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<document id="DD77BA34B0EB26A6EEC97AA5154E01B9" ID-CLB-Dataset="31106" ID-DOI="10.11646/zootaxa.4365.1.1" ID-GBIF-Dataset="8e2d27e3-dec0-4ebb-bcd3-dd16e56c8092" ID-ISSN="1175-5326" ID-Zenodo-Dep="1125219" ID-ZooBank="5F132840-6E2F-42E6-8DF2-9EF191DE243C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513931221510" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2017" docId="03FC87DCFFA99D33FF0D3B2BFD1CFABC" docLanguage="en" docName="zootaxa.4365.1.1.pdf.imf" docOrigin="Zootaxa 4365 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium puetzi Shavrin &amp; Smetana 2017, sp.n." docType="treatment" docVersion="8" lastPageNumber="22" masterDocId="FFC5FFA4FFBB9D26FF9A3F56FFB8FFFD" masterDocTitle="A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups" masterLastPageNumber="39" masterPageNumber="1" pageNumber="19" updateTime="1698511183827" updateUser="ExternalLinkService">
<mods:mods id="E3A2C04CFF6218F32483E8CE627F6419" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B4BB1418B3E4A8CF4BF5CE27731E55CA">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups</mods:title>
</mods:titleInfo>
<mods:name id="1777008BA10E45E5418D5FAF347D075B" type="personal">
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<mods:namePart id="6AD056CAD35A06B509BBDC006956AC29">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="F8FEA5BB727B53DB50A65617259D731B">Smetana, Aleš</mods:namePart>
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<mods:title id="FDC942B5A0CC3580B03484234C32221D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CF2FFFDD1000BA977E7B033F58D1A9BC">
<mods:date id="FB04450BB86F7221702D399A250E0C90">2017</mods:date>
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<mods:number id="81D817A6E0B21C4D2395B3793074435E">2017-12-15</mods:number>
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<mods:number id="4DBCDE432CF30CE753BBF8A2313F2E3C">4365</mods:number>
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<mods:classification id="D70F5210DD9684410AD5188C154E78B9">journal article</mods:classification>
<mods:identifier id="0D687A2F1D3598CDEE449B45F9057A88" type="CLB-Dataset">31106</mods:identifier>
<mods:identifier id="6458F5A7C9A4C071E14FC2CADD72D04A" type="DOI">10.11646/zootaxa.4365.1.1</mods:identifier>
<mods:identifier id="436D0E34936A7A31CC7A539FD0C8029D" type="GBIF-Dataset">8e2d27e3-dec0-4ebb-bcd3-dd16e56c8092</mods:identifier>
<mods:identifier id="78B4E0005FD03A0649B39E2980FE7B07" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FC87DCFFA99D33FF0D3B2BFD1CFABC" ID-DOI="http://doi.org/10.5281/zenodo.6017646" ID-GBIF-Taxon="139444205" ID-Zenodo-Dep="6017646" LSID="urn:lsid:plazi:treatment:03FC87DCFFA99D33FF0D3B2BFD1CFABC" httpUri="http://treatment.plazi.org/id/03FC87DCFFA99D33FF0D3B2BFD1CFABC" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19">
<subSubSection id="C34F6541FFA99D34FF0D3B2BFDA8FB6A" box="[151,528,1149,1175]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BEA36CAFFA99D34FF0D3B2BFDA8FB6A" blockId="18.[151,528,1149,1208]" box="[151,528,1149,1175]" pageId="18" pageNumber="19">
<heading id="D0A281A6FFA99D34FF0D3B2BFDA8FB6A" bold="true" box="[151,528,1149,1175]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B921EAD8FFA99D34FF0D3B2BFDA8FB6A" bold="true" box="[151,528,1149,1175]" pageId="18" pageNumber="19">
3.2.2.3.
