176 lines
17 KiB
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176 lines
17 KiB
XML
<document id="58096E2AB06ACFEF4122D07475925665" ID-DOI="10.11646/zootaxa.3999.2.3" ID-GBIF-Dataset="8f47b303-98a7-43ce-b5bb-d1b4bf02555b" ID-ISSN="1175-5326" ID-Zenodo-Dep="237045" ID-ZooBank="E4B0B754-1C94-4B61-B612-848804B059A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461280765185" checkinUser="plazi" docAuthor="Mcmahan, Caleb D., Matamoros, Wilfredo A., Piller, Kyle R. & Chakrabarty, Prosanta" docDate="2015" docId="36162773FFB7FFB6AB8EFF16FF6C0716" docLanguage="en" docName="zt03999p234.pdf" docOrigin="Zootaxa 3999 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichromis Mcmahan, Matamoros, Piller & Chakrabarty, 2015, gen. nov." docType="treatment" docUuid="BEC5E416-152F-42DD-835E-D17CFDAE2038" docUuidSource="ZooBank" docVersion="9" lastPageNumber="231" masterDocId="CA2F5F0BFFA3FFA2AB19FF8EFF850318" masterDocTitle="Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera" masterLastPageNumber="234" masterPageNumber="211" pageNumber="231" updateTime="1698613983299" updateUser="plazi">
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<mods:titleInfo id="DE1BBA14D7A931BD33CCF1E2CBAC5A3A">
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<mods:title id="84F76C7D28A0CA1127FCAC5546AD2DE5">Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera</mods:title>
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<mods:roleTerm id="C6D1B1A86AEB39835943FC3EA43E4898">Author</mods:roleTerm>
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<mods:namePart id="D7C9EAA4CDFFE1A76F11C4F9B7CE6537">Mcmahan, Caleb D.</mods:namePart>
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<mods:namePart id="B0865EF427AB75AE884EC97802A03ABD">Matamoros, Wilfredo A.</mods:namePart>
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<mods:namePart id="7A003AE93E54A990F56544E12CAB399C">Piller, Kyle R.</mods:namePart>
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<mods:name id="082DC0884DEC6589306CC8D27D6BF854" type="personal">
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<mods:roleTerm id="6D9B755C231C20A13AC91B1A72C44B65">Author</mods:roleTerm>
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<mods:namePart id="F98AFD9A91A5BA5441920CDA182DF95F">Chakrabarty, Prosanta</mods:namePart>
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<mods:typeOfResource id="199CB5CA0106F7DB6E8ECB75A636360A">text</mods:typeOfResource>
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<mods:title id="65195F98D37CD255106A09A478FC3AD0">Zootaxa</mods:title>
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<mods:part id="07384E7B3467E170BACC15A534745563">
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<mods:date id="31ACB33A6945FE802855EBEB98F85CDB">2015</mods:date>
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<mods:detail id="8789A2F77402913ED7F8B775FDF34536" type="volume">
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<mods:number id="D67417D3C8239DF3599E271E3E5CD5C2">3999</mods:number>
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<mods:detail id="8D908A0326C6A0B44A6F87CE0FF4624E" type="issue">
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<mods:number id="5389B808A67DF8396CB0762E51080108">2</mods:number>
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<mods:classification id="524C82208D9168A3C0B7F44B59F9CEBF">journal article</mods:classification>
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<mods:identifier id="F79793D6C65932DD26FDAE610499C8BC" type="DOI">10.11646/zootaxa.3999.2.3</mods:identifier>
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<treatment id="36162773FFB7FFB6AB8EFF16FF6C0716" ID-DOI="http://doi.org/10.5281/zenodo.5672278" ID-GBIF-Taxon="119640595" ID-Zenodo-Dep="5672278" LSID="urn:lsid:zoobank.org:act:BEC5E416-152F-42DD-835E-D17CFDAE2038" httpUri="http://treatment.plazi.org/id/36162773FFB7FFB6AB8EFF16FF6C0716" lastPageNumber="231" pageId="20" pageNumber="231">
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<subSubSection id="F6A5C5EEFFB7FFB6AB8EFF16FD300201" pageId="20" pageNumber="231" type="nomenclature">
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<paragraph id="BE009665FFB7FFB6AB8EFF16FC1203AA" blockId="20.[151,919,152,211]" box="[151,919,152,178]" pageId="20" pageNumber="231">
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<heading id="E5482109FFB7FFB6AB8EFF16FC1203AA" bold="true" box="[151,919,152,178]" fontSize="11" level="1" pageId="20" pageNumber="231" reason="1">
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<emphasis id="8CCB4A77FFB7FFB6AB8EFF16FC1203AA" bold="true" box="[151,919,152,178]" pageId="20" pageNumber="231">
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Genus
