treatments-xml/data/03/E6/87/03E687D6BB02FFB0F0C9F99BFE97F007.xml

<document id="3F0E3D82CC7FF4373C4ED0A44CD12981" ID-CLB-Dataset="24070" ID-DOI="10.11646/zootaxa.4732.2.3" ID-GBIF-Dataset="e82cebbb-f29b-47ec-9c64-4a60a5bf1630" ID-ISSN="1175-5326" ID-Zenodo-Dep="3663449" ID-ZooBank="51E842C7-FE20-44BD-A0E9-DE983EF676D8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581497104097" checkinUser="plazi" docAuthor="Podenas, Sigitas, Podeniene, Virginija, Seo, Hong-Yul, Kim, A-Young, Park, Sun-Jae, Byun, Hye-Woo, Kim, Heung-Chul &amp; Aukštikalnienė, Rasa" docDate="2020" docId="03E687D6BB02FFB0F0C9F99BFE97F007" docLanguage="en" docName="zootaxa.4732.2.3.pdf" docOrigin="Zootaxa 4732 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė 2020, sp. nov." docType="treatment" docUuid="E4E09E3A-7545-4407-B6FC-B2054F8AC3B8" docUuidSource="ZooBank" docVersion="8" lastPageNumber="292" masterDocId="FFDFFFAEBB07FFBBF05EFFA2FF98F670" masterDocTitle="New Atypophthalmus (Diptera: Limoniidae) from South Korea" masterLastPageNumber="294" masterPageNumber="281" pageNumber="286" updateTime="1698782730213" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0DCD042AEC7DF12381E766E182A27ED8">New Atypophthalmus (Diptera: Limoniidae) from South Korea</mods:title>
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<mods:namePart id="F663732EED387263C5F2198034CDF340">Podenas, Sigitas</mods:namePart>
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<mods:namePart id="4A20A53F495AA0690DB97C8D2FF74F28">Podeniene, Virginija</mods:namePart>
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<mods:namePart id="3043D0D6F9BF52A1DECB3C14C230008B">Seo, Hong-Yul</mods:namePart>
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<mods:namePart id="E8C4C4DBE5947DB01008CB98318F5922">Kim, A-Young</mods:namePart>
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<mods:namePart id="7E82B9B3594080DD3CEDFE95996D282E">Byun, Hye-Woo</mods:namePart>
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<mods:namePart id="BBFEEAC044B4289C422EEADFC590DA2F">Kim, Heung-Chul</mods:namePart>
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<subSubSection id="C355654BBB02FFBEF0C9F99BFC40F023" box="[151,984,1593,1620]" pageId="5" pageNumber="286" type="nomenclature">
<paragraph id="8BF036C0BB02FFBEF0C9F99BFC40F023" blockId="5.[151,984,1593,1655]" box="[151,984,1593,1620]" pageId="5" pageNumber="286">
<heading id="D0B881ACBB02FFBEF0C9F99BFC40F023" bold="true" box="[151,984,1593,1620]" fontSize="11" level="1" pageId="5" pageNumber="286" reason="1">
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<taxonomicName id="4C4F4D43BB02FFBEF0C9F99BFCF6F023" ID-CoL="8GZPS" authority="Podenas &amp; Podeniene" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[151,878,1593,1620]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB02FFBEF0C9F99BFEC6F024" bold="true" box="[151,350,1593,1620]" italics="true" pageId="5" pageNumber="286">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB02FFBEF131F99BFD8DF024" bold="true" box="[367,533,1593,1620]" italics="true" pageId="5" pageNumber="286">Microlimonia</emphasis>
) 
<emphasis id="B93BEAD2BB02FFBEF279F99BFDCDF023" bold="true" box="[551,597,1593,1619]" italics="true" pageId="5" pageNumber="286">jeju</emphasis>
Podenas &amp; Podeniene
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, 
<taxonomicNameLabel id="A20857A9BB02FFBEF324F99BFC40F023" box="[890,984,1593,1619]" pageId="5" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C355654BBB02FFBEF0C9F9FFFED2F007" box="[151,330,1628,1655]" pageId="5" pageNumber="286" type="description">
<paragraph id="8BF036C0BB02FFBEF0C9F9FFFED2F007" blockId="5.[151,984,1593,1655]" box="[151,330,1628,1655]" pageId="5" pageNumber="286">
(
<figureCitation id="13742A45BB02FFBEF0C1F9FFFF77F007" box="[159,239,1628,1655]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1932,1957]" captionTargetBox="[285,1294,185,1900]" captionTargetId="figure@2.[276,1312,174,1913]" captionTargetPageId="2" captionText="FIGURES 1–6. Wings of Korean Atypophthalmus. 1: A. (Atypophthalmus) crinitus; 2: A. (A.) kleini Podenas, sp. nov.; 3: A. (A.) stylacanthus; 4: A. (Microlimonia) inelegans; 5: A. (M.) jeju Podenas, Podeniene, sp. nov.; 6: A. (M.) machidai. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663451" httpUri="https://zenodo.org/record/3663451/files/figure.png" pageId="5" pageNumber="286">Figs. 5</figureCitation>
, 
<figureCitation id="13742A45BB02FFBEF0A5F9FFFEDAF007" box="[251,322,1629,1655]" captionStart-0="FIGURES 14–17" captionStart-1="FIGURES 18–27" captionStart-2="FIGURES 28–32" captionStartId-0="7.[151,264,1763,1788]" captionStartId-1="9.[151,264,1907,1932]" captionStartId-2="10.[151,264,1925,1950]" captionTargetBox-0="[171,1425,350,1735]" captionTargetBox-1="[184,1413,180,1883]" captionTargetBox-2="[161,1428,703,1897]" captionTargetId-0="figure@7.[151,1435,337,1744]" captionTargetId-1="figure@9.[173,1415,174,1887]" captionTargetId-2="figure@10.[151,1436,697,1905]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 14–17. Atypophthalmus (Microlimonia) jeju Podenas, Podeniene, sp. nov. 14: male, holotype, general view; 15: male antenna; 16: male genitalia, dorsal view; 17: ovipositor, lateral view." captionText-1="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." captionText-2="FIGURES 28–32. Female pupa of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3663457" figureDoi-1="http://doi.org/10.5281/zenodo.3663459" figureDoi-2="http://doi.org/10.5281/zenodo.3663463" httpUri-0="https://zenodo.org/record/3663457/files/figure.png" httpUri-1="https://zenodo.org/record/3663459/files/figure.png" httpUri-2="https://zenodo.org/record/3663463/files/figure.png" pageId="5" pageNumber="286">14–32</figureCitation>
)
</paragraph>
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<subSubSection id="C355654BBB02FFBDF0C9F906FB5AF76F" lastPageId="6" lastPageNumber="287" pageId="5" pageNumber="286" type="diagnosis">
<paragraph id="8BF036C0BB02FFBDF0C9F906FB5AF76F" blockId="5.[151,1437,1700,2015]" lastBlockId="6.[151,1437,153,2015]" lastPageId="6" lastPageNumber="287" pageId="5" pageNumber="286">
<emphasis id="B93BEAD2BB02FFBEF0C9F906FE8AF0CF" bold="true" box="[151,274,1700,1727]" pageId="5" pageNumber="286">Diagnosis.</emphasis>
A small to medium-sized crane fly, with body length reaching approximately 
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in the male and nearly 
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in the female. Antennae 15-segmented. Thorax pale yellow with distinct dark brown stripe along the pleuron, abdomen brownish gray dorsally and somewhat lighter ventrally. Male: Gonocoxite of male terminalia with large elongate ventro-mesial lobe with a straight distal end, and smaller rounded lobe at its base. Gonostylus egg-shaped with large, arched sclerotized rostral prolongation and smaller rounded subbasal tubercle reaching to the middle of the gonostylus. Female: Cercus of ovipositor short and narrow, distal part slightly turned upwards. Hypovalva long and straight, nearly reaching tip of cercus, with distinct black spot at base. Larva: Dorsoventrally depressed and slightly reduced head capsule, dorsal suture reaching only one third of the head capsule length. Labrum elongate-oval with dentate anterior part, mandible with six teeth, hypostoma with twelve teeth. Spiracular field with five short and blunt lobes, lateral lobe twice as wide as long, without sclerites, ventral lobe as long as width at the base and bearing a dark sclerotized, elongate and angulate sclerite. Pupa: Female pupa with flattened triangular pronotal horns, bases of which are very close to each other, apices separated. Legs reaching almost to the middle of fourth abdominal segment. Sheath of cerci and sheath of valvae are similar in length.
</paragraph>
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<subSubSection id="C355654BBB01FFBDF099FE8AFE8CF413" pageId="6" pageNumber="287" type="materials_examined">
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<emphasis id="B93BEAD2BB01FFBDF099FE8AFE2BF733" bold="true" box="[199,435,296,323]" pageId="6" pageNumber="287">Material examined:</emphasis>
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specimens 
<specimenCount id="9D49FD49BB01FFBDF22DFE8BFD42F733" box="[627,730,297,323]" pageId="6" pageNumber="287" type="male">24 males</specimenCount>
, 
<specimenCount id="9D49FD49BB01FFBDF2B8FE8BFCCDF733" box="[742,853,297,323]" pageId="6" pageNumber="287" type="female">7 females</specimenCount>
, 
<specimenCount id="9D49FD49BB01FFBDF33EFE8BFC24F733" box="[864,956,297,323]" pageId="6" pageNumber="287" type="larva">4 larvae</specimenCount>
, 
<specimenCount id="9D49FD49BB01FFBDF399FE8BFBB0F733" box="[967,1064,297,323]" pageId="6" pageNumber="287" type="female">1 female</specimenCount>
pupa.
</paragraph>
<paragraph id="8BF036C0BB01FFBDF099FEEFFF59F7DF" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="287">
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<typeStatus id="54F48862BB01FFBDF099FEEFFEA0F717" box="[199,312,333,359]" pageId="6" pageNumber="287" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (pinned) (
<figureCitation id="13742A45BB01FFBDF1A4FEEFFDD7F717" box="[506,591,333,359]" captionStart="FIGURES 14–17" captionStartId="7.[151,264,1763,1788]" captionTargetBox="[171,1425,350,1735]" captionTargetId="figure@7.[151,1435,337,1744]" captionTargetPageId="7" captionText="FIGURES 14–17. Atypophthalmus (Microlimonia) jeju Podenas, Podeniene, sp. nov. 14: male, holotype, general view; 15: male antenna; 16: male genitalia, dorsal view; 17: ovipositor, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3663457" httpUri="https://zenodo.org/record/3663457/files/figure.png" pageId="6" pageNumber="287">Fig. 14</figureCitation>
), S. 
<collectingCountry id="F3587650BB01FFBDF2D7FEEFFD48F717" box="[649,720,333,359]" name="South Korea" pageId="6" pageNumber="287">Korea</collectingCountry>
, 
<collectingRegion id="498BF822BB01FFBDF280FEEFFCA8F717" box="[734,816,333,359]" country="South Korea" name="Jeju" pageId="6" pageNumber="287">Jeju-do</collectingRegion>
(do = 
<collectingCountry id="F3587650BB01FFBDF3DBFEEFFC56F718" box="[901,974,333,360]" name="Iceland" pageId="6" pageNumber="287">Island</collectingCountry>
), 
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wetland, 
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, 
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, alt. 
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<collectingDate id="EFB5E9E8BB01FFBDF274FED2FD30F7FB" box="[554,680,368,395]" pageId="6" pageNumber="287" value="2017-05-23">2017.05.23</collectingDate>
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(1), coll. 
