; three white streaks arising from dorsal margin: first one from dorsal 1/5 to base of fold, second one from dorsal 2/5 to basal 1/3 of upper margin of cell, broad and slightly arched, third one from dorsal 2/3 to lower margin of cell at distal 1/5; tornus with a small triangular spot; an ill-defined dark-gray streak along distal 1/5 of lower margin of cell; apex with a large wedge-like spot extending along termen, mixed with some black scales; cilia dark ochreous yellow, dark gray along distal part of costal and dorsal margins. Hindwing and cilia gray. Foreleg black, tibia with white spot near base and middle on dorsal surface, with a tuft of short white scales at apex, tarsus with white spots on dorsal surface; midleg with tibia and tarsus yellow on ventral surface, black on dorsal surface, tibia white in basal 1/3, and with a few white scales at middle on dorsal surface, with a tuft of long white scales at apex, tarsus with white spots on dorsal surface; hindleg yellow on ventral surface, black on dorsal surface, tarsus with white spots on dorsal surface.
</paragraph>
<paragraph id="514136880A21D65BA0C9D03DFCEA229F" blockId="16.[151,1436,1230,2011]" lastBlockId="17.[151,1436,151,752]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
Male genitalia (
<figureCitation id="C9C52A0D0A21D65AA173D03DFE3524C2" box="[381,468,1914,1938]" captionStart="FIGURES 4146" captionStartId="38.[151,263,1874,1897]" captionTargetBox="[255,1341,183,1850]" captionTargetId="figure-109@38.[206,1381,151,1879]" captionTargetPageId="38" captionText="FIGURES 4146. Male genitalia of Promalactis spp. 41, P. extumida sp. nov., holotype, slide No. ZMUC-NK004; 42, P. fascisetacea sp. nov., holotype, 42a, enlarged papillary processes of uncus, slide No. MNHU-NK015; 43, P.flagellaris sp. nov., holotype, slide No. ZMUC-NK017; 44, P. foraminulata sp. nov., holotype, 44a, enlarged basal process of costa and left lateral process of juxta, slide No. BMNH-33560; 45, P. fortijuxtalis sp. nov., holotype, slide No. ZMUC-NK013; 46, P.lancea sp. nov., holotype, slide No. ZMUC-NK011." figureDoi="http://doi.org/10.5281/zenodo.10098017" httpUri="https://zenodo.org/record/10098017/files/figure.png" pageId="16" pageNumber="417">Fig. 45</figureCitation>
). Uncus small, nearly rounded, with a small U-shaped notch at middle on posterior margin. Gnathos about 3/5 length of uncus, subtriangular, curved backward; lateral arms narrow, about same length as gnathos. Tegumen branched from middle, rounded anteriorly. Valva very broad and short; costa very short, slightly concave; ventral margin with basal 3/4 convex, distal 1/4 concave; apex deeply concave, forming two apical processes: dorso-apical process short, triangular, basal 2/3 with a triangular fold setose apically; ventroapical process elongate, basally slender, distally broad elliptic and setose, serrate dorsally. Sacculus very broad, subtriangular, reaching basal 3/4 of ventral margin of valva. Saccus very broad, slightly longer than uncus, nearly semi-ovate, bluntly rounded anteriorly. Juxta very strong, sclerotized, narrow at base, with a heavily sclerotized, cymbiform basal process reaching apex of saccus, gradually broadened to 1/5, posterior 4/5 broadly U shaped, margined with dense strong spines dorsally, forming two knife-like lateral lobes, sinuate on posterior margin, setose, reaching base of uncus. Aedeagus curved, slightly shorter than juxta, distally sclerotized, narrowed triangularly, margined with blunt teeth; cornutus absent.