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<taxonomicName id="79BFEDE6FFB7FFB6ABF7FF16FEF603AA" ID-CoL="7ZRF" ID-ENA="1830341" box="[238,371,152,178]" class="Actinopterygii" family="Cichlidae" genus="Trichromis" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="231" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="8CCB4A77FFB7FFB6ABF7FF16FEF603AA" bold="true" box="[238,371,152,178]" italics="true" pageId="20" pageNumber="231">Trichromis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="97F8F70CFFB7FFB6AA62FF16FE6C03AA" box="[379,489,152,178]" pageId="20" pageNumber="231" rank="genus">gen. nov.</taxonomicNameLabel>
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<bibRefCitation id="DA2EEB94FFB7FFB6AAE9FF16FC1203AA" author="McMahan" box="[496,919,152,178]" pageId="20" pageNumber="231" refString="Matamoros, W. A., McMahan, C. D., Chakrabarty, P., Albert, J. S. & Schaefer, J. F. (2015) Derivation of the freshwater fish fauna of Central America revisited: Myers's hypothesis in the twenty-first century. Cladistics, 31 (2), 177 - 188. http: // dx. doi. org / 10.1111 / cla. 12081" type="journal article" year="2015">McMahan and Chakrabarty 2015</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BE009665FFB7FFB6AB8EFF37FF7803CB" blockId="20.[151,919,152,211]" box="[151,253,185,211]" pageId="20" pageNumber="231">
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(
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<figureCitation id="26848AE0FFB7FFB6AB86FF37FF7103CB" box="[159,244,185,211]" captionStart="FIGURE 20" captionStartId="20.[151,250,1726,1748]" captionTargetBox="[166,1417,1105,1689]" captionTargetId="figure@20.[151,1436,1090,1704]" captionTargetPageId="20" captionText="FIGURE 20. Trichromis salvini, FMNH 109065, 69.76 mm SL." httpUri="https://zenodo.org/record/237067/files/figure.png" pageId="20" pageNumber="231">Fig. 20</figureCitation>
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)
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</paragraph>
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<paragraph id="BE009665FFB7FFB6AB8EFE8FFD300201" blockId="20.[151,1437,257,1038]" box="[151,693,257,282]" pageId="20" pageNumber="231">
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<emphasis id="8CCB4A77FFB7FFB6AB8EFE8FFEE60202" bold="true" box="[151,355,257,282]" pageId="20" pageNumber="231">Inclusive species.</emphasis>
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<taxonomicName id="79BFEDE6FFB7FFB6AA73FE8CFE550202" box="[362,464,257,282]" class="Actinopterygii" family="Cichlidae" genus="Trichromis" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="231" phylum="Chordata" rank="species" species="salvini">
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<emphasis id="8CCB4A77FFB7FFB6AA73FE8CFEF20201" box="[362,375,258,281]" italics="true" pageId="20" pageNumber="231">T</emphasis>
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.
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<emphasis id="8CCB4A77FFB7FFB6AA9CFE8FFE550202" box="[389,464,257,282]" italics="true" pageId="20" pageNumber="231">salvini</emphasis>
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</taxonomicName>
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(
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<typeStatus id="610428C7FFB7FFB6AAC7FE8FFD950201" box="[478,528,257,281]" pageId="20" pageNumber="231">type</typeStatus>
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by monotypy)
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFB7FFB6ABDEFEABFDBE02EA" pageId="20" pageNumber="231" type="diagnosis">
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<paragraph id="BE009665FFB7FFB6ABDEFEABFDBE02EA" blockId="20.[151,1437,257,1038]" pageId="20" pageNumber="231">
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<emphasis id="8CCB4A77FFB7FFB6ABDEFEABFEC40226" bold="true" box="[199,321,293,318]" pageId="20" pageNumber="231">Diagnosis.</emphasis>
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<taxonomicName id="79BFEDE6FFB7FFB6AA53FEABFE410226" box="[330,452,293,318]" class="Actinopterygii" family="Cichlidae" genus="Trichromis" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="8CCB4A77FFB7FFB6AA53FEABFE410226" box="[330,452,293,318]" italics="true" pageId="20" pageNumber="231">Trichromis</emphasis>
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</taxonomicName>
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is diagnosed by the presence of two dark lines down the length of the body, one just above the distal tip of the pectoral fin, and the other right below the dorsal fin. The mouth is small and terminal and the overall body shape is oval or oblong. Dark, narrow bars or lines are present, typically three or four, from in between the eyes to the predorsal region anterior to the dorsal fin. The interorbital markings are not as broad as those in