<taxonomicName id="4C554D49FFA99D34FF6C3B2BFE76FB6A" ID-CoL="5VCFK" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[246,462,1149,1175]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAD8FFA99D34FF6C3B2BFE76FB6A" bold="true" box="[246,462,1149,1175]" italics="true" pageId="18" pageNumber="19">Anthobium puetzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257A3FFA99D34FE4F3B2BFDA8FB6A" box="[469,528,1149,1175]" pageId="18" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA99D34FF0D3BC9FE3AFB45" box="[151,386,1183,1208]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BEA36CAFFA99D34FF0D3BC9FE3AFB45" blockId="18.[151,528,1149,1208]" box="[151,386,1183,1208]" pageId="18" pageNumber="19">
(
<figureCitation id="136E2A4FFFA99D34FF053BC9FF44FB45" box="[159,252,1183,1208]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1852,1875]" captionTargetBox="[175,1412,205,1815]" captionTargetId="figure@16.[150,1436,193,1831]" captionTargetPageId="16" captionText="FIGURES 39–41. Habitus of Anthobium morchella group: 39—A. morchella (paratype), 40—A. hydraenoides (paratype), 41—A.puetzi (holotype).Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1125237/files/figure.png" pageId="18" pageNumber="19">Figs. 41</figureCitation>
,
<figureCitation id="136E2A4FFFA99D34FE923BC9FEC1FB45" box="[264,377,1183,1208]" captionStart-0="FIGURES 56–61" captionStart-1="FIGURE62" captionStartId-0="19.[151,264,1976,1999]" captionStartId-1="20.[151,250,1942,1964]" captionTargetBox-0="[208,1373,208,1954]" captionTargetBox-1="[187,1400,1309,1921]" captionTargetId-0="figure@19.[170,1417,183,1968]" captionTargetId-1="figure@20.[187,1400,1309,1921]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 56–61. Aedeagus of Anthobium morchella (56, 59—paratype), A. hydraenoides (57, 60—paratype) and A. puetzi (58, 61—paratype): 56–58—parameral view, 59–61—lateral view.Scale bars: 0.1 mm." captionText-1="FIGURE62.Distribution of Anthobium morchella group inChina:A.morchella(black square), A.hydraenoides(black circle), A. puetzi (black triangle)." httpUri-0="https://zenodo.org/record/1125239/files/figure.png" httpUri-1="https://zenodo.org/record/1125241/files/figure.png" pageId="18" pageNumber="19">58, 61–62</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA99D34FF0D3BB1FED1F9B9" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BEA36CAFFA99D34FF0D3BB1FDA4FA91" blockId="18.[151,1437,1255,2037]" pageId="18" pageNumber="19">
<emphasis id="B921EAD8FFA99D34FF0D3BB1FE71FAFD" bold="true" box="[151,457,1255,1280]" pageId="18" pageNumber="19">
<typeStatus id="54EE8868FFA99D34FF0D3BB1FF6AFAFD" box="[151,210,1255,1280]" pageId="18" pageNumber="19">Type</typeStatus>
material examined:
</emphasis>
<materialsCitation id="3B3D3C97FFA99D34FE4E3BB1FDA4FA91" ID-GBIF-Occurrence="1799096744" collectingDate="20. / 23. V. 1997" collectionCode="SNSD" collectorName="A. Pütz’" country="China" elevation="3000" latitude="30.03" location="Daxue Shan" longitude="102" municipality="Ganzi pref" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="54EE8868FFA99D34FE4E3BB1FD84FAFD" box="[468,572,1255,1280]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
<specimenCount id="9D53FD43FFA99D34FDDD3BBEFDE7FB02" box="[583,607,1256,1279]" pageId="18" pageNumber="19" type="male">♂</specimenCount>
: ‘
<collectingCountry id="F342765AFFA99D34FDE93BB1FD07FAFD" box="[627,703,1255,1280]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
:
<collectingRegion id="4991F828FFA99D34FD573BB1FC92FAFD" box="[717,810,1255,1280]" country="China" name="Sichuan" pageId="18" pageNumber="19">Sichuan</collectingRegion>
,
<collectingMunicipality id="6B8EACB0FFA99D34FCA03BB1FC00FAFD" box="[826,952,1255,1280]" pageId="18" pageNumber="19">Ganzi pref</collectingMunicipality>
. |
<location id="8E8A6011FFA99D34FC4D3BBEFBDEFAFD" LSID="urn:lsid:plazi:treatment:03FC87DCFFA99D33FF0D3B2BFD1CFABC:8E8A6011FFA99D34FC4D3BBEFBDEFAFD" box="[983,1126,1255,1280]" country="China" latitude="30.03" municipality="Ganzi pref" name="Daxue Shan" pageId="18" pageNumber="19" stateProvince="Sichuan">Daxue Shan</location>
,
<geoCoordinate id="EE61500DFFA99D34FBEF3BBEFB7AFAFD" box="[1141,1218,1256,1280]" pageId="18" pageNumber="19">102.00</geoCoordinate>
<geoCoordinate id="EE61500DFFA99D34FB573BBEFA96FB02" box="[1229,1326,1256,1280]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="555" value="30.03">30.03 N</geoCoordinate>
, |
<quantity id="4CAD9B2FFFA99D34FAD43BB1FA38FAFD" box="[1358,1408,1255,1280]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="km" value="5.0">
5
<location id="8E8A6011FFA99D34FAC03BB1FA38FAFD" LSID="urn:lsid:plazi:treatment:03FC87DCFFA99D33FF0D3B2BFD1CFABC:8E8A6011FFA99D34FAC03BB1FA38FAFD" box="[1370,1408,1255,1280]" country="China" latitude="30.03" municipality="Ganzi pref" name="km E Kangding, river valley" pageId="18" pageNumber="19" stateProvince="Sichuan">km</location>
</quantity>
E Kangding, river valley | ca.