<collectorName id="26BA5316BB01FFBDF348FED3FC0CF7FB" box="[790,916,369,395]" pageId="6" pageNumber="287">S. Podenas</collectorName>
, evergreen broad-leaved forest, 
<collectingMethod id="520E4ED7BB01FFBDF4A2FED3FF25F7DF" pageId="6" pageNumber="287">entomological net</collectingMethod>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF036C0BB01FFBDF099FE1BFE8CF413" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="287">
<emphasis id="B93BEAD2BB01FFBDF099FE1BFED0F7A3" bold="true" box="[199,328,441,467]" pageId="6" pageNumber="287">
<typeStatus id="54F48862BB01FFBDF099FE1BFEDBF7A3" box="[199,323,441,467]" pageId="6" pageNumber="287" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D49FD49BB01FFBDF110FE1BFE2CF7A3" box="[334,436,441,467]" pageId="6" pageNumber="287" type="male">18 males</specimenCount>
, 
<specimenCount id="9D49FD49BB01FFBDF1E0FE1BFDB1F7A3" box="[446,553,441,467]" pageId="6" pageNumber="287" type="female">9 females</specimenCount>
(pinned), 
<specimenCount id="9D49FD49BB01FFBDF2C4FE1BFC98F7A3" box="[666,768,441,467]" pageId="6" pageNumber="287" type="male">22 males</specimenCount>
, 
<specimenCount id="9D49FD49BB01FFBDF354FE1BFCEDF7A3" box="[778,885,441,467]" pageId="6" pageNumber="287" type="female">6 females</specimenCount>
(in EtOH), 
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<specimenCount id="9D49FD49BB01FFBDF3A9FE1BFBD7F7A3" box="[1015,1103,441,467]" pageId="6" pageNumber="287" type="larva">3 larvae</specimenCount>
and 
<specimenCount id="9D49FD49BB01FFBDF4DAFE1BFB7BF7A3" box="[1156,1251,441,467]" pageId="6" pageNumber="287" type="female">1 female</specimenCount>
pupa (in EtOH), 1 larva (slide-mounted), topotypic, coll. 
<collectorName id="26BA5316BB01FFBDF23DFE7FFD7AF787" box="[611,738,477,503]" pageId="6" pageNumber="287">S. Podenas</collectorName>
, 
<collectionCode id="ED5EAE05BB01FFBDF2B1FE7FFC9BF787" box="[751,771,477,503]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="287" type="Museum">V</collectionCode>
. 
<collectingMunicipality id="6B94ACBABB01FFBDF351FE7FFC1FF787" box="[783,903,477,503]" pageId="6" pageNumber="287">Podeniene</collectingMunicipality>
, four of these males preserved at the 
<location id="8E90601BBB01FFBDF56FFE7FFEEBF46B" LSID="urn:lsid:plazi:treatment:03E687D6BB02FFB0F0C9F99BFE97F007:8E90601BBB01FFBDF56FFE7FFEEBF46B" country="United States of America" municipality="Podeniene" name="Academy of Natural Sciences" pageId="6" pageNumber="287">Academy of Natural Sciences</location>
of Drexel University, 
<collectorName id="26BA5316BB01FFBDF230FDA3FCB4F46B" box="[622,812,513,539]" pageId="6" pageNumber="287">Philadelphia, PA</collectorName>
, 
<collectingCountry id="F3587650BB01FFBDF369FDA3FCF0F46B" box="[823,872,513,539]" name="United States of America" pageId="6" pageNumber="287">U.S.</collectingCountry>
</materialsCitation>
A; 
<specimenCount id="9D49FD49BB01FFBDF3D2FDA3FC4FF46B" box="[908,983,513,539]" pageId="6" pageNumber="287" type="male">1 male</specimenCount>
(pinned), 
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<specimenCount id="9D49FD49BB01FFBDF412FDA3FB35F46B" box="[1100,1197,513,539]" pageId="6" pageNumber="287" type="female">1 female</specimenCount>
(in EtOH), S. 
<collectingCountry id="F3587650BB01FFBDF50FFDA3FA00F46B" box="[1361,1432,513,539]" name="South Korea" pageId="6" pageNumber="287">Korea</collectingCountry>
, 
<collectingRegion id="498BF822BB01FFBDF0C9FD87FF74F44F" box="[151,236,549,575]" country="South Korea" name="Jeju" pageId="6" pageNumber="287">Jeju-do</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="3B273C9DBB01FFBDF0A6FD87FE88F413" ID-GBIF-Occurrence="2565010305" collectingDate="2017-05-24" collectorName="S. Podenas" country="South Korea" elevation="577" latitude="33.43222" location="Hallasan National Forest" longLatPrecision="1" longitude="126.59776" pageId="6" pageNumber="287" specimenCount="1" stateProvince="Jeju" typeStatus="paratype">
<location id="8E90601BBB01FFBDF0A6FD87FD89F44F" LSID="urn:lsid:plazi:treatment:03E687D6BB02FFB0F0C9F99BFE97F007:8E90601BBB01FFBDF0A6FD87FD89F44F" box="[248,529,549,575]" country="South Korea" latitude="33.43222" longLatPrecision="1" longitude="126.59776" name="Hallasan National Forest" pageId="6" pageNumber="287" stateProvince="Jeju">Hallasan National Forest</location>
, 
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, 
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, alt. 
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<elevation id="0062D1F3BB01FFBDF32BFD86FC27F44F" box="[885,959,548,575]" metricMagnitude="2" metricUnit="m" metricValue="5.77" pageId="6" pageNumber="287" unit="m" value="577.0">577 m</elevation>
</quantity>
, 
<date id="FFF11000BB01FFBDF394FD86FBD0F44F" box="[970,1096,548,575]" pageId="6" pageNumber="287" value="2017-05-24">
<collectingDate id="EFB5E9E8BB01FFBDF394FD86FBD0F44F" box="[970,1096,548,575]" pageId="6" pageNumber="287" value="2017-05-24">2017.05.24</collectingDate>
</date>
(1), coll. 
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, 
<collectingMethod id="520E4ED7BB01FFBDF561FD87FE88F413" pageId="6" pageNumber="287">entomological net</collectingMethod>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C355654BBB01FFB3F099FDCEFC8FF0EB" lastPageId="8" lastPageNumber="289" pageId="6" pageNumber="287" type="description">
<paragraph id="8BF036C0BB01FFBDF099FDCEFD6FF4F8" blockId="6.[151,1437,153,2015]" box="[199,759,620,648]" pageId="6" pageNumber="287">
<emphasis id="B93BEAD2BB01FFBDF099FDCEFEC1F4F7" bold="true" box="[199,345,620,647]" pageId="6" pageNumber="287">Description.</emphasis>
Adult, male (N=42), female (N=16).
</paragraph>
<paragraph id="8BF036C0BB01FFBDF099FD33FF52F4BF" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="287">
General body color yellow with gray to brown abdomen. Body length of male 3.8–5.0 mm, of female 
<quantity id="4CB79B25BB01FFBDF516FD32FF5EF4BF" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.7" metricValueMin="3.4" pageId="6" pageNumber="287" unit="mm" value="4.55" valueMax="5.7" valueMin="3.4">3.4–5.7 mm</quantity>
.
</paragraph>
<paragraph id="8BF036C0BB01FFBDF099FD7BFBFCF5BB" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="287">
<emphasis id="B93BEAD2BB01FFBDF099FD7BFE93F483" box="[199,267,729,755]" italics="true" pageId="6" pageNumber="287">Head.</emphasis>
Black dusted with grayish pruinosity. Eyes large, holoptic, nearly reaching each other dorsally. Antennae (
<figureCitation id="13742A45BB01FFBDF091FD5FFEBAF567" box="[207,290,764,791]" captionStart="FIGURES 14–17" captionStartId="7.[151,264,1763,1788]" captionTargetBox="[171,1425,350,1735]" captionTargetId="figure@7.[151,1435,337,1744]" captionTargetPageId="7" captionText="FIGURES 14–17. Atypophthalmus (Microlimonia) jeju Podenas, Podeniene, sp. nov. 14: male, holotype, general view; 15: male antenna; 16: male genitalia, dorsal view; 17: ovipositor, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3663457" httpUri="https://zenodo.org/record/3663457/files/figure.png" pageId="6" pageNumber="287">Fig. 15</figureCitation>
) entirely brown, 
<quantity id="4CB79B25BB01FFBDF1B9FD5FFDE8F567" box="[487,624,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="6" pageNumber="287" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
long in male, 
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in female, nearly reaching wing base if bent backwards. Scape elongate, approximately 1.5 times as long as wide, slightly widened distally. Pedicel egg-shaped, nearly as long as scape. Flagellum 13-segmented, flagellomeres oval with small apical pedicels, twelfth flagellomere elongate, apical flagellomere very small, distinctly shorter than the penultimate flagellomere. Longest verticils nearly as long as respective segments. Flagellomeres covered with sparse whitish pubescence. Rostrum yellow, slightly brownish dorsally at base. Palpus yellow, distal part of apical segment grayish.
</paragraph>
<paragraph id="8BF036C0BB01FFBDF099FC77FEECF177" blockId="6.[151,1437,153,2015]" pageId="6" pageNumber="287">
<emphasis id="B93BEAD2BB01FFBDF099FC77FEB8F59F" box="[199,288,981,1007]" italics="true" pageId="6" pageNumber="287">Thorax.</emphasis>
Generally yellow. Cervical sclerites dorsally grayish, laterally dark brown. Pronotum yellowish gray with dark brown lateral margin. Mesonotal prescutum grayish yellow, stripes absent, darkening, intensity of which depends on specimen, at the middle before posterior margin of sclerite. Scutal lobe with large drop-shaped brownish gray spot, widely yellow at frontal margin and pale yellow postero-lateral corner. Both lobes widely separated by pale yellow color. Scutellum grayish, widely yellow laterally. Mediotergite light gray frontally, dark gray posteriorly, with pale yellow frontal margin and narrow dark line along middle at posterior half. Pleuron pale yellow dorsally, whitish ventrally with distinct dark brown stripe extending from cervical sclerites through the base of the halter to the basal tergite of abdomen. Few setae on the katepisternum. Wing (
<figureCitation id="13742A45BB01FFBDF3A1FB73FBD9F29B" box="[1023,1089,1232,1259]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1932,1957]" captionTargetBox="[285,1294,185,1900]" captionTargetId="figure@2.[276,1312,174,1913]" captionTargetPageId="2" captionText="FIGURES 1–6. Wings of Korean Atypophthalmus. 1: A. (Atypophthalmus) crinitus; 2: A. (A.) kleini Podenas, sp. nov.; 3: A. (A.) stylacanthus; 4: A. (Microlimonia) inelegans; 5: A. (M.) jeju Podenas, Podeniene, sp. nov.; 6: A. (M.) machidai. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663451" httpUri="https://zenodo.org/record/3663451/files/figure.png" pageId="6" pageNumber="287">Fig. 5</figureCitation>
) subhyaline, unpatterned, with drop-shaped brownish stigma. Length 4.7–6.0 mm (male), 
<quantity id="4CB79B25BB01FFBDF36BFB57FC25F37F" box="[821,957,1269,1295]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="6.3" metricValueMin="4.6" pageId="6" pageNumber="287" unit="mm" value="5.449999999999999" valueMax="6.3" valueMin="4.6">4.6–6.3 mm</quantity>
(female). Veins brown, pale at wing base. Venation: 
<emphasis id="B93BEAD2BB01FFBDF157FABBFEBBF343" box="[265,291,1305,1331]" italics="true" pageId="6" pageNumber="287">Sc</emphasis>
medium-long, 
<emphasis id="B93BEAD2BB01FFBDF190FABBFE68F348" box="[462,496,1305,1336]" italics="true" pageId="6" pageNumber="287">
Sc 
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</emphasis>
reaching to about one third of 
<emphasis id="B93BEAD2BB01FFBDF31BFAB8FCF9F343" box="[837,865,1306,1331]" italics="true" pageId="6" pageNumber="287">Rs</emphasis>
length. 
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Sc 
<subScript id="17CB3485BB01FFBDF388FA88FC46F348" attach="left" box="[982,990,1322,1336]" fontSize="6" pageId="6" pageNumber="287">2</subScript>
</emphasis>
slightly before apex of 
<emphasis id="B93BEAD2BB01FFBDF4BBFABBFA9FF348" box="[1253,1287,1305,1336]" italics="true" pageId="6" pageNumber="287">
Sc 
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</emphasis>
. 
<emphasis id="B93BEAD2BB01FFBDF54BFAB8FAA9F343" box="[1301,1329,1306,1331]" italics="true" pageId="6" pageNumber="287">Rs</emphasis>
mediumlong, slightly arched, or nearly straight, alternately angulate and short-spurred; variations occur on each specimen. Free end of 
<emphasis id="B93BEAD2BB01FFBDF17DFAC0FEA5F3F0" box="[291,317,1378,1408]" italics="true" pageId="6" pageNumber="287">
R 
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</emphasis>
short, transverse, very light and indistinct. 
<emphasis id="B93BEAD2BB01FFBDF372FAC0FCDDF3F0" box="[812,837,1378,1408]" italics="true" pageId="6" pageNumber="287">
R 
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</emphasis>
some distance beyond tip of 
<emphasis id="B93BEAD2BB01FFBDF4C7FAC0FB2AF3F0" box="[1177,1202,1378,1408]" italics="true" pageId="6" pageNumber="287">
R 
<subScript id="17CB3485BB01FFBDF4F4FAD0FB2AF3F0" attach="left" box="[1194,1202,1394,1408]" fontSize="6" pageId="6" pageNumber="287">1</subScript>
</emphasis>
. 
<emphasis id="B93BEAD2BB01FFBDF49CFAC0FB43F3F0" box="[1218,1243,1378,1408]" italics="true" pageId="6" pageNumber="287">
R 
<subScript id="17CB3485BB01FFBDF48DFAD0FB43F3F0" attach="left" box="[1235,1243,1394,1408]" fontSize="6" pageId="6" pageNumber="287">3</subScript>
</emphasis>
and 
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R 
<subScript id="17CB3485BB01FFBDF578FAD0FADAF3F0" attach="left" box="[1318,1346,1394,1408]" fontSize="6" pageId="6" pageNumber="287">4+5</subScript>
</emphasis>
arched, parallel to each other. Specimens with short-spurred base of 
<emphasis id="B93BEAD2BB01FFBDF360FA24FCC2F3EF" box="[830,858,1414,1439]" italics="true" pageId="6" pageNumber="287">Rs</emphasis>
usually have short-spurred base of 
<emphasis id="B93BEAD2BB01FFBDF4B4FA24FA8FF3D4" box="[1258,1303,1414,1444]" italics="true" pageId="6" pageNumber="287">
R 
<subScript id="17CB3485BB01FFBDF4A5FA34FA8FF3D4" attach="left" box="[1275,1303,1430,1444]" fontSize="6" pageId="6" pageNumber="287">4+5</subScript>
</emphasis>
. Discal cell missing by atrophy of basal part of 
<emphasis id="B93BEAD2BB01FFBDF27BFA08FDDCF3B8" box="[549,580,1450,1480]" italics="true" pageId="6" pageNumber="287">
M 
<subScript id="17CB3485BB01FFBDF262FA18FDDCF3B8" attach="left" box="[572,580,1466,1480]" fontSize="6" pageId="6" pageNumber="287">3</subScript>
</emphasis>
. Cell 
<emphasis id="B93BEAD2BB01FFBDF2D6FA08FD2FF3B8" box="[648,695,1450,1480]" italics="true" pageId="6" pageNumber="287">
m 
<subScript id="17CB3485BB01FFBDF2C2FA18FD2FF3B8" attach="left" box="[668,695,1466,1480]" fontSize="6" pageId="6" pageNumber="287">1+2</subScript>
</emphasis>
short-petiolate, with stem being nearly three times as short as cell itself. Basal deflection of 
<emphasis id="B93BEAD2BB01FFBDF1E9FA6FFE69F39C" box="[439,497,1485,1516]" italics="true" pageId="6" pageNumber="287">
CuA 
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</emphasis>
close to branching point of 
<emphasis id="B93BEAD2BB01FFBDF374FA6CFCD9F397" box="[810,833,1486,1511]" italics="true" pageId="6" pageNumber="287">M</emphasis>
, sometimes slightly before or slightly beyond it. Both anal veins nearly straight, diverging. Anal lobe wide. Length of halter 0.7–1.0 mm (male) and 
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(female). Halter uniformly grayish, with distinctly pale base. Coxae pale, just frontal face of fore coxa distinctly darkened depending on specimen, distal margin narrowly dark brown. Trochanters pale with narrowly blackish distal margin. Legs uniformly brownish except yellowish base of femur. Male femur I: 4.0– 
<quantity id="4CB79B25BB01FFBDF419F9FFFB06F007" box="[1095,1182,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="287" unit="mm" value="4.3">4.3 mm</quantity>
long, II: 
<quantity id="4CB79B25BB01FFBDF552F9FEFA00F007" box="[1292,1432,1628,1655]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="6" pageNumber="287" unit="mm" value="4.55" valueMax="4.6" valueMin="4.5">4.5–4.6 mm</quantity>
, III: 
<quantity id="4CB79B25BB01FFBDF09DF922FED5F0EB" box="[195,333,1664,1691]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="6" pageNumber="287" unit="mm" value="4.8" valueMax="5.1" valueMin="4.5">4.5–5.1 mm</quantity>
, tibiae I: 
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, II: 
<quantity id="4CB79B25BB01FFBDF22EF922FD62F0EB" box="[624,762,1664,1691]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.1" metricValueMin="4.7" pageId="6" pageNumber="287" unit="mm" value="4.9" valueMax="5.1" valueMin="4.7">4.7–5.1 mm</quantity>
, III: 
<quantity id="4CB79B25BB01FFBDF36CF922FC24F0EB" box="[818,956,1664,1691]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.5" metricValueMin="4.8" pageId="6" pageNumber="287" unit="mm" value="5.15" valueMax="5.5" valueMin="4.8">4.8–5.5 mm</quantity>
, tarsus I: 
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, II: 4.3–5.0 mm, III: 
<quantity id="4CB79B25BB01FFBDF0C9F906FEBAF0CF" box="[151,290,1700,1727]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.5" metricValueMin="3.8" pageId="6" pageNumber="287" unit="mm" value="4.15" valueMax="4.5" valueMin="3.8">3.8–4.5 mm</quantity>
. Female femur I: 
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long, II: 
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, III: 
<quantity id="4CB79B25BB01FFBDF3C2F906FBBEF0CF" box="[924,1062,1700,1727]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.6" metricValueMin="4.2" pageId="6" pageNumber="287" unit="mm" value="4.9" valueMax="5.6" valueMin="4.2">4.2–5.6 mm</quantity>
, tibiae I: 
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, II: 
<quantity id="4CB79B25BB01FFBDF516F906FF5DF093" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.4" metricValueMin="4.2" pageId="6" pageNumber="287" unit="mm" value="4.8" valueMax="5.4" valueMin="4.2">4.2–5.4 mm</quantity>
, III: 
<quantity id="4CB79B25BB01FFBDF0A1F96AFE12F093" box="[255,394,1736,1763]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.1" metricValueMin="3.4" pageId="6" pageNumber="287" unit="mm" value="4.25" valueMax="5.1" valueMin="3.4">3.4–5.1 mm</quantity>
, tarsus I: 4.2–5.0 mm, II: 
<quantity id="4CB79B25BB01FFBDF2E9F96AFCDAF093" box="[695,834,1736,1763]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="6" pageNumber="287" unit="mm" value="4.6" valueMax="5.1" valueMin="4.1">4.1–5.1 mm</quantity>
, III: 3.6–4.0 mm. Claw with subbasal spine in male, spineless in female.
</paragraph>
<paragraph id="8BF036C0BB01FFBCF099F8B3FAA4F76F" blockId="6.[151,1437,153,2015]" lastBlockId="7.[151,1437,153,323]" lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287">
<emphasis id="B93BEAD2BB01FFBDF099F8B3FEA1F15B" box="[199,313,1809,1835]" italics="true" pageId="6" pageNumber="287">Abdomen.</emphasis>
Abdominal tergites brownish gray. Membrane between tergites and sternites pale with distinct dark spots where separate segments meet. Basal sternite pale yellow, succeeding three sternites pale at base and grayish distally, remaining sternites grayish, lighter than tergites. Male terminalia (
<figureCitation id="13742A45BB01FFBDF3BBF8FBFBA3F104" box="[997,1083,1881,1908]" captionStart="FIGURES 14–17" captionStartId="7.[151,264,1763,1788]" captionTargetBox="[171,1425,350,1735]" captionTargetId="figure@7.[151,1435,337,1744]" captionTargetPageId="7" captionText="FIGURES 14–17. Atypophthalmus (Microlimonia) jeju Podenas, Podeniene, sp. nov. 14: male, holotype, general view; 15: male antenna; 16: male genitalia, dorsal view; 17: ovipositor, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3663457" httpUri="https://zenodo.org/record/3663457/files/figure.png" pageId="6" pageNumber="287">Fig. 16</figureCitation>
): yellowish dark gray with distinctly pale yellow aedeagus. Posterior margin of ninth tergite widely rounded with indistinct emargination medially. Gonocoxite elongate with large elongate ventro-mesial lobe, distal apex straight rather than rounded, smaller round- ed lobe at its base. Gonostylus egg-shaped with large, arched, sclerotized rostral prolongation and smaller rounded subbasal bump reaching to the middle of gonostylus. Paramere with wide blackened hook-shaped prolongation of mesial margin. Penis wide and long, reaching beyond tip of paramere, apex shallowly bifid. Female: Ovipositor yellow (
<figureCitation id="13742A45BB00FFBCF0ADFF43FEDFF68B" box="[243,327,225,251]" captionStart="FIGURES 14–17" captionStartId="7.[151,264,1763,1788]" captionTargetBox="[171,1425,350,1735]" captionTargetId="figure@7.[151,1435,337,1744]" captionTargetPageId="7" captionText="FIGURES 14–17. Atypophthalmus (Microlimonia) jeju Podenas, Podeniene, sp. nov. 14: male, holotype, general view; 15: male antenna; 16: male genitalia, dorsal view; 17: ovipositor, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3663457" httpUri="https://zenodo.org/record/3663457/files/figure.png" pageId="7" pageNumber="288">Fig. 17</figureCitation>
). Tenth tergite slightly darkened along dorso-posterior margin. Cercus short and narrow, distal part slightly raised. Hypovalva long and straight, nearly reaching tip of cercus, with distinct black spot at base.
</paragraph>
<paragraph id="8BF036C0BB00FFBCF099FE8BFC22F733" blockId="7.[151,1437,153,323]" box="[199,954,296,323]" pageId="7" pageNumber="288">
<emphasis id="B93BEAD2BB00FFBCF099FE8BFE88F733" bold="true" box="[199,272,297,323]" pageId="7" pageNumber="288">Larva</emphasis>
(N=4). Length 
<quantity id="4CB79B25BB00FFBCF19EFE8BFDD1F733" box="[448,585,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" metricValueMax="9.8" metricValueMin="8.8" pageId="7" pageNumber="288" unit="mm" value="9.3" valueMax="9.8" valueMin="8.8">8.8–9.8 mm</quantity>
, width 
<quantity id="4CB79B25BB00FFBCF2C2FE8AFCBEF733" box="[668,806,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="7" pageNumber="288" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
. Body white.
</paragraph>
<caption id="DF306648BB00FFBCF0C9F941FC1EF150" ID-DOI="http://doi.org/10.5281/zenodo.3663457" ID-Zenodo-Dep="3663457" httpUri="https://zenodo.org/record/3663457/files/figure.png" pageId="7" pageNumber="288" startId="7.[151,264,1763,1788]" targetBox="[171,1425,350,1735]" targetPageId="7">
<paragraph id="8BF036C0BB00FFBCF0C9F941FC1EF150" blockId="7.[151,1436,1763,1824]" pageId="7" pageNumber="288">
<emphasis id="B93BEAD2BB00FFBCF0C9F941FECEF08C" bold="true" box="[151,342,1763,1788]" pageId="7" pageNumber="288">FIGURES 14–17.</emphasis>
<taxonomicName id="4C4F4D43BB00FFBCF101F946FC36F08C" authority="Podenas, Podeniene" authorityName="Podenas, Podeniene" authorityYear="2020" box="[351,942,1763,1788]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB00FFBCF101F946FD9CF08C" box="[351,516,1764,1788]" italics="true" pageId="7" pageNumber="288">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB00FFBCF24BF946FD07F08C" box="[533,671,1764,1788]" italics="true" pageId="7" pageNumber="288">Microlimonia</emphasis>
) 
<emphasis id="B93BEAD2BB00FFBCF2EEF947FD4EF08C" box="[688,726,1765,1788]" italics="true" pageId="7" pageNumber="288">jeju</emphasis>
Podenas, Podeniene
</taxonomicName>
, 
<emphasis id="B93BEAD2BB00FFBCF3E5F946FB96F08C" bold="true" box="[955,1038,1764,1788]" pageId="7" pageNumber="288">
<taxonomicNameLabel id="A20857A9BB00FFBCF3E5F946FB96F08C" box="[955,1038,1764,1788]" pageId="7" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
14: male, holotype, general view; 15: male antenna; 16: male genitalia, dorsal view; 17: ovipositor, lateral view.
</paragraph>
</caption>
<paragraph id="8BF036C0BB00FFB3F099F8ECFBDDF4DB" blockId="7.[151,1437,1870,2041]" lastBlockId="8.[151,1437,153,2015]" lastPageId="8" lastPageNumber="289" pageId="7" pageNumber="288">
<emphasis id="B93BEAD2BB00FFBCF099F8ECFE93F118" box="[199,267,1870,1896]" italics="true" pageId="7" pageNumber="288">Head.</emphasis>
Head capsule 1.0 mm long and 
<quantity id="4CB79B25BB00FFBCF2DEF8ECFD40F119" box="[640,728,1870,1897]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="288" unit="mm" value="0.6">0.6 mm</quantity>
wide, oval in shape, depressed dorsoventrally and slightly reduced, dorsal suture reaches only one third of head capsule length (
<figureCitation id="13742A45BB00FFBCF3CCF8D0FC47F1FD" box="[914,991,1906,1933]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="7" pageNumber="288">Fig.18</figureCitation>
). Labrum elongate-oval with numerous sensory structures: short seta medially of anterior part and three short setae on antero-lateral part, sensory circle area with two sensory papillae inside and two separate sensory pits located near the postero-lateral side. Anterior part of labrum dentate (
<figureCitation id="13742A45BB00FFBCF131F87CFE5CF188" box="[367,452,2014,2040]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="7" pageNumber="288">Fig. 19</figureCitation>
). Clypeus distinctly divided from labrum, consisting of sclerotized narrow postclypeus and membranous wide preclypeus. A long seta located near the antero-lateral side of preclypeus and sensory pit below it. A long seta and sensory pit located on postero-lateral part of preclypeus. Frons separated from clypeus, fused with internolateralia, wide, and T-shaped. A single sensory pit near the base of antenna and four sensory pits along the lateral margin of the widened part of frons. All five sensory pits arranged along straight axis. Caudal end of frons rounded (
<figureCitation id="13742A45BB0FFFB3F13FFE8BFE2CF733" box="[353,436,297,323]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 18</figureCitation>
). Basal segment of antenna cylindrical, less than 1.5 times as long as wide, with few very short sensory structures apically. Apical segment short, button-shaped. Sensory pit located at the middle of basal segment´s length (
<figureCitation id="13742A45BB0FFFB3F13DFED3FE2EF7FB" box="[355,438,369,395]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 20</figureCitation>
). Mandible conus-shaped, with two large teeth similar in shape and size apically (
<figureCitation id="13742A45BB0FFFB3F563FED3FA16F7FB" box="[1341,1422,369,395]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 21</figureCitation>
), three smaller triangular teeth on ventral side (basal tooth is the largest) and a well-developed sharp tooth on dorsal side. Maxilla well developed (
<figureCitation id="13742A45BB0FFFB3F1ABFE1BFDD3F7A3" box="[501,587,441,467]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 22</figureCitation>
), stipes and galea large and similar in size. Stipes: two-thirds sclerotized, apical papilla button shaped with sensory structures on the apex, a small papilla (on outer margin) and sensory pit (on inner margin) below. Galea: basal part sclerotized, sensory papilla and sensory pit on inner margin of ventral sclerite, apical part covered with short setae, large sensory papilla (with small sensory structures inside) and small sensory papilla on apex. Cardo large, wedge shaped, long seta located near inner margin and short seta and sensory pit near outer margin of sclerite. Hypopharynx and prementum dentate (
<figureCitation id="13742A45BB0FFFB3F397FDCFFB8EF4F8" box="[969,1046,621,648]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig.23</figureCitation>
). Hypostoma bearing 12 teeth. All (except the middle and the most lateral teeth) are similar in shape and size (
<figureCitation id="13742A45BB0FFFB3F3BAFD33FBA1F4DB" box="[996,1081,657,683]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FD17FAE0F4BF" blockId="8.[151,1437,153,2015]" box="[199,1400,693,719]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FD17FEB8F4BF" box="[199,288,693,719]" italics="true" pageId="8" pageNumber="289">Thorax.</emphasis>
All thoracic segments wider than long. Second and third segments each with ventral creeping welt.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FD7BFC84F52F" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FD7BFEA1F483" box="[199,313,729,755]" italics="true" pageId="8" pageNumber="289">Abdomen.</emphasis>
First, seventh and last abdominal segment less than 1.5 times as long as wide. Second abdominal segment more than twice as long as wide. Abdominal segments III–VI more than three times as long as wide. First segment with ventral creeping welt only, segments II–VII with dorsal and ventral creeping welts (
<figureCitation id="13742A45BB0FFFB3F48EFC83FABBF54B" box="[1232,1315,800,827]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 25</figureCitation>
). Creeping welts with brown spines, arranged into longitudinal rows.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FCCBFC8FF247" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FCCBFEC6F5F3" box="[199,350,873,899]" italics="true" pageId="8" pageNumber="289">Anal division</emphasis>
. Spiracular field very small, with five short blunt lobes, dorsal and lateral part inclined inward (
<figureCitation id="13742A45BB0FFFB3F0FEFC2FFF6BF5D8" box="[160,243,909,936]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 26</figureCitation>
). Dorsal lobe vestigial with a pair of long setae at apex. Lateral lobe twice as wide as long, not covered with sclerites, two long and one short seta located at the margin. Ventral lobe as long as width at the base and bears a dark sclerotized, elongate and angulate sclerite. One short and three long setae located at the apex of each lobe. Spiracular field fringed with long firm setae (
<figureCitation id="13742A45BB0FFFB3F24AFC5BFDFFF263" box="[532,615,1017,1043]" captionStart="FIGURES 18–27" captionStartId="9.[151,264,1907,1932]" captionTargetBox="[184,1413,180,1883]" captionTargetId="figure@9.[173,1415,174,1887]" captionTargetPageId="9" captionText="FIGURES 18–27. Larva of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25)." figureDoi="http://doi.org/10.5281/zenodo.3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="8" pageNumber="289">Fig. 27</figureCitation>
). Spiracle large, oblong, placed obliquely and inclined dorsally.Anal field consists of two pairs of blunt, white, fleshy anal papillae.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FBE3FAD2F20F" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FBE3FE94F22B" bold="true" box="[199,268,1089,1115]" pageId="8" pageNumber="289">Pupa,</emphasis>
female (N=1). Male pupa unknown. Female pupa 
<quantity id="4CB79B25BB0FFFB3F361FBE2FC0CF22B" box="[831,916,1088,1115]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="289" unit="mm" value="8.5">8.5 mm</quantity>
long, 
<quantity id="4CB79B25BB0FFFB3F387FBE3FBB6F22B" box="[985,1070,1089,1115]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="289" unit="mm" value="1.3">1.3 mm</quantity>
wide. Abdomen yellowish white (
<figureCitation id="13742A45BB0FFFB3F0C1FBC7FEB1F20F" box="[159,297,1125,1151]" captionStart="FIGURES 28–32" captionStartId="10.[151,264,1925,1950]" captionTargetBox="[161,1428,703,1897]" captionTargetId="figure@10.[151,1436,697,1905]" captionTargetPageId="10" captionText="FIGURES 28–32. Female pupa of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663463" httpUri="https://zenodo.org/record/3663463/files/figure.png" pageId="8" pageNumber="289">Figs. 28, 29</figureCitation>
). Head, thorax, wings, legs and terminal segment dark brown, darker than the rest of the body.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FB2BFE80F29B" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FB2BFE93F2D3" box="[199,267,1161,1187]" italics="true" pageId="8" pageNumber="289">Head.</emphasis>
Cephalic crest absent. Surface non-setose. Antennal sheaths short, only slightly extending beyond base of wing (
<figureCitation id="13742A45BB0FFFB3F080FB0FFEAAF2B7" box="[222,306,1197,1223]" captionStart="FIGURES 28–32" captionStartId="10.[151,264,1925,1950]" captionTargetBox="[161,1428,703,1897]" captionTargetId="figure@10.[151,1436,697,1905]" captionTargetPageId="10" captionText="FIGURES 28–32. Female pupa of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663463" httpUri="https://zenodo.org/record/3663463/files/figure.png" pageId="8" pageNumber="289">Fig. 30</figureCitation>
). Labrum trapezoid with bluntly rounded apex. Labial lobe diamond-shaped. Maxillary palpus broad, transversal.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FB57FA1BF30B" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FB57FEB8F37F" box="[199,288,1269,1295]" italics="true" pageId="8" pageNumber="289">Thorax.</emphasis>
Pronotal horns flattened, triangular, wide at the base and gradually tapering toward the apex, twice as long as width at the base (
<figureCitation id="13742A45BB0FFFB3F1E7FABBFD95F343" box="[441,525,1305,1331]" captionStart="FIGURES 28–32" captionStartId="10.[151,264,1925,1950]" captionTargetBox="[161,1428,703,1897]" captionTargetId="figure@10.[151,1436,697,1905]" captionTargetPageId="10" captionText="FIGURES 28–32. Female pupa of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663463" httpUri="https://zenodo.org/record/3663463/files/figure.png" pageId="8" pageNumber="289">Fig. 31</figureCitation>
). Base of horns very close to each other, apices separated. Dorsum of thorax nonsetose. Apex of wing reaching the end of second abdominal segment (
<figureCitation id="13742A45BB0FFFB3F3C9FA9FFC72F327" box="[919,1002,1341,1367]" captionStart="FIGURES 28–32" captionStartId="10.[151,264,1925,1950]" captionTargetBox="[161,1428,703,1897]" captionTargetId="figure@10.[151,1436,697,1905]" captionTargetPageId="10" captionText="FIGURES 28–32. Female pupa of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663463" httpUri="https://zenodo.org/record/3663463/files/figure.png" pageId="8" pageNumber="289">Fig. 30</figureCitation>
). Legs reaching almost to the middle of fourth abdominal segment. The innermost pair of legs are the longest, the outermost pair – the shortest (
<figureCitation id="13742A45BB0FFFB3F57CFAC3FAEEF30B" box="[1314,1398,1377,1403]" captionStart="FIGURES 28–32" captionStartId="10.[151,264,1925,1950]" captionTargetBox="[161,1428,703,1897]" captionTargetId="figure@10.[151,1436,697,1905]" captionTargetPageId="10" captionText="FIGURES 28–32. Female pupa of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663463" httpUri="https://zenodo.org/record/3663463/files/figure.png" pageId="8" pageNumber="289">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FA27FD9DF397" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FA27FEA1F3EF" box="[199,313,1413,1439]" italics="true" pageId="8" pageNumber="289">Abdomen.</emphasis>
Segments III–VII with dorsal and ventral creeping welts. Surface of abdominal segments non-setose. Spiracles hardly visible. Terminal segment elongate (
<figureCitation id="13742A45BB0FFFB3F2ADFA0BFCD1F3B3" box="[755,841,1449,1475]" captionStart="FIGURES 28–32" captionStartId="10.[151,264,1925,1950]" captionTargetBox="[161,1428,703,1897]" captionTargetId="figure@10.[151,1436,697,1905]" captionTargetPageId="10" captionText="FIGURES 28–32. Female pupa of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov. 28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663463" httpUri="https://zenodo.org/record/3663463/files/figure.png" pageId="8" pageNumber="289">Fig. 32</figureCitation>
). Sheath of cerci and sheath of valvae are similar in length. Additional spines absent.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099FA52FE42F023" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099FA52FEB2F07B" bold="true" box="[199,298,1520,1547]" pageId="8" pageNumber="289">Habitat.</emphasis>
Deciduous broad-leaved forest (
<figureCitation id="13742A45BB0FFFB3F2C5FA53FD77F07B" box="[667,751,1521,1547]" captionStart="FIGURE 33" captionStartId="12.[151,250,1035,1060]" captionTargetBox="[151,1435,166,1022]" captionTargetId="figure@12.[151,1435,166,1022]" captionTargetPageId="12" captionText="FIGURE 33. Habitat (type locality) of Atypophthalmus (Microlimonia) jeju Podenas &amp; Podeniene, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3663465" httpUri="https://zenodo.org/record/3663465/files/figure.png" pageId="8" pageNumber="289">Fig. 33</figureCitation>
). 
<taxonomicName id="4C4F4D43BB0FFFB3F35AFA53FA99F07B" authority="Podenas &amp; Podeniene" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[772,1281,1521,1547]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="jeju" subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0FFFB3F35AFA53FC85F07B" box="[772,797,1521,1547]" italics="true" pageId="8" pageNumber="289">A.</emphasis>
(
<emphasis id="B93BEAD2BB0FFFB3F370FA53FC5FF07B" box="[814,967,1521,1547]" italics="true" pageId="8" pageNumber="289">Microlimonia</emphasis>
) 
<emphasis id="B93BEAD2BB0FFFB3F386FA50FB9AF07B" box="[984,1026,1522,1547]" italics="true" pageId="8" pageNumber="289">jeju</emphasis>
Podenas &amp; Podeniene
</taxonomicName>
was the most abundant crane fly species at the 
<typeStatus id="54F48862BB0FFFB3F250F9B7FDA6F05F" box="[526,574,1557,1583]" pageId="8" pageNumber="289">type</typeStatus>
locality. Larvae and pupa were found under the bark of truncated deciduous tree trunks in sap with fungi.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099F9FEFE55F007" blockId="8.[151,1437,153,2015]" box="[199,461,1628,1655]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099F9FEFED8F007" bold="true" box="[199,320,1628,1655]" pageId="8" pageNumber="289">Elevation.</emphasis>
<quantity id="4CB79B25BB0FFFB3F119F9FFFE51F007" box="[327,457,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="8" pageNumber="289" unit="m" value="350.0" valueMax="600.0" valueMin="100.0">100–600 m</quantity>
.
</paragraph>
<paragraph id="8BF036C0BB0FFFB3F099F922FC8FF0EB" blockId="8.[151,1437,153,2015]" box="[199,791,1664,1691]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099F922FE02F0EB" bold="true" box="[199,410,1664,1691]" pageId="8" pageNumber="289">Period of activity.</emphasis>
Collected only at the end of May.
</paragraph>
</subSubSection>
<subSubSection id="C355654BBB0FFFB3F099F906FBCDF0CF" box="[199,1109,1700,1727]" pageId="8" pageNumber="289" type="distribution">
<paragraph id="8BF036C0BB0FFFB3F099F906FBCDF0CF" blockId="8.[151,1437,153,2015]" box="[199,1109,1700,1727]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099F906FEF9F0CF" bold="true" box="[199,353,1700,1727]" pageId="8" pageNumber="289">Distribution.</emphasis>
Currently known only from Jeju Island, 
<collectingCountry id="F3587650BB0FFFB3F374F907FC60F0CF" box="[810,1016,1701,1727]" name="South Korea" pageId="8" pageNumber="289">Republic of Korea</collectingCountry>
(
<collectingCountry id="F3587650BB0FFFB3F456F907FBD1F0CF" box="[1032,1097,1701,1727]" name="South Korea" pageId="8" pageNumber="289">ROK</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C355654BBB0FFFB3F099F96BFBB0F093" box="[199,1064,1737,1763]" pageId="8" pageNumber="289" type="etymology">
<paragraph id="8BF036C0BB0FFFB3F099F96BFBB0F093" blockId="8.[151,1437,153,2015]" box="[199,1064,1737,1763]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099F96BFED6F093" bold="true" box="[199,334,1737,1763]" pageId="8" pageNumber="289">Etymology.</emphasis>
Species is named after Jeju Island, 
<collectingCountry id="F3587650BB0FFFB3F283F96BFC83F093" box="[733,795,1737,1763]" name="South Korea" pageId="8" pageNumber="289">ROK</collectingCountry>
, where it was collected.
</paragraph>
</subSubSection>
<subSubSection id="C355654BBB0FFFB0F099F94FFE97F007" lastPageId="11" lastPageNumber="292" pageId="8" pageNumber="289" type="discussion">
<paragraph id="8BF036C0BB0FFFB3F099F94FFD1BF13F" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="289">
<emphasis id="B93BEAD2BB0FFFB3F099F94FFEA3F177" bold="true" box="[199,315,1773,1799]" pageId="8" pageNumber="289">Remarks.</emphasis>
Analysis of mitochondrial DNA Cytochrome c oxidase subunit I barcoding region showed no difference in sequences of adults and larvae (GenBank accession numbers 
<accessionNumber id="941CAB23BB0FFFB3F391F8B2FBC8F15B" box="[975,1104,1808,1835]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN715236" pageId="8" pageNumber="289">MN715236</accessionNumber>
and 
<accessionNumber id="941CAB23BB0FFFB3F4D5F8B2FA88F15B" box="[1163,1296,1808,1835]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN715237" pageId="8" pageNumber="289">MN715237</accessionNumber>
), leaving no doubts that they belong to the same species.
</paragraph>
<paragraph id="8BF036C0BB0FFFB1F099F8FBFF6BF4DB" blockId="8.[151,1437,153,2015]" lastBlockId="10.[151,1437,153,683]" lastPageId="10" lastPageNumber="291" pageId="8" pageNumber="289">
<taxonomicName id="4C4F4D43BB0FFFB3F099F8FBFE36F103" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[199,430,1881,1907]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">
<emphasis id="B93BEAD2BB0FFFB3F099F8FBFE36F103" box="[199,430,1881,1907]" italics="true" pageId="8" pageNumber="289">Atypophthalmus jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0FFFB3F1EBF8FBFD96F103" bold="true" box="[437,526,1881,1907]" pageId="8" pageNumber="289">
<taxonomicNameLabel id="A20857A9BB0FFFB3F1EBF8FBFD96F103" box="[437,526,1881,1907]" pageId="8" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
could be grouped with two other East Palearctic species, 
<taxonomicName id="4C4F4D43BB0FFFB3F4CDF8FBFF41F1E7" authority="(Alexander, 1938)" baseAuthorityName="Alexander" baseAuthorityYear="1938" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="egressus">
<emphasis id="B93BEAD2BB0FFFB3F4CDF8FBFA89F103" box="[1171,1297,1881,1907]" italics="true" pageId="8" pageNumber="289">A. egressus</emphasis>
(
<bibRefCitation id="EFDE4B31BB0FFFB3F57EF8FBFF48F1E7" author="Alexander, C. P." pageId="8" pageNumber="289" pagination="129 - 166" refId="ref7802" refString="Alexander, C. P. (1938) New or little-known Tipulidae from eastern Asia (Diptera). XL. Philippine Journal of Science, 67, 129 - 166." type="journal article" year="1938">Alexander, 1938</bibRefCitation>
)
</taxonomicName>
from 
<collectingRegion id="498BF822BB0FFFB3F17FF8DFFEE6F1E7" box="[289,382,1917,1943]" country="China" name="Sichuan" pageId="8" pageNumber="289">Sichuan</collectingRegion>
, 
<collectingCountry id="F3587650BB0FFFB3F1D5F8DFFE56F1E7" box="[395,462,1917,1943]" name="China" pageId="8" pageNumber="289">China</collectingCountry>
and 
<taxonomicName id="4C4F4D43BB0FFFB3F256F8DFFC1DF1E7" authority="(Alexander, 1954)" baseAuthorityName="Alexander" baseAuthorityYear="1954" box="[520,901,1916,1943]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="omogoensis">
<emphasis id="B93BEAD2BB0FFFB3F256F8DFFD35F1E7" box="[520,685,1917,1943]" italics="true" pageId="8" pageNumber="289">A. omogoensis</emphasis>
(
<bibRefCitation id="EFDE4B31BB0FFFB3F2E0F8DFFCE4F1E7" author="Alexander, C. P." box="[702,892,1916,1943]" pageId="8" pageNumber="289" pagination="263 - 308" refId="ref7835" refString="Alexander, C. P. (1954) Records and descriptions of Japanese Tipulidae (Diptera). Part III. The crane-flies of Shikoku. III. Philippine Journal of Science, 82, 263 - 308." type="journal article" year="1954">Alexander, 1954</bibRefCitation>
)
</taxonomicName>
from Shikoku, 
<collectingCountry id="F3587650BB0FFFB3F464F8DFFBE5F1E7" box="[1082,1149,1917,1943]" name="Japan" pageId="8" pageNumber="289">Japan</collectingCountry>
. All have similar general appearance, structures of male terminalia, and wing venation. Closest in general structure is 
<taxonomicName id="4C4F4D43BB0FFFB3F4F1F803FABBF1CB" box="[1199,1315,1953,1979]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="egressa">
<emphasis id="B93BEAD2BB0FFFB3F4F1F803FABBF1CB" box="[1199,1315,1953,1979]" italics="true" pageId="8" pageNumber="289">A. egressa</emphasis>
</taxonomicName>
. The male terminalia of all three species have an elongate gonocoxite with a very large elongate bilobed ventro-mesial lobe. Both lobes in A. jeju sp. nov. are distinctly different in size, the dorsal being distinctly shorter than the ventral, while both lobes are nearly equal in size in A. egressus and A. omogoensis. The gonostylus is single in all three species, but in A. omogoensis it is trilobed, rostral prolongation twice as long as gonostylus itself, bilobed in A. jeju 
<taxonomicNameLabel id="A20857A9BB0DFFB1F51DFF43FA04F68B" box="[1347,1436,225,251]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C4F4D43BB0DFFB1F099FEA7FEDDF76F" baseAuthorityName="Alexander" baseAuthorityYear="1938" box="[199,325,261,287]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="egressus">
<emphasis id="B93BEAD2BB0DFFB1F099FEA7FEDDF76F" box="[199,325,261,287]" italics="true" pageId="10" pageNumber="291">A. egressus</emphasis>
</taxonomicName>
, but in 
<taxonomicName id="4C4F4D43BB0DFFB1F1C2FEA7FD82F76F" baseAuthorityName="Alexander" baseAuthorityYear="1938" box="[412,538,261,287]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="egressus">
<emphasis id="B93BEAD2BB0DFFB1F1C2FEA7FD82F76F" box="[412,538,261,287]" italics="true" pageId="10" pageNumber="291">A. egressus</emphasis>
</taxonomicName>
, the gonostylus is smaller, bifid at apex with a very elongate rostral lobe, which is distinctly longer than the gonostylus. The gonostylus in 
<taxonomicName id="4C4F4D43BB0DFFB1F37CFE8BFCF3F733" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[802,875,297,323]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">
<emphasis id="B93BEAD2BB0DFFB1F37CFE8BFCF3F733" box="[802,875,297,323]" italics="true" pageId="10" pageNumber="291">A. jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0DFFB1F32FFE8BFC53F733" bold="true" box="[881,971,297,323]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F32FFE8BFC53F733" box="[881,971,297,323]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinctly larger, the dorsal lobe is very small and does not reach the tip of the ventral lobe, the rostral prolongation distinctly shorter and not exceeding the length of the gonostylus. The meso-apical lobe of the paramere narrow with apex acute in 
<taxonomicName id="4C4F4D43BB0DFFB1F4D0FED3FA94F7FB" baseAuthorityName="Alexander" baseAuthorityYear="1938" box="[1166,1292,369,395]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="egressus">
<emphasis id="B93BEAD2BB0DFFB1F4D0FED3FA94F7FB" box="[1166,1292,369,395]" italics="true" pageId="10" pageNumber="291">A. egressus</emphasis>
</taxonomicName>
and 
<emphasis id="B93BEAD2BB0DFFB1F51DFED3FF73F7DF" italics="true" pageId="10" pageNumber="291">A. omo- goensis</emphasis>
, but wider and blunt at apex in 
<taxonomicName id="4C4F4D43BB0DFFB1F212FE37FD0CF7DF" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[588,660,405,431]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">
<emphasis id="B93BEAD2BB0DFFB1F212FE37FD0CF7DF" box="[588,660,405,431]" italics="true" pageId="10" pageNumber="291">A. jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0DFFB1F2C5FE37FD6CF7DF" bold="true" box="[667,756,405,431]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F2C5FE37FD6CF7DF" box="[667,756,405,431]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The wing has an open discal cell in 
<taxonomicName id="4C4F4D43BB0DFFB1F4D7FE37FB4AF7DF" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[1161,1234,405,431]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">
<emphasis id="B93BEAD2BB0DFFB1F4D7FE37FB4AF7DF" box="[1161,1234,405,431]" italics="true" pageId="10" pageNumber="291">A. jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0DFFB1F486FE37FAAAF7DF" bold="true" box="[1240,1330,405,431]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F486FE37FAAAF7DF" box="[1240,1330,405,431]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, while 
<emphasis id="B93BEAD2BB0DFFB1F5DAFE37FF6EF7A3" italics="true" pageId="10" pageNumber="291">
<taxonomicName id="4C4F4D43BB0DFFB1F5DAFE37FA05F7DF" baseAuthorityName="Alexander" baseAuthorityYear="1938" box="[1412,1437,405,431]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="egressus">A.</taxonomicName>
egressus
</emphasis>
and 
<taxonomicName id="4C4F4D43BB0DFFB1F172FE1BFE56F7A3" baseAuthorityName="Alexander" baseAuthorityYear="1954" box="[300,462,441,467]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="omogoensis">
<emphasis id="B93BEAD2BB0DFFB1F172FE1BFE56F7A3" box="[300,462,441,467]" italics="true" pageId="10" pageNumber="291">A. omogoensis</emphasis>
</taxonomicName>
have a more a typical closed discal cell, as within the genus. Cell 
<emphasis id="B93BEAD2BB0DFFB1F4F2FE18FB43F7A8" box="[1196,1243,442,472]" italics="true" pageId="10" pageNumber="291">
m 
<subScript id="17CB3485BB0DFFB1F49EFE68FB43F7A8" attach="left" box="[1216,1243,458,472]" fontSize="6" pageId="10" pageNumber="291">1+2</subScript>
</emphasis>
is short-petiolate in 
<taxonomicName id="4C4F4D43BB0DFFB1F0EAFE7FFF66F787" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[180,254,477,503]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">
<emphasis id="B93BEAD2BB0DFFB1F0EAFE7FFF66F787" box="[180,254,477,503]" italics="true" pageId="10" pageNumber="291">A. jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0DFFB1F15BFE7FFEC7F787" bold="true" box="[261,351,477,503]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F15BFE7FFEC7F787" box="[261,351,477,503]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with the stem nearly three times shorter than the cell, and long-petiolate in 
<taxonomicName id="4C4F4D43BB0DFFB1F4EAFE7FFAAAF787" baseAuthorityName="Alexander" baseAuthorityYear="1938" box="[1204,1330,477,503]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="egressus">
<emphasis id="B93BEAD2BB0DFFB1F4EAFE7FFAAAF787" box="[1204,1330,477,503]" italics="true" pageId="10" pageNumber="291">A. egressus</emphasis>
</taxonomicName>
, with the stem nearly as long as cell. The tip of vein 
<emphasis id="B93BEAD2BB0DFFB1F22DFDA3FD0DF450" box="[627,661,513,544]" italics="true" pageId="10" pageNumber="291">
Sc 
<subScript id="17CB3485BB0DFFB1F2D3FDB0FD0DF450" attach="left" box="[653,661,530,544]" fontSize="6" pageId="10" pageNumber="291">1</subScript>
</emphasis>
reaches to about one third of 
<emphasis id="B93BEAD2BB0DFFB1F381FDA0FC63F46B" box="[991,1019,514,539]" italics="true" pageId="10" pageNumber="291">Rs</emphasis>
length in 
<taxonomicName id="4C4F4D43BB0DFFB1F434FDA3FB2BF46B" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[1130,1203,513,539]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">
<emphasis id="B93BEAD2BB0DFFB1F434FDA3FB2BF46B" box="[1130,1203,513,539]" italics="true" pageId="10" pageNumber="291">A. jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0DFFB1F4E7FDA3FA8AF46B" bold="true" box="[1209,1298,513,539]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F4E7FDA3FA8AF46B" box="[1209,1298,513,539]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but beyond the mid-length of 
<emphasis id="B93BEAD2BB0DFFB1F13FFD84FEE5F44F" box="[353,381,550,575]" italics="true" pageId="10" pageNumber="291">Rs</emphasis>
in 
<emphasis id="B93BEAD2BB0DFFB1F1FCFD87FDE0F44F" box="[418,632,549,575]" italics="true" pageId="10" pageNumber="291">
<taxonomicName id="4C4F4D43BB0DFFB1F1FCFD87FDBCF44F" baseAuthorityName="Alexander" baseAuthorityYear="1938" box="[418,548,549,575]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="egressus">A. egressus</taxonomicName>
. 
<taxonomicName id="4C4F4D43BB0DFFB1F270FD87FDE0F44F" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[558,632,549,575]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">A. jeju</taxonomicName>
</emphasis>
<emphasis id="B93BEAD2BB0DFFB1F221FD87FD41F44F" bold="true" box="[639,729,549,575]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F221FD87FD41F44F" box="[639,729,549,575]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs by yellow coloration of rostrum and palpus, which are black or brown in both of the other species. 
<taxonomicName id="4C4F4D43BB0DFFB1F2E4FDEBFC9DF413" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[698,773,585,611]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov.">
<emphasis id="B93BEAD2BB0DFFB1F2E4FDEBFC9DF413" box="[698,773,585,611]" italics="true" pageId="10" pageNumber="291">A. jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0DFFB1F353FDEBFCF0F413" bold="true" box="[781,872,585,611]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F353FDEBFCF0F413" box="[781,872,585,611]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only Korean 
<taxonomicName id="4C4F4D43BB0DFFB1F415FDEBFA99F413" baseAuthorityName="Savchenko &amp; Krivolutskaya" baseAuthorityYear="1976" box="[1099,1281,585,611]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAD2BB0DFFB1F415FDEBFA99F413" box="[1099,1281,585,611]" italics="true" pageId="10" pageNumber="291">Atypophthalmus</emphasis>
</taxonomicName>
with an open discal cell, except some specimens of 
<taxonomicName id="4C4F4D43BB0DFFB1F21BFDCFFD68F4F7" baseAuthorityName="Alexander" baseAuthorityYear="1971" box="[581,752,621,647]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="stylacanthus">
<emphasis id="B93BEAD2BB0DFFB1F21BFDCFFD68F4F7" box="[581,752,621,647]" italics="true" pageId="10" pageNumber="291">A. stylacanthus</emphasis>
</taxonomicName>
that may have an open discal cell (
<figureCitation id="13742A45BB0DFFB1F420FDCFFB43F4F7" box="[1150,1243,621,647]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1932,1957]" captionTargetBox="[285,1294,185,1900]" captionTargetId="figure@2.[276,1312,174,1913]" captionTargetPageId="2" captionText="FIGURES 1–6. Wings of Korean Atypophthalmus. 1: A. (Atypophthalmus) crinitus; 2: A. (A.) kleini Podenas, sp. nov.; 3: A. (A.) stylacanthus; 4: A. (Microlimonia) inelegans; 5: A. (M.) jeju Podenas, Podeniene, sp. nov.; 6: A. (M.) machidai. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3663451" httpUri="https://zenodo.org/record/3663451/files/figure.png" pageId="10" pageNumber="291">Fig. 2 C</figureCitation>
in 
<bibRefCitation id="EFDE4B31BB0DFFB1F4A1FDCFFA04F4F7" author="Podenas, S. &amp; Aukstikalniene, R. &amp; Byun, H. - W. &amp; Klein, T. A. &amp; Kim, H. C. &amp; Kim, T. - W. &amp; Kang, T. - H. &amp; Seo, H. - Y." box="[1279,1436,621,647]" pageId="10" pageNumber="291" pagination="258 - 279" refId="ref8385" refString="Podenas, S., Aukstikalniene, R., Byun, H. - W., Klein, T. A., Kim, H. C., Kim, T. - W., Kang, T. - H. &amp; Seo, H. - Y. (2017) Limoniinae crane flies (Diptera: Limoniinae) new to Korea II. Journal of Species Research, 6 (3), 258 - 279. https: // doi. org / 10.12651 / JSR. 2017.6.3.258." type="journal article" year="2017">
Podenas 
<emphasis id="B93BEAD2BB0DFFB1F53AFDCCFA0DF4F7" box="[1380,1429,621,647]" italics="true" pageId="10" pageNumber="291">et al</emphasis>
.
</bibRefCitation>
(2017)).
</paragraph>
<caption id="DF306648BB0EFFB2F0C9F8D1FAD0F188" ID-DOI="http://doi.org/10.5281/zenodo.3663459" ID-Zenodo-Dep="3663459" httpUri="https://zenodo.org/record/3663459/files/figure.png" pageId="9" pageNumber="290" startId="9.[151,264,1907,1932]" targetBox="[184,1413,180,1883]" targetPageId="9">
<paragraph id="8BF036C0BB0EFFB2F0C9F8D1FAD0F188" blockId="9.[151,1436,1907,2040]" pageId="9" pageNumber="290">
<emphasis id="B93BEAD2BB0EFFB2F0C9F8D1FECFF1FC" bold="true" box="[151,343,1907,1932]" pageId="9" pageNumber="290">FIGURES 18–27.</emphasis>
Larva of 
<taxonomicName id="4C4F4D43BB0EFFB2F19DF8D1FBB3F1FC" authority="Podenas &amp; Podeniene" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[451,1067,1907,1932]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0EFFB2F19DF8D1FDF0F1FB" box="[451,616,1907,1931]" italics="true" pageId="9" pageNumber="290">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB0EFFB2F227F8D1FC9BF1FB" box="[633,771,1907,1931]" italics="true" pageId="9" pageNumber="290">Microlimonia</emphasis>
) 
<emphasis id="B93BEAD2BB0EFFB2F34BF8D6FCA3F1FB" box="[789,827,1908,1931]" italics="true" pageId="9" pageNumber="290">jeju</emphasis>
Podenas &amp; Podeniene
</taxonomicName>
, 
<emphasis id="B93BEAD2BB0EFFB2F467F8D6FB14F1FC" bold="true" box="[1081,1164,1908,1932]" pageId="9" pageNumber="290">
<taxonomicNameLabel id="A20857A9BB0EFFB2F467F8D6FB14F1FC" box="[1081,1164,1908,1932]" pageId="9" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
18: general view of head capsule, dorsal aspect; 19: labrum and clypeus, dorsal view; 20: right antenna, dorsal view; 21: right mandible, dorsal view; 22: right maxilla, ventral view; 23: hypopharynx and prementum, ventral view; 24: hypostoma, ventral view; 25: general view, lateral aspect; 26: terminal segments, dorsal view; 27: spiracular field. Scale bars: 0.1 mm (18–24, 26, 27), 1 mm (25).
</paragraph>
</caption>
<caption id="DF306648BB0DFFB1F0C9F827FB8CF195" ID-DOI="http://doi.org/10.5281/zenodo.3663463" ID-Zenodo-Dep="3663463" httpUri="https://zenodo.org/record/3663463/files/figure.png" pageId="10" pageNumber="291" startId="10.[151,264,1925,1950]" targetBox="[161,1428,703,1897]" targetPageId="10">
<paragraph id="8BF036C0BB0DFFB1F0C9F827FB8CF195" blockId="10.[151,1436,1925,2022]" pageId="10" pageNumber="291">
<emphasis id="B93BEAD2BB0DFFB1F0C9F827FECCF1EE" bold="true" box="[151,340,1925,1950]" pageId="10" pageNumber="291">FIGURES 28–32.</emphasis>
Female pupa of 
<taxonomicName id="4C4F4D43BB0DFFB1F1A1F824FBC1F1EE" authority="Podenas &amp; Podeniene" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[511,1113,1925,1950]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0DFFB1F1A1F824FD3CF1EE" box="[511,676,1926,1950]" italics="true" pageId="10" pageNumber="291">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB0DFFB1F2ECF824FCA4F1EE" box="[690,828,1926,1950]" italics="true" pageId="10" pageNumber="291">Microlimonia</emphasis>
) 
<emphasis id="B93BEAD2BB0DFFB1F312F825FCEAF1EE" box="[844,882,1927,1950]" italics="true" pageId="10" pageNumber="291">jeju</emphasis>
Podenas &amp; Podeniene
</taxonomicName>
, 
<emphasis id="B93BEAD2BB0DFFB1F43DF824FB2CF1EE" bold="true" box="[1123,1204,1926,1950]" pageId="10" pageNumber="291">
<taxonomicNameLabel id="A20857A9BB0DFFB1F43DF824FB2CF1EE" box="[1123,1204,1926,1950]" pageId="10" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
28: general view, ventral aspect; 29: general view, lateral aspect; 30: head, thorax, I–IV abdominal segments, ventral view; 31: head, thorax, I–IV abdominal segments, dorsal view; 32: terminal segments, lateral view. Scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BF036C0BB0CFFB0F099FF3BFBA2F76F" blockId="11.[151,1437,153,1655]" pageId="11" pageNumber="292">
The larva of 
<emphasis id="B93BEAD2BB0CFFB0F107FF3BFEF2F6C3" box="[345,362,153,179]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. 
<taxonomicName id="4C4F4D43BB0CFFB0F127FF3BFD82F6C3" baseAuthorityName="Alexander" baseAuthorityYear="1921" box="[377,538,153,179]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="machidai" subGenus="Microlimonia">
(
<emphasis id="B93BEAD2BB0CFFB0F1DCFF38FE01F6C3" box="[386,409,154,179]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F1ECFF3BFD82F6C3" box="[434,538,153,179]" italics="true" pageId="11" pageNumber="292">machidai</emphasis>
</taxonomicName>
was described by 
<bibRefCitation id="EFDE4B31BB0CFFB0F2B5FF3BFC52F6C3" author="Krivosheina, N. P." box="[747,970,153,179]" pageId="11" pageNumber="292" pagination="135 - 140" refId="ref8046" refString="Krivosheina, N. P. (2010) On the larval morphology and ecology of the limoniid Microlimonia machidai (Diptera, Limoniidae). Entomological Review, 90, 135 - 140. https: // doi. org / 10.1134 / S 0013873810010124" type="journal article" year="2010">Krivosheina (2010)</bibRefCitation>
. Neither the head capsule nor the general appearance of that larva looks like a typical Limoniinae. We have doubts if Krivosheina’s identification is true, because no information is given on rearing. Because of that, no comparison is made between larvae of these two species. The larvae of 
<emphasis id="B93BEAD2BB0CFFB0F1CCFEA7FE3BF76F" box="[402,419,261,287]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. 
<taxonomicName id="4C4F4D43BB0CFFB0F1EFFEA7FDC9F76F" baseAuthorityName="Alexander" baseAuthorityYear="1921" box="[433,593,261,287]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="machidai" subGenus="Microlimonia">
(
<emphasis id="B93BEAD2BB0CFFB0F1E4FEA4FE49F76F" box="[442,465,262,287]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F1B7FEA7FDC9F76F" box="[489,593,261,287]" italics="true" pageId="11" pageNumber="292">machidai</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4F4D43BB0CFFB0F2D9FEA7FC91F76F" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[647,777,261,287]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F2D9FEA7FD00F76F" box="[647,664,261,287]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F2EEFEA4FD5FF76F" box="[688,711,262,287]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F281FEA4FC91F76F" box="[735,777,262,287]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F34EFEA7FCF1F76F" bold="true" box="[784,873,261,287]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F34EFEA7FCF1F76F" box="[784,873,261,287]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are very different.
</paragraph>
<paragraph id="8BF036C0BB0CFFB0F099FE8BFF57F787" blockId="11.[151,1437,153,1655]" pageId="11" pageNumber="292">
In general appearance the larvae of the subgenera 
<taxonomicName id="4C4F4D43BB0CFFB0F2A5FE8BFC29F733" baseAuthorityName="Savchenko &amp; Krivolutskaya" baseAuthorityYear="1976" box="[763,945,297,323]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B93BEAD2BB0CFFB0F2A5FE8BFC29F733" box="[763,945,297,323]" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857A9BB0CFFB0F3E7FE8BFC45F733" box="[953,989,297,323]" pageId="11" pageNumber="292" sensu="stricto">s.s.</taxonomicNameLabel>
and 
<taxonomicName id="4C4F4D43BB0CFFB0F44AFE8BFB35F733" authorityName="Savchenko" authorityYear="1976" box="[1044,1197,297,323]" class="Insecta" family="Limoniidae" genus="Microlimonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAD2BB0CFFB0F44AFE8BFB35F733" box="[1044,1197,297,323]" italics="true" pageId="11" pageNumber="292">Microlimonia</emphasis>
</taxonomicName>
are very similar. The body is white with dorsal and ventral creeping welts on abdominal segments II–VII. Head capsules of the subgenera 
<taxonomicName id="4C4F4D43BB0CFFB0F09FFED3FEEFF7FB" baseAuthorityName="Savchenko &amp; Krivolutskaya" baseAuthorityYear="1976" box="[193,375,369,395]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B93BEAD2BB0CFFB0F09FFED3FEEFF7FB" box="[193,375,369,395]" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857A9BB0CFFB0F121FED3FE3BF7FB" box="[383,419,369,395]" pageId="11" pageNumber="292" sensu="stricto">s.s.</taxonomicNameLabel>
and 
<taxonomicName id="4C4F4D43BB0CFFB0F185FED3FDECF7FB" authorityName="Savchenko" authorityYear="1976" box="[475,628,369,395]" class="Insecta" family="Limoniidae" genus="Microlimonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAD2BB0CFFB0F185FED3FDECF7FB" box="[475,628,369,395]" italics="true" pageId="11" pageNumber="292">Microlimonia</emphasis>
</taxonomicName>
are similar: slightly reduced with short dorsal sutures, frons fused with internolateralia, elongate-oval labrum, clypeus divided into preclypeus and postclypeus, complete and dentate hypostoma. The larvae of both species have spiracular lobes reduced and spiracular fields with ventral pair of sclerites only.
</paragraph>
<paragraph id="8BF036C0BB0CFFB0F099FDA3FBACF54B" blockId="11.[151,1437,153,1655]" pageId="11" pageNumber="292">
Differences were noted in the shape of the frons, as well as the number of mandibular teeth, the number of hypostomal teeth, and the arrangement of sensory structures on the labrum. 
<taxonomicName id="4C4F4D43BB0CFFB0F3ADFD87FB0DF44F" baseAuthorityName="Meigen" baseAuthorityYear="1818" box="[1011,1173,549,575]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="inustus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F3ADFD87FB9CF44F" box="[1011,1028,549,575]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F442FD87FBB5F44F" box="[1052,1069,549,575]" italics="true" pageId="11" pageNumber="292">A</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F418FD84FB0DF44F" box="[1094,1173,550,575]" italics="true" pageId="11" pageNumber="292">inustus</emphasis>
</taxonomicName>
larvae have eleven hypostomal teeth and a tridentate caudal end of frons, whereas 
<taxonomicName id="4C4F4D43BB0CFFB0F31CFDEBFC5CF413" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[834,964,585,611]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F31CFDEBFCCBF413" box="[834,851,585,611]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F335FDE8FC1AF413" box="[875,898,586,611]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F3C4FDE8FC5CF413" box="[922,964,586,611]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F392FDEBFBBEF413" bold="true" box="[972,1062,585,611]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F392FDEBFBBEF413" box="[972,1062,585,611]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has twelve hypostomal teeth and a rounded caudal end of frons. Abdominal segments III–VI of 
<taxonomicName id="4C4F4D43BB0CFFB0F302FDCFFC47F4F7" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[860,991,621,647]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F302FDCFFCF5F4F7" box="[860,877,621,647]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F3D8FDCCFC3DF4F7" box="[902,933,622,647]" italics="true" pageId="11" pageNumber="292">M.</emphasis>
) 
<emphasis id="B93BEAD2BB0CFFB0F3EBFDCCFC47F4F7" box="[949,991,622,647]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F3B9FDCFFBDAF4F7" bold="true" box="[999,1090,621,647]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F3B9FDCFFBDAF4F7" box="[999,1090,621,647]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are almost three times as long as wide, whereas in 
<taxonomicName id="4C4F4D43BB0CFFB0F122FD33FD86F4DB" baseAuthorityName="Meigen" baseAuthorityYear="1818" box="[380,542,657,683]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="inustus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F122FD33FE15F4DB" box="[380,397,657,683]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F1F8FD33FE26F4DB" box="[422,446,657,683]" italics="true" pageId="11" pageNumber="292">A.</emphasis>
) 
<emphasis id="B93BEAD2BB0CFFB0F191FD30FD86F4DB" box="[463,542,658,683]" italics="true" pageId="11" pageNumber="292">inustus</emphasis>
</taxonomicName>
it is only twice as long as wide. The spiracular field of 
<taxonomicName id="4C4F4D43BB0CFFB0F4CBFD33FAAFF4DB" baseAuthorityName="Meigen" baseAuthorityYear="1818" box="[1173,1335,657,683]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="inustus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F4CBFD33FB3EF4DB" box="[1173,1190,657,683]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F4E1FD33FB48F4DB" box="[1215,1232,657,683]" italics="true" pageId="11" pageNumber="292">A</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F4B6FD30FAAFF4DB" box="[1256,1335,658,683]" italics="true" pageId="11" pageNumber="292">inustus</emphasis>
</taxonomicName>
has well developed ventral lobes, whereas 
<taxonomicName id="4C4F4D43BB0CFFB0F24DFD17FD0CF4BF" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[531,660,693,719]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F24DFD17FDBCF4BF" box="[531,548,693,719]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F262FD14FDCBF4BF" box="[572,595,694,719]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F234FD14FD0CF4BF" box="[618,660,694,719]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F2C2FD17FD6EF4BF" bold="true" box="[668,758,693,719]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F2C2FD17FD6EF4BF" box="[668,758,693,719]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has lateral lobes and a vestigial dorsal lobe, all of which are inclined inward. 
<taxonomicName id="4C4F4D43BB0CFFB0F10AFD7BFE6CF483" baseAuthorityName="Meigen" baseAuthorityYear="1818" box="[340,500,729,755]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="inustus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F10AFD7BFEFDF483" box="[340,357,729,755]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F122FD7BFE15F483" box="[380,397,729,755]" italics="true" pageId="11" pageNumber="292">A</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F1FBFD78FE6CF483" box="[421,500,730,755]" italics="true" pageId="11" pageNumber="292">inustus</emphasis>
</taxonomicName>
has a single short seta on the lateral margin and two long widely separated setae on the dorsal margin of the spiracular field. 
<taxonomicName id="4C4F4D43BB0CFFB0F200FD5FFD46F567" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[606,734,765,791]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F200FD5FFDF7F567" box="[606,623,765,791]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F2D8FD5CFD05F567" box="[646,669,766,791]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F2EAFD5CFD46F567" box="[692,734,766,791]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F2BBFD5FFCA6F567" bold="true" box="[741,830,765,791]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F2BBFD5FFCA6F567" box="[741,830,765,791]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has two long and one short setae on the lateral margin and two long setae, close in proximity, on the dorsal margin of the spiracular field.
</paragraph>
<paragraph id="8BF036C0BB0CFFB0F099FCE7FCEBF5D7" blockId="11.[151,1437,153,1655]" pageId="11" pageNumber="292">
The pupae of 
<taxonomicName id="4C4F4D43BB0CFFB0F101FCE7FD8DF52F" baseAuthorityName="Savchenko &amp; Krivolutskaya" baseAuthorityYear="1976" box="[351,533,837,863]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B93BEAD2BB0CFFB0F101FCE7FD8DF52F" box="[351,533,837,863]" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857A9BB0CFFB0F245FCE7FDA7F52F" box="[539,575,837,863]" pageId="11" pageNumber="292" sensu="stricto">s.s.</taxonomicNameLabel>
and 
<taxonomicName id="4C4F4D43BB0CFFB0F22AFCE7FC95F52F" authorityName="Savchenko" authorityYear="1976" box="[628,781,837,863]" class="Insecta" family="Limoniidae" genus="Microlimonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAD2BB0CFFB0F22AFCE7FC95F52F" box="[628,781,837,863]" italics="true" pageId="11" pageNumber="292">Microlimonia</emphasis>
</taxonomicName>
are very similar in general appearance. Pupal head, thorax, wings, legs and terminal segment are dark brown, abdomen yellowish-white. Cephalic crest absent. The surface of head and thorax is smooth with laterally flattened pronotal horns.
</paragraph>
<paragraph id="8BF036C0BB0CFFB0F099FC13FE24F2D3" blockId="11.[151,1437,153,1655]" pageId="11" pageNumber="292">
The most significant difference is the shape of the pronotal horns, which are triangular in shape and wide at the base and narrowing toward the apex in 
<taxonomicName id="4C4F4D43BB0CFFB0F2DCFC77FC9EF59F" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[642,774,981,1007]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F2DCFC77FD0BF59F" box="[642,659,981,1007]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F2F2FC74FD5BF59F" box="[684,707,982,1007]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F282FC74FC9EF59F" box="[732,774,982,1007]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F350FC77FCF0F59F" bold="true" box="[782,872,981,1007]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F350FC77FCF0F59F" box="[782,872,981,1007]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, whereas in 
<taxonomicName id="4C4F4D43BB0CFFB0F3A7FC77FB03F59F" baseAuthorityName="Meigen" baseAuthorityYear="1818" box="[1017,1179,981,1007]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="inustus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F3A7FC77FB92F59F" box="[1017,1034,981,1007]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F47DFC77FBACF59F" box="[1059,1076,981,1007]" italics="true" pageId="11" pageNumber="292">A</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F412FC74FB03F59F" box="[1100,1179,982,1007]" italics="true" pageId="11" pageNumber="292">inustus</emphasis>
</taxonomicName>
they are narrow at the base and become wider medially. The shape of the pronotal horns of 
<taxonomicName id="4C4F4D43BB0CFFB0F3C3FC5BFB87F263" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[925,1055,1017,1043]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F3C3FC5BFC36F263" box="[925,942,1017,1043]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F398FC58FC45F263" box="[966,989,1018,1043]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F3ABFC58FB87F263" box="[1013,1055,1018,1043]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F478FC5BFB18F263" bold="true" box="[1062,1152,1017,1043]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F478FC5BFB18F263" box="[1062,1152,1017,1043]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is more similar to that of 
<taxonomicName id="4C4F4D43BB0CFFB0F0C9FBBFFD35F247" authority="(Loew, 1873)" baseAuthorityName="Loew" baseAuthorityYear="1873" box="[151,685,1053,1079]" class="Insecta" family="Limoniidae" genus="Achyrolimonia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="decemmmaculata">
<emphasis id="B93BEAD2BB0CFFB0F0C9FBBFFD91F247" box="[151,521,1053,1079]" italics="true" pageId="11" pageNumber="292">Achyrolimonia decemmmaculata</emphasis>
(
<bibRefCitation id="EFDE4B31BB0CFFB0F244FBBFFD3DF247" author="Loew, H." box="[538,677,1053,1079]" pageId="11" pageNumber="292" refId="ref8180" refString="Loew, H. (1873) Beschreibungen europaischer Dipteren. Vol. 3. H. W. Schmidt, Halle, viii + 320 pp." type="book" year="1873">Loew, 1873</bibRefCitation>
)
</taxonomicName>
. Another difference was in the length of the leg sheath. In 
<taxonomicName id="4C4F4D43BB0CFFB0F50FFBBFFF7EF22B" baseAuthorityName="Meigen" baseAuthorityYear="1818" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="inustus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F50FFBBFFAFAF247" box="[1361,1378,1053,1079]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F525FBBFFA14F247" box="[1403,1420,1053,1079]" italics="true" pageId="11" pageNumber="292">A</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F0C9FBE0FF7EF22B" box="[151,230,1090,1115]" italics="true" pageId="11" pageNumber="292">inustus</emphasis>
</taxonomicName>
leg sheaths reach the fifth abdominal segment and end at the same level, whereas in 
<taxonomicName id="4C4F4D43BB0CFFB0F4CAFBE3FA8CF22B" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[1172,1300,1089,1115]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F4CAFBE3FB3DF22B" box="[1172,1189,1089,1115]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F4E2FBE0FB4BF22B" box="[1212,1235,1090,1115]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F4B4FBE0FA8CF22B" box="[1258,1300,1090,1115]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F545FBE3FAECF22B" bold="true" box="[1307,1396,1089,1115]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F545FBE3FAECF22B" box="[1307,1396,1089,1115]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the leg sheaths reach nearly to the middle of fourth abdominal segment, the innermost pair longer and the outermost pair shorter than mid-pair.
</paragraph>
<paragraph id="8BF036C0BB0CFFB0F099FB0FFECEF05F" blockId="11.[151,1437,153,1655]" pageId="11" pageNumber="292">
The larval and pupal characters of 
<taxonomicName id="4C4F4D43BB0CFFB0F20DFB0FFC25F2B7" authority="" box="[595,957,1197,1223]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="subGenus" subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F20DFB0FFC91F2B7" box="[595,777,1197,1223]" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB0CFFB0F345FB0FFC2CF2B7" box="[795,948,1197,1223]" italics="true" pageId="11" pageNumber="292">Microlimonia</emphasis>
)
</taxonomicName>
that are listed in this paper allow preliminary placement of the subgenus among the Limoniinae. Characters were analyzed with the use of cladogram of nonquantitative analysis of phylogenetic relationship based on larval and pupal characters made by 
<bibRefCitation id="EFDE4B31BB0CFFB0F4ABFB57FEC4F343" author="Oosterbroek, P. &amp; Theowald, B." pageId="11" pageNumber="292" pagination="211 - 267" refId="ref8338" refString="Oosterbroek, P. &amp; Theowald, B. (1991) Phylogeny of the Tipuloidea based on characters of larvae and pupae (Diptera, Nematocera) with an index to the literature except Tipulidae. Tijdschrift voor Entomologie, 134, 211 - 267." type="journal article" year="1991">Oosterbroek &amp; Theowald (1991)</bibRefCitation>
, which is the most suitable. 
<taxonomicName id="4C4F4D43BB0CFFB0F2C0FABBFB9FF343" authority="" box="[670,1031,1305,1331]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="subGenus" subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F2C0FABBFCCCF343" box="[670,852,1305,1331]" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB0CFFB0F33BFABBFC66F343" box="[869,1022,1305,1331]" italics="true" pageId="11" pageNumber="292">Microlimonia</emphasis>
)
</taxonomicName>
possess characters 18 (inner maxillary lobe with separate sclerotized plate), 19 (externolateralia strengthened), but lacks synapomorphy 9 (pupae in silken cocoon) and belongs to the clade Limoniinae. 
<taxonomicName id="4C4F4D43BB0CFFB0F281FAC3FBDEF30B" authority="" box="[735,1094,1377,1403]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="subGenus" subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F281FAC3FC0DF30B" box="[735,917,1377,1403]" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB0CFFB0F3FBFAC3FBA6F30B" box="[933,1086,1377,1403]" italics="true" pageId="11" pageNumber="292">Microlimonia</emphasis>
)
</taxonomicName>
has synapomorphy 100 (pupal creeping welts elliptical), 101 (spiracles oblong or elliptical), 102 (spiracles placed oblique), 103 (larval segments 2 to 10 with creeping welts), but lacks character 99 (larvae in tube or case), placing this subgenus in 
<taxonomicName id="4C4F4D43BB0CFFB0F557FA0BFAF5F3B3" authorityName="Schiner" authorityYear="1863" box="[1289,1389,1449,1475]" class="Insecta" family="Limoniidae" genus="Elliptera" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAD2BB0CFFB0F557FA0BFAF5F3B3" box="[1289,1389,1449,1475]" italics="true" pageId="11" pageNumber="292">Elliptera</emphasis>
</taxonomicName>
and 
<emphasis id="B93BEAD2BB0CFFB0F0C9FA6FFE56F397" box="[151,462,1485,1511]" italics="true" pageId="11" pageNumber="292">
<taxonomicName id="4C4F4D43BB0CFFB0F0C9FA6FFED5F397" baseAuthorityName="Savchenko &amp; Krivolutskaya" baseAuthorityYear="1976" box="[151,333,1485,1511]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">Atypophthalmus</taxonomicName>
– 
<taxonomicName id="4C4F4D43BB0CFFB0F105FA6FFE56F397" box="[347,462,1485,1511]" class="Insecta" family="Limoniidae" genus="Discobola" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">Discobola</taxonomicName>
</emphasis>
lineages. Presence of character 104 (spiracular lobes reduced) and absence of character 105 (caudal end of head capsule tridentate) separates this subgenus from the clade 
<taxonomicName id="4C4F4D43BB0CFFB0F41DFA53FF54F05F" authority="(Atypophthalmus)" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="subGenus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F41DFA53FB61F07B" box="[1091,1273,1521,1547]" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
(
<emphasis id="B93BEAD2BB0CFFB0F554FA53FF5CF05F" italics="true" pageId="11" pageNumber="292">Atypophthalmus</emphasis>
)
</taxonomicName>
– 
<taxonomicName id="4C4F4D43BB0CFFB0F082F9B7FED7F05F" box="[220,335,1557,1583]" class="Insecta" family="Limoniidae" genus="Discobola" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAD2BB0CFFB0F082F9B7FED7F05F" box="[220,335,1557,1583]" italics="true" pageId="11" pageNumber="292">Discobola</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF036C0BB0CFFB0F099F99BFE97F007" blockId="11.[151,1437,153,1655]" pageId="11" pageNumber="292">
The habitats of 
<taxonomicName id="4C4F4D43BB0CFFB0F124F99BFD85F023" baseAuthorityName="Meigen" baseAuthorityYear="1818" box="[378,541,1593,1619]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="inustus" subGenus="Atypophthalmus">
<emphasis id="B93BEAD2BB0CFFB0F124F99BFE13F023" box="[378,395,1593,1619]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F1FAF99BFE2DF023" box="[420,437,1593,1619]" italics="true" pageId="11" pageNumber="292">A</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F190F998FD85F023" box="[462,541,1594,1619]" italics="true" pageId="11" pageNumber="292">inustus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4F4D43BB0CFFB0F208F99BFD42F023" authority="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė, 2020" authorityName="Podenas &amp; Podeniene &amp; Seo &amp; Kim &amp; Park &amp; Byun &amp; Kim &amp; Aukštikalnienė" authorityYear="2020" box="[598,730,1593,1619]" class="Insecta" family="Limoniidae" genus="Atypophthalmus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="jeju" status="sp. nov." subGenus="Microlimonia">
<emphasis id="B93BEAD2BB0CFFB0F208F99BFDFFF023" box="[598,615,1593,1619]" italics="true" pageId="11" pageNumber="292">A</emphasis>
. (
<emphasis id="B93BEAD2BB0CFFB0F2DEF998FD0FF023" box="[640,663,1594,1619]" italics="true" pageId="11" pageNumber="292">M</emphasis>
.) 
<emphasis id="B93BEAD2BB0CFFB0F2EEF998FD42F023" box="[688,730,1594,1619]" italics="true" pageId="11" pageNumber="292">jeju</emphasis>
</taxonomicName>
<emphasis id="B93BEAD2BB0CFFB0F2BDF99BFCA5F023" bold="true" box="[739,829,1593,1619]" pageId="11" pageNumber="292">
<taxonomicNameLabel id="A20857A9BB0CFFB0F2BDF99BFCA5F023" box="[739,829,1593,1619]" pageId="11" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar, with larvae developing under the bark of dead trees.
</paragraph>
</subSubSection>
</treatment>
</document>