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<taxonomicName id="79BFEDE6FFB7FFB6ABEEFE3BFEFA02D6" box="[247,383,437,462]" class="Actinopterygii" family="Cichlidae" genus="Maskaheros" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="8CCB4A77FFB7FFB6ABEEFE3BFEFA02D6" box="[247,383,437,462]" italics="true" pageId="20" pageNumber="231">Maskaheros</emphasis>
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</taxonomicName>
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. Both sexes of species within this genus are colorful (yellow, red, blue/green), and coloration becomes more vivid during breeding.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFB7FFB6ABDEFE73FB14010E" box="[199,1169,509,534]" pageId="20" pageNumber="231" type="distribution">
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<paragraph id="BE009665FFB7FFB6ABDEFE73FB14010E" blockId="20.[151,1437,257,1038]" box="[199,1169,509,534]" pageId="20" pageNumber="231">
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<emphasis id="8CCB4A77FFB7FFB6ABDEFE73FEE5010E" bold="true" box="[199,352,509,534]" pageId="20" pageNumber="231">Distribution.</emphasis>
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Rivers of the Atlantic slope of southern
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<collectingCountry id="C6A8D6F5FFB7FFB6A831FE73FC05010E" box="[808,896,509,534]" name="Mexico" pageId="20" pageNumber="231">Mexico</collectingCountry>
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,
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<collectingCountry id="C6A8D6F5FFB7FFB6A892FE73FC50010E" box="[907,981,509,534]" name="Belize" pageId="20" pageNumber="231">Belize</collectingCountry>
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, and
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<collectingCountry id="C6A8D6F5FFB7FFB6AF09FE73FB08010E" box="[1040,1165,509,534]" name="Guatemala" pageId="20" pageNumber="231">Guatemala</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFB7FFB6ABDEFDAFFCAE0146" pageId="20" pageNumber="231" type="etymology">
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<paragraph id="BE009665FFB7FFB6ABDEFDAFFCAE0146" blockId="20.[151,1437,257,1038]" pageId="20" pageNumber="231">
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<emphasis id="8CCB4A77FFB7FFB6ABDEFDAFFECB0122" bold="true" box="[199,334,545,570]" pageId="20" pageNumber="231">Etymology.</emphasis>
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Gender feminine. “Tri” is Greek for three, in reference to the three vibrant colors defining this genus (red, yellow, and blue). “-chromis” is Greek for fish.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFB7FFB6ABDEFDE7FE9A00BA" pageId="20" pageNumber="231" type="discussion">
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<paragraph id="BE009665FFB7FFB6ABDEFDE7FE9A00BA" blockId="20.[151,1437,257,1038]" pageId="20" pageNumber="231">
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<emphasis id="8CCB4A77FFB7FFB6ABDEFDE7FECA019A" bold="true" box="[199,335,617,642]" pageId="20" pageNumber="231">Comments.</emphasis>
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This species of northern Middle American cichlid has been recovered in several different phylogenetic positions across studies. Most of these discrepancies are likely due to differences in taxonomic sampling among studies.
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<bibRefCitation id="DA2EEB94FFB7FFB6AAA0FD3CFD4B01D2" box="[441,718,689,714]" pageId="20" pageNumber="231" refString="Matamoros, W. A., McMahan, C. D., Chakrabarty, P., Albert, J. S. & Schaefer, J. F. (2015) Derivation of the freshwater fish fauna of Central America revisited: Myers's hypothesis in the twenty-first century. Cladistics, 31 (2), 177 - 188. http: // dx. doi. org / 10.1111 / cla. 12081" type="journal article">
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Matamoros
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<emphasis id="8CCB4A77FFB7FFB6A958FD3CFDF101D2" box="[577,628,689,714]" italics="true" pageId="20" pageNumber="231">et al</emphasis>
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. (2015)
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</bibRefCitation>
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recovered this species outside of the herichthyins and sister to
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<taxonomicName id="79BFEDE6FFB7FFB6AB8EFD5BFE2001F6" box="[151,421,725,750]" class="Actinopterygii" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="231" phylum="Chordata" rank="species" species="nanoluteus">
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<emphasis id="8CCB4A77FFB7FFB6AB8EFD5BFE2001F6" box="[151,421,725,750]" italics="true" pageId="20" pageNumber="231">Cryptoheros nanoluteus</emphasis>
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</taxonomicName>
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.