<elevation id="0078D1F9FFA99D34FE7C3A5AFD8FFAD9" box="[486,567,1292,1316]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="m" value="3000.0">
<quantity id="4CAD9B2FFFA99D34FE7C3A5AFD8FFAD9" box="[486,567,1292,1316]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="m" value="3000.0">3000m</quantity>
</elevation>
,
<collectingDate id="EFAFE9E2FFA99D34FDDC3A5AFD5EFAD9" box="[582,742,1292,1316]" pageId="18" pageNumber="19">
20./
<date id="FFEB100AFFA99D34FDE83A5AFD5EFAD9" box="[626,742,1292,1316]" pageId="18" pageNumber="19" value="1997-05-23">23.V.1997</date>
</collectingDate>
, leg.
<collectorName id="26A0531CFFA99D34FCB13A5AFC35FAD9" box="[811,909,1292,1316]" pageId="18" pageNumber="19">A. Pütz’</collectorName>
&lt;
<locationDeviation id="9A582FE8FFA99D34FC323A5AFE04FA90" pageId="18" pageNumber="19">
rectangular label, printed&gt;, ‘HOLOTYPE |
<taxonomicName id="4C554D49FFA99D34FF0D3A66FEAAFAB5" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[151,274,1328,1352]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B921EAD8FFA99D34FF0D3A66FEAAFAB5" box="[151,274,1328,1352]" italics="true" pageId="18" pageNumber="19">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C554D49FFA99D34FEBF3A66FED0FABA" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[293,360,1328,1351]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAD8FFA99D34FEBF3A66FED0FABA" box="[293,360,1328,1351]" italics="true" pageId="18" pageNumber="19">puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFA99D34FEF53A79FE1FFAB5" bold="true" box="[367,423,1327,1352]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A21257A3FFA99D34FEF53A79FE1FFAB5" box="[367,423,1327,1352]" pageId="18" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. 2017’ &lt;red rectangular label, printed&gt; (temporarily in CAP, to be eventually deposited in
</locationDeviation>
<collectionCode id="ED44AE0FFFA99D34FE593A02FDB7FA91" box="[451,527,1364,1388]" pageId="18" pageNumber="19">SNSD</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEA36CAFFA99D34FF5D3A2EFED1F9B9" blockId="18.[151,1437,1255,2037]" pageId="18" pageNumber="19">
Paratypes (3 specimens): 1 ♂, 1 ♀: same label as the holotype (CAP); 1 ♂ [plastic plate with aedeagus, abdominal tergite VIII, sternite VIII and apical segment in Canadian balsam was pinned under the plate with beetle]: ‘CHINA: W-Sichuan | Aba Tibetian Aut. Pref. | Weizhou Co., Qionglai Shan | Wolong Valley,
<quantity id="4CAD9B2FFFA99D34FA8B3AE9FAE9FA25" box="[1297,1361,1471,1496]" metricMagnitude="4" metricUnit="m" metricValue="6.9" pageId="18" pageNumber="19" unit="km" value="69.0">69km</quantity>
WSW | Guanxian,
<quantity id="4CAD9B2FFFA99D34FE833AB2FECBFA01" box="[281,371,1508,1533]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="18" pageNumber="19" unit="m" value="3900.0">3900 m</quantity>
, 30.53,67N | 102.54,87E,
<date id="FFEB100AFFA99D34FD383AB2FC93FA00" box="[674,811,1508,1533]" pageId="18" pageNumber="19" value="1999-07-15">15.VII.1999</date>
| leg. A. Pütz’ &lt;rectangular label printed&gt; (CAS). All paratypes with additional red rectangular printed label: ‘PARATYPE |
<taxonomicName id="4C554D49FFA99D34FC50395EFBFDF9DD" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[970,1093,1544,1568]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B921EAD8FFA99D34FC50395EFBFDF9DD" box="[970,1093,1544,1568]" italics="true" pageId="18" pageNumber="19">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C554D49FFA99D34FBC5395EFB1AF9E2" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1119,1186,1544,1567]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAD8FFA99D34FBC5395EFB1AF9E2" box="[1119,1186,1544,1567]" italics="true" pageId="18" pageNumber="19">puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFA99D34FB373951FB5CF9DD" bold="true" box="[1197,1252,1543,1568]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A21257A3FFA99D34FB373951FB5CF9DD" box="[1197,1252,1543,1568]" pageId="18" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. 2017’.
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA99D32FF5D3919FB49FC58" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph id="8BEA36CAFFA99D34FF5D3919FB78F94D" blockId="18.[151,1437,1255,2037]" pageId="18" pageNumber="19">
<emphasis id="B921EAD8FFA99D34FF5D3919FEE9F995" bold="true" box="[199,337,1615,1640]" pageId="18" pageNumber="19">Description</emphasis>
. Measurements (n=4): HW: 0.85–0.92, HL: 0.52–0.59; AL (
<typeStatus id="54EE8868FFA99D34FB9B3919FBD2F995" box="[1025,1130,1615,1640]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
): 2.03; OL: 0.24–0.25; PL: 0.79–0.82; PW: 1.26–1.37; ESL: 1.90–2.00; EW: 1.75–1.97; AW: 1.35–1.55; MTbL (
<typeStatus id="54EE8868FFA99D34FB083922FB43F970" box="[1170,1275,1652,1677]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
): 1.12, MTrL (
<typeStatus id="54EE8868FFA99D34FF0539C1FEB0F94D" box="[159,264,1687,1712]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
): 0.52 (MTrL 1–4: 0.32; MTrL 5: 0.20); AedL: 0.90–0.97; TL: 3.75 (
<typeStatus id="54EE8868FFA99D34FB9739C1FBCBF94D" box="[1037,1139,1687,1712]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
)–4.75.
</paragraph>
<paragraph id="8BEA36CAFFA99D34FF5D39EBFC87F809" blockId="18.[151,1437,1255,2037]" pageId="18" pageNumber="19">
In external appearance similar to
<taxonomicName id="4C554D49FFA99D34FDD639EAFD66F929" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[588,734,1724,1748]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="morchella" status="sp. nov.">
<emphasis id="B921EAD8FFA99D34FDD639EAFD66F929" box="[588,734,1724,1748]" italics="true" pageId="18" pageNumber="19">A. morchella</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFA99D34FD7239EAFC98F928" bold="true" box="[744,800,1724,1749]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A21257A3FFA99D34FD7239EAFC98F928" box="[744,800,1724,1749]" pageId="18" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
Body moderately elongated. Reddish-brown to brown; antennomeres 3–11 brown; lateral and basal portions of pronotum, latero-apical and apical portions of elytra, apical segments of abdomen yellow-brown; mouthparts, antennomeres 1 and 2 and legs yellow. Head behind postocular ridges and lateral portions of neck with fine small cellular sculpture, larger and smoothed in middle of neck. Punctation of infraorbital ridges moderately regular, markedly denser than that on middle part of head; middle part of neck with transverse, irregular row of deep large punctures; pronotum with coarse and deep punctures, denser on medio-basal portion, irregular and somewhat smaller on medio-apical part, more rugose on lateral portions; scutellum with several small punctures; punctation of elytra as that on pronotum, slightly denser and smaller in prescutellar and basal area near suture. Habitus as in
<figureCitation id="136E2A4FFFA99D34FD73388AFC83F809" box="[745,827,2012,2037]" captionStart="FIGURES 39–41" captionStartId="16.[151,264,1852,1875]" captionTargetBox="[175,1412,205,1815]" captionTargetId="figure@16.[150,1436,193,1831]" captionTargetPageId="16" captionText="FIGURES 39–41. Habitus of Anthobium morchella group: 39—A. morchella (paratype), 40—A. hydraenoides (paratype), 41—A.puetzi (holotype).Scale bars:1.0 mm." httpUri="https://zenodo.org/record/1125237/files/figure.png" pageId="18" pageNumber="19">Fig. 41</figureCitation>
.
</paragraph>
<caption id="DF2A6642FFA89D35FF0D38EEFBB1F80D" httpUri="https://zenodo.org/record/1125239/files/figure.png" pageId="19" pageNumber="20" targetBox="[208,1373,208,1954]" targetPageId="19">
<paragraph id="8BEA36CAFFA89D35FF0D38EEFBB1F80D" blockId="19.[151,1435,1976,2032]" pageId="19" pageNumber="20">
<emphasis id="B921EAD8FFA89D35FF0D38EEFEEDF833" bold="true" box="[151,341,1976,1999]" pageId="19" pageNumber="20">FIGURES 56–61.</emphasis>
Aedeagus of
<taxonomicName id="4C554D49FFA89D35FE7C38EFFD7BF832" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[486,707,1977,1999]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="morchella">
<emphasis id="B921EAD8FFA89D35FE7C38EFFD7BF832" box="[486,707,1977,1999]" italics="true" pageId="19" pageNumber="20">Anthobium morchella</emphasis>
</taxonomicName>
(56, 59—paratype),
<taxonomicName id="4C554D49FFA89D35FC0638ECFBFAF832" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[924,1090,1977,1999]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hydraenoides">
<emphasis id="B921EAD8FFA89D35FC0638ECFBFAF832" box="[924,1090,1977,1999]" italics="true" pageId="19" pageNumber="20">A. hydraenoides</emphasis>
</taxonomicName>
(57, 60—paratype) and
<taxonomicName id="4C554D49FFA89D35FAD838ECFA23F832" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1346,1435,1978,1999]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="B921EAD8FFA89D35FAD838ECFA23F832" box="[1346,1435,1978,1999]" italics="true" pageId="19" pageNumber="20">A. puetzi</emphasis>
</taxonomicName>
(58, 61—paratype): 56–58—parameral view, 59–61—lateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEA36CAFFAF9D32FF5D3FC1FBA7FE2D" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
Head 1.5–1.6 times as wide as long, slightly widened posteriad; middle portion of head and infraorbital ridges slightly elevated; median portion with a pair of indistinct medial impressions behind level of anterior margin of eyes and small transverse impression on level of posterior margin of eyes, with very deep elongated grooves in front of ocelli reaching apical third of length of eye; head gradually narrowed toward neck behind distinctly smoothed postocular ridges; anterior portion between antennal insertion and anterior margin of eye with wide, moderately deep notch. Ocelli situated slightly behind of level of postocular ridge, distance between ocelli slightly larger than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp less than three times as long as preceeding segment. Measurements of antennomeres (
<typeStatus id="54EE8868FFAF9D32FC283EC2FBA3FE50" box="[946,1051,404,429]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
): 1: 0.21 × 0.09; 2: 0.15 × 0.06; 3: 0.18 × 0.06; 4–7: 0.19 × 0.06; 8: 0.17 × 0.06; 9–10: 0.17 × 0.07; 11: 0.22 × 0.07.
</paragraph>
<paragraph id="8BEA36CAFFAF9D32FF5D3E8BFCCFFD9D" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">Pronotum 1.5–1.6 times as wide as long, 1.4 times as wide as head, widest near middle; lateral edge with moderately uniform crenulation, distinctly larger in middle; middle elevation of pronotum with vague, moderately wide longitudinal and a pair of suboval medio-basal impressions; small area above medio-basal impressions markedly elevated; lateral portions narrow, indistinctly explanate.</paragraph>
<paragraph id="8BEA36CAFFAF9D32FF5D3D3AFDA6FD0D" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
Elytra more than twice as long as pronotum, gradually widened apicad, reaching basal part of abdominal tergite V–VII; shoulders indistinctly protruded anteriad; middle parts of each elytron with vague irregular elevations between punctures (one
<typeStatus id="54EE8868FFAF9D32FDBE3DE3FD3DFD30" box="[548,645,693,717]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
with indistinct longitudinal elevation beginning behind shoulders and stretching to apical third of elytra).
</paragraph>
<paragraph id="8BEA36CAFFAF9D32FF5D3DAAFCA5FC7D" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
Male. Apical margins of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (
<figureCitation id="136E2A4FFFAF9D32FA993DAAFAE0FCE8" box="[1283,1368,764,789]" captionStart="FIGURES 56–61" captionStartId="19.[151,264,1976,1999]" captionTargetBox="[208,1373,208,1954]" captionTargetId="figure@19.[170,1417,183,1968]" captionTargetPageId="19" captionText="FIGURES 56–61. Aedeagus of Anthobium morchella (56, 59—paratype), A. hydraenoides (57, 60—paratype) and A. puetzi (58, 61—paratype): 56–58—parameral view, 59–61—lateral view.Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1125239/files/figure.png" pageId="20" pageNumber="21">Fig. 58</figureCitation>
) with wide basal part slightly narrowed towards subtruncate apex of median lobe; parameres significantly exceeding apex of median lobe, with moderately long apical and two preapical setae; internal sac narrow and long, markedly more sclerotized basally. Aedeagus laterally as in
<figureCitation id="136E2A4FFFAF9D32FD5D3C31FCA1FC7D" box="[711,793,871,896]" captionStart="FIGURES 56–61" captionStartId="19.[151,264,1976,1999]" captionTargetBox="[208,1373,208,1954]" captionTargetId="figure@19.[170,1417,183,1968]" captionTargetPageId="19" captionText="FIGURES 56–61. Aedeagus of Anthobium morchella (56, 59—paratype), A. hydraenoides (57, 60—paratype) and A. puetzi (58, 61—paratype): 56–58—parameral view, 59–61—lateral view.Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/1125239/files/figure.png" pageId="20" pageNumber="21">Fig. 61</figureCitation>
.
</paragraph>
<paragraph id="8BEA36CAFFAF9D32FF5D3CDAFB49FC58" blockId="20.[151,1437,151,1256]" box="[199,1265,908,933]" pageId="20" pageNumber="21">Female. Apical margins of abdominal tergite VIII and sternite VIII straight to slightly rounded.</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFAF9D32FF5D3CF9FEAAFB5D" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BEA36CAFFAF9D32FF5D3CF9FEAAFB5D" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
<emphasis id="B921EAD8FFAF9D32FF5D3CF9FE09FC35" bold="true" box="[199,433,943,968]" pageId="20" pageNumber="21">Comparative notes.</emphasis>
Based on the presence of indistinct irregular elevations between punctures on the elytra and parameres significantly exceeding apex of median lobe of the aedeagus,
<taxonomicName id="4C554D49FFAF9D32FBB53C82FB2FFC11" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1071,1175,980,1004]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAD8FFAF9D32FBB53C82FB2FFC11" box="[1071,1175,980,1004]" italics="true" pageId="20" pageNumber="21">A. puetzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257A3FFAF9D32FB3E3C82FB63FC10" box="[1188,1243,980,1005]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B921EAD8FFAF9D32FB3E3C82FB63FC10" bold="true" box="[1188,1243,980,1005]" pageId="20" pageNumber="21">sp.n.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C554D49FFAF9D32FA1E3C82FEB0FBED" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morchella" status="sp. nov.">
<emphasis id="B921EAD8FFAF9D32FA1E3C82FEB0FBED" italics="true" pageId="20" pageNumber="21">A. morchella</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFAF9D32FE953CA1FEFFFBED" bold="true" box="[271,327,1015,1040]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A21257A3FFAF9D32FE953CA1FEFFFBED" box="[271,327,1015,1040]" pageId="20" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, from which it differs by the head behind eyes slightly widened posteriad, by the larger and more rugose punctation of the pronotum with vague longitudinal and medio-basal impressions on middle elevation and by the relatively shorter parameres.
<taxonomicName id="4C554D49FFAF9D32FDAE3B16FD44FBAA" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[564,764,1088,1112]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puetzi" status="sp. nov.">
<emphasis id="B921EAD8FFAF9D32FDAE3B16FD44FBAA" box="[564,764,1088,1112]" italics="true" pageId="20" pageNumber="21">Anthobium puetzi</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFAF9D32FC9F3B69FC84FBA5" bold="true" box="[773,828,1087,1112]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A21257A3FFAF9D32FC9F3B69FC84FBA5" box="[773,828,1087,1112]" pageId="20" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from remaining two species of the
<taxonomicName id="4C554D49FFAF9D32FAB13B16FA23FBA5" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1323,1435,1088,1112]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morchella">
<emphasis id="B921EAD8FFAF9D32FAB13B16FA23FBA5" box="[1323,1435,1088,1112]" italics="true" pageId="20" pageNumber="21">morchella</emphasis>
</taxonomicName>
group by wider median lobe of the aedeagus and by the presence of two long preapical setae on apical portions of parameres.
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFAF9D32FF5D3BFAFEDEFB15" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BEA36CAFFAF9D32FF5D3BFAFEDEFB15" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
<emphasis id="B921EAD8FFAF9D32FF5D3BFAFED8FB38" bold="true" box="[199,352,1196,1221]" pageId="20" pageNumber="21">Distribution.</emphasis>
The new species is known from two locations (
<figureCitation id="136E2A4FFFAF9D32FCE93BFAFC7CFB38" box="[883,964,1196,1221]" captionStart="FIGURE62" captionStartId="20.[151,250,1942,1964]" captionTargetBox="[187,1400,1309,1921]" captionTargetId="figure@20.[187,1400,1309,1921]" captionTargetPageId="20" captionText="FIGURE62.Distribution of Anthobium morchella group inChina:A.morchella(black square), A.hydraenoides(black circle), A. puetzi (black triangle)." httpUri="https://zenodo.org/record/1125241/files/figure.png" pageId="20" pageNumber="21">Fig. 62</figureCitation>
) in Daxue
<collectingRegion id="4991F828FFAF9D32FBA53BFAFBCFFB38" box="[1087,1143,1196,1221]" country="Myanmar" name="Shan" pageId="20" pageNumber="21">Shan</collectingRegion>
and Qionglai
<collectingRegion id="4991F828FFAF9D32FA8E3BFAFAF4FB38" box="[1300,1356,1196,1221]" country="Myanmar" name="Shan" pageId="20" pageNumber="21">Shan</collectingRegion>
ranges in
<collectingRegion id="4991F828FFAF9D32FF2E3B99FEA9FB15" box="[180,273,1231,1256]" country="China" name="Sichuan" pageId="20" pageNumber="21">Sichuan</collectingRegion>
,
<collectingCountry id="F342765AFFAF9D32FE863B99FEDBFB15" box="[284,355,1231,1256]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF2A6642FFAF9D32FF0D38C0FE20F836" httpUri="https://zenodo.org/record/1125241/files/figure.png" pageId="20" pageNumber="21" targetBox="[187,1400,1309,1921]" targetPageId="20">
<paragraph id="8BEA36CAFFAF9D32FF0D38C0FE20F836" blockId="20.[151,1436,1942,1995]" pageId="20" pageNumber="21">
<emphasis id="B921EAD8FFAF9D32FF0D38C0FEA7F856" bold="true" box="[151,287,1942,1964]" pageId="20" pageNumber="21">FIGURE 62.</emphasis>
Distribution of
<taxonomicName id="4C554D49FFAF9D32FE2538C0FD22F851" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[447,666,1942,1964]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morchella">
<emphasis id="B921EAD8FFAF9D32FE2538C0FD22F851" box="[447,666,1942,1964]" italics="true" pageId="20" pageNumber="21">Anthobium morchella</emphasis>
</taxonomicName>
group in China:
<taxonomicName id="4C554D49FFAF9D32FCDE38C1FC7DF851" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[836,965,1942,1964]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morchella">
<emphasis id="B921EAD8FFAF9D32FCDE38C1FC7DF851" box="[836,965,1942,1964]" italics="true" pageId="20" pageNumber="21">A. morchella</emphasis>
</taxonomicName>
(black square),
<taxonomicName id="4C554D49FFAF9D32FBFC38C1FAB3F851" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1126,1291,1942,1964]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hydraenoides">
<emphasis id="B921EAD8FFAF9D32FBFC38C1FAB3F851" box="[1126,1291,1942,1964]" italics="true" pageId="20" pageNumber="21">A. hydraenoides</emphasis>
</taxonomicName>
(black circle),
<taxonomicName id="4C554D49FFAF9D32FF0D38E0FF57F836" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[151,239,1974,1995]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="B921EAD8FFAF9D32FF0D38E0FF57F836" box="[151,239,1974,1995]" italics="true" pageId="20" pageNumber="21">A. puetzi</emphasis>
</taxonomicName>
(black triangle).
</paragraph>
</caption>
<caption id="DF2A6642FFAE9D33FF0D3B29FF40FB49" httpUri="https://zenodo.org/record/1125243/files/figure.png" pageId="21" pageNumber="22" targetBox="[169,1418,193,1130]" targetPageId="21">
<paragraph id="8BEA36CAFFAE9D33FF0D3B29FF40FB49" blockId="21.[151,1436,1151,1204]" pageId="21" pageNumber="22">
<emphasis id="B921EAD8FFAE9D33FF0D3B29FE9CFB69" bold="true" box="[151,292,1151,1173]" pageId="21" pageNumber="22">FIGURE 63.</emphasis>
Type locality of
<taxonomicName id="4C554D49FFAE9D33FE793B29FD7BFB68" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[483,707,1151,1173]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="morchella">
<emphasis id="B921EAD8FFAE9D33FE793B29FD7BFB68" box="[483,707,1151,1173]" italics="true" pageId="21" pageNumber="22">Anthobium morchella</emphasis>
</taxonomicName>
in the Baima Shan range, northern Yunnan, China (photograph: M. Schülke).
</paragraph>
</caption>
<subSubSection id="C34F6541FFAE9D33FF5D3BB6FD1CFABC" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="8BEA36CAFFAE9D33FF5D3BB6FD1CFABC" blockId="21.[151,1436,1248,1345]" pageId="21" pageNumber="22">
<emphasis id="B921EAD8FFAE9D33FF5D3BB6FEF2FB04" bold="true" box="[199,330,1248,1273]" pageId="21" pageNumber="22">Bionomics.</emphasis>
Specimens were taken by sifting forest floor litter at elevations from
<elevation id="0078D1F9FFAE9D33FBCC3BB7FAFAFB04" box="[1110,1346,1248,1273]" metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.9" metricValueMin="3.0" pageId="21" pageNumber="22" unit="m" value="3450.0" valueMax="3900.0" valueMin="3000.0">
<quantity id="4CAD9B2FFFAE9D33FBCC3BB7FABEFB05" box="[1110,1286,1249,1273]" metricMagnitude="3" metricUnit="m" metricValue="3.45" metricValueMax="3.9" metricValueMin="3.0" pageId="21" pageNumber="22" unit="m" value="3450.0" valueMax="3900.0" valueMin="3000.0">3000 to 3900 m</quantity>
a.s.l.
</elevation>
<emphasis id="B921EAD8FFAE9D33FF5D3A52FEF6FAE0" bold="true" box="[199,334,1284,1309]" pageId="21" pageNumber="22">Etymology.</emphasis>
Patronymic, the species is named to honour our colleague
<location id="8E8A6011FFAE9D33FBBD3A52FB7DFAE0" LSID="urn:lsid:plazi:treatment:03FC87DCFFA99D33FF0D3B2BFD1CFABC:8E8A6011FFAE9D33FBBD3A52FB7DFAE0" box="[1063,1221,1284,1309]" country="Germany" municipality="Eisenhuttenstadt" name="Andreas Putz" pageId="21" pageNumber="22">Andreas Pütz</location>
(
<collectingMunicipality id="6B8EACB0FFAE9D33FB403A52FA20FAE0" box="[1242,1432,1284,1309]" pageId="21" pageNumber="22">Eisenhüttenstadt</collectingMunicipality>
,
<collectingCountry id="F342765AFFAE9D33FF0D3A7FFEBCFABC" box="[151,260,1321,1345]" name="Germany" pageId="21" pageNumber="22">Germany</collectingCountry>
), the collector of the
<typeStatus id="54EE8868FFAE9D33FE6A3A7FFD98FABC" box="[496,544,1321,1345]" pageId="21" pageNumber="22">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
</treatment>
</document>
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