</paragraph>
<paragraph id="514136880A20D65BA0C9D69BFE7122A4" blockId="17.[151,1436,151,752]" box="[199,400,476,500]" pageId="17" pageNumber="418">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="19E465030A20D65BA0C9D6B8FB9921D4" pageId="17" pageNumber="418" type="diagnosis">
<paragraph id="514136880A20D65BA0C9D6B8FB9921D4" blockId="17.[151,1436,151,752]" pageId="17" pageNumber="418">
<emphasis id="638AEA9A0A20D65BA0C9D6B8FEA02148" bold="true" box="[199,321,511,536]" inLineHeading="true" pageId="17" pageNumber="418" reason="1">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="96FE4D0B0A20D65BA2A1D546FC2F2147" authority="Wang, 2006" authorityName="Wang" authorityYear="2006" box="[687,974,511,536]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="spatulata">
<emphasis id="638AEA9A0A20D65BA2A1D546FCD42148" box="[687,821,512,536]" italics="true" pageId="17" pageNumber="418">P. spatulata</emphasis>
<bibRefCitation id="356F4B790A20D65BA34ED547FC2F2147" author="Wang, S. X." box="[832,974,511,536]" pageId="17" pageNumber="418" refId="ref47248" refString="Wang, S. X. (2006) Oecophoridae of China (Insecta: Lepidoptera). Science Press, Beijing, viii + 258 pp. + 15 color plates." type="book" year="2006">Wang, 2006</bibRefCitation>
</taxonomicName>
. It can be separated by the dorso-apical process of the valva with a triangular fold in basal 2/3, ventro-apical process broad and elliptic distally and the very strong juxta broadly U shaped in the male genitalia. In
<taxonomicName id="96FE4D0B0A20D65BA306D50EFC6D2130" authorityName="Wang" authorityYear="2006" box="[776,908,584,608]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="spatulata">
<emphasis id="638AEA9A0A20D65BA306D50EFC6D2130" box="[776,908,584,608]" italics="true" pageId="17" pageNumber="418">P. spatulata</emphasis>
</taxonomicName>
, dorso-apical process of the valva has no fold, the ventro-apical process is large thorn-like and the juxta is slender in the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="19E465030A20D65BA0C9D5C8FD7721F7" box="[199,662,655,680]" pageId="17" pageNumber="418" type="distribution">
<paragraph id="514136880A20D65BA0C9D5C8FD7721F7" blockId="17.[151,1436,151,752]" box="[199,662,655,680]" pageId="17" pageNumber="418">
<emphasis id="638AEA9A0A20D65BA0C9D5C8FE8121F8" bold="true" box="[199,352,655,680]" inLineHeading="true" pageId="17" pageNumber="418" reason="1">Distribution.</emphasis>
<collectingCountry id="29E976180A20D65BA169D5C8FE2B21F7" box="[359,458,655,679]" name="Thailand" pageId="17" pageNumber="418">Thailand</collectingCountry>
(
<collectingRegion id="933AF86A0A20D65BA1D7D5C8FD6B21F7" box="[473,650,655,679]" country="Thailand" name="Nakhon Nayok" pageId="17" pageNumber="418">Nakhon Nayok</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="19E465030A20D65BA0C9D5F3FD1721BF" pageId="17" pageNumber="418" type="etymology">
<paragraph id="514136880A20D65BA0C9D5F3FD1721BF" blockId="17.[151,1436,151,752]" pageId="17" pageNumber="418">
<emphasis id="638AEA9A0A20D65BA0C9D5F3FEAC219C" bold="true" box="[199,333,692,716]" inLineHeading="true" pageId="17" pageNumber="418" reason="1">Etymology.</emphasis>
The specific name is derived from the Latin prefix
<emphasis id="638AEA9A0A20D65BA39ED5F3FC23219C" box="[912,962,692,716]" italics="true" pageId="17" pageNumber="418">fort-</emphasis>
(= strong) and the Latin
<emphasis id="638AEA9A0A20D65BA4D4D5F3FACC219C" box="[1242,1325,692,716]" italics="true" pageId="17" pageNumber="418">juxtalis</emphasis>
(= juxta), referring to the very strong juxta in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>