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<bibRefCitation id="DA2EEB94FFB7FFB6AAADFD5BFD0F01F6" box="[436,650,725,750]" pageId="20" pageNumber="231" refString="Rican, O., Pialek, L., Zardoya, R., Doadrio, I. & Zrzav, Y. (2013) Biogeography of the Mesoamerican Cichlidae (Teleostei: Heroini): colonization through the GAARlandia land bridge and early diversification. Journal of Biogeography, 40 (3), 579 - 593. http: // dx. doi. org / 10.1111 / jbi. 12023" type="journal article">
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Říčan
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<emphasis id="8CCB4A77FFB7FFB6AAE4FD58FDB501F6" box="[509,560,725,750]" italics="true" pageId="20" pageNumber="231">et al</emphasis>
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. (2013)
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</bibRefCitation>
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recovered this species as the sister group to
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<taxonomicName id="79BFEDE6FFB7FFB6AF9EFD5BFA8801F6" box="[1159,1293,725,750]" class="Actinopterygii" family="Cichlidae" genus="Thorichthys" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="231" phylum="Chordata" rank="genus">
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<emphasis id="8CCB4A77FFB7FFB6AF9EFD5BFA8801F6" box="[1159,1293,725,750]" italics="true" pageId="20" pageNumber="231">Thorichthys</emphasis>
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</taxonomicName>
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, and López- Fernández
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<emphasis id="8CCB4A77FFB7FFB6AA0DFD74FEC3000A" box="[276,326,761,786]" italics="true" pageId="20" pageNumber="231">et al</emphasis>
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. (2010) found this species to be outside of the herichthyin cichlids and the sister group to a large clade of several different genera. In nearly all cases (including the present study),
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<taxonomicName id="79BFEDE6FFB7FFB6AF29FC90FB12002E" box="[1072,1175,797,822]" class="Actinopterygii" family="Cichlidae" genus="Trichromis" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="231" phylum="Chordata" rank="species" species="salvini">
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<emphasis id="8CCB4A77FFB7FFB6AF29FC90FBB8002D" box="[1072,1085,798,821]" italics="true" pageId="20" pageNumber="231">T</emphasis>
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.
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<emphasis id="8CCB4A77FFB7FFB6AF55FC93FB12002E" box="[1100,1175,797,822]" italics="true" pageId="20" pageNumber="231">salvini</emphasis>
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</taxonomicName>
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is recovered in its own clade and not the sister group to any species or clade of species even remotely similar morphologically. Thus, given the phylogenetic position and the morphological distinctiveness of this species, we describe a new genus to contain this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFB7FFB6ABDEFC23FF6C0716" pageId="20" pageNumber="231" type="materials_examined">
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<paragraph id="BE009665FFB7FFB6ABDEFC23FF6C0716" blockId="20.[151,1437,257,1038]" pageId="20" pageNumber="231">
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<emphasis id="8CCB4A77FFB7FFB6ABDEFC23FE3600DE" bold="true" box="[199,435,941,966]" pageId="20" pageNumber="231">Material examined.</emphasis>
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<collectionCode id="D8AE0EA0FFB7FFB6AAA7FC20FD9800DE" box="[446,541,942,966]" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="20" pageNumber="231">LSUMZ</collectionCode>
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16257 [n=2,
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<collectingCountry id="C6A8D6F5FFB7FFB6A9A7FC23FCB900DE" box="[702,828,941,966]" name="Guatemala" pageId="20" pageNumber="231">Guatemala</collectingCountry>
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: Lago Peten-Itza],
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<collectionCode id="D8AE0EA0FFB7FFB6AF03FC20FBFC00DE" box="[1050,1145,942,966]" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="20" pageNumber="231">LSUMZ</collectionCode>
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16366 [n=4,
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<collectingCountry id="C6A8D6F5FFB7FFB6AE02FC23FA1C00DE" box="[1307,1433,941,966]" name="Guatemala" pageId="20" pageNumber="231">Guatemala</collectingCountry>
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: Río La Pasion],
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<collectionCode id="D8AE0EA0FFB7FFB6AA44FC5CFE3900F2" box="[349,444,978,1002]" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="20" pageNumber="231">LSUMZ</collectionCode>
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16417 [n=4,
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<collectingCountry id="C6A8D6F5FFB7FFB6A97DFC5FFD6700F2" box="[612,738,977,1002]" name="Guatemala" pageId="20" pageNumber="231">Guatemala</collectingCountry>
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: Lago Yaxhá],
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<collectionCode id="D8AE0EA0FFB7FFB6A882FC5CFC6900F1" LSID="urn:lsid:biocol.org:col:34795" box="[923,1004,978,1001]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="231">FMNH</collectionCode>
|
|
109065 [n=18,
|
|
<collectingCountry id="C6A8D6F5FFB7FFB6AFA9FC5FFAAB00F2" box="[1200,1326,977,1002]" name="Guatemala" pageId="20" pageNumber="231">Guatemala</collectingCountry>
|
|
: Río
|
|
<collectingCountry id="C6A8D6F5FFB7FFB6AE6BFC5CFF5B0716" name="Saint Pierre and Miquelon" pageId="20" pageNumber="231">San Pedro</collectingCountry>
|
|
].
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |