304 lines
39 KiB
XML
304 lines
39 KiB
XML
<document id="9D4AAEC65E0065EEE0DAFF8FA5B9BD9A" ID-DOI="10.11646/zootaxa.3736.5.8" ID-GBIF-Dataset="c231e1f1-ac35-406c-871b-a5ad87995af9" ID-ISSN="1175-5326" ID-Zenodo-Dep="249099" ID-ZooBank="D961FDF8-B2FB-4B63-A01A-45A1978FC6D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460699311845" checkinUser="plazi" docAuthor="Giłka, Wojciech, Zakrzewska, Marta, Dominiak, Patrycja & Urbanek, Aleksandra" docDate="2013" docId="03E687CEEA1AFFFBFF0BFBD04B69FA20" docLanguage="en" docName="zt03736p586.pdf" docOrigin="Zootaxa 3736 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus congregabilis Gilka et Zakrzewska, sp. nov." docType="treatment" docVersion="9" lastPageNumber="583" masterDocId="FFDFFFB6EA16FFF5FF9CFFA6482BFFF9" masterDocTitle="Non-biting midges of the tribe Tanytarsini in Eocene amber from the Rovno region (Ukraine): a pioneer systematic study with notes on the phylogeny (Diptera: Chironomidae)" masterLastPageNumber="586" masterPageNumber="569" pageNumber="581" updateTime="1698354172026" updateUser="plazi">
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<mods:mods id="9BBF318C998B2A7E113D22E3944FC47B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8947DBA8F4D5F417C52F641E6F50E746">
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<mods:title id="D978202CB2BFBD7D6C47544D860501C2">Non-biting midges of the tribe Tanytarsini in Eocene amber from the Rovno region (Ukraine): a pioneer systematic study with notes on the phylogeny (Diptera: Chironomidae)</mods:title>
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<mods:name id="B1E651C1EE04CAC25032729B292BB17E" type="personal">
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<mods:roleTerm id="92E28664888E5E74488D3C164FBA6490">Author</mods:roleTerm>
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<mods:namePart id="0A41FAD75E3A16844F2125315405373F">Giłka, Wojciech</mods:namePart>
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</mods:name>
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<mods:roleTerm id="26364B630D72FE0CB188ECFAE56649B9">Author</mods:roleTerm>
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<mods:namePart id="7696109EE0815A862BCCB41E7A510857">Zakrzewska, Marta</mods:namePart>
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</mods:name>
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<mods:roleTerm id="1E98C2519D8532299B4BC547D553B38E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="72D2ABBB37B1F85E0B6DA7967AEC3AAD">Dominiak, Patrycja</mods:namePart>
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</mods:name>
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<mods:name id="6D7E0C8837A5DEFD7FBEC44B766421AB" type="personal">
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<mods:roleTerm id="00158B4F5E1EBD34F9255B3CDE7C3C63">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="60365A3D02070B3B3F602B07F8CCE5AB">Urbanek, Aleksandra</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="6B3CA996D18B39A8D72941179A295589">text</mods:typeOfResource>
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<mods:title id="909B003D2922C521868EB2218DBFC324">Zootaxa</mods:title>
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<mods:part id="B494F5BFA5971A77C0A08D917923E70B">
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<mods:date id="DE521FBEAAC882AE9AEE8F4D5D46FA3E">2013</mods:date>
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<mods:detail id="7625C2DAC41473C694B2A349D7664745" type="volume">
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<mods:number id="8F35EB0BFA554270E7C6CD27DD5678F9">3736</mods:number>
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<mods:number id="184021320204F51F26D1916C0C222481">5</mods:number>
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<mods:start id="4E4365652BEC4664C11A8E3D64D05BB6">569</mods:start>
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<mods:classification id="E0D365AE561D5682335D7A33D3492EDD">journal article</mods:classification>
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<mods:identifier id="BAA576C38AEDF60B7A6F2FDEFA0AEF14" type="DOI">10.11646/zootaxa.3736.5.8</mods:identifier>
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<mods:identifier id="B9D34CA44F7860958C38FFC088052A38" type="GBIF-Dataset">c231e1f1-ac35-406c-871b-a5ad87995af9</mods:identifier>
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<mods:identifier id="044262AFF34BF772FA4C948B22E7D31F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8E92471E7232BE7F213D7CC990C63482" type="Zenodo-Dep">249099</mods:identifier>
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<mods:identifier id="85953B8F8D2B86BEF296C621C2493E08" type="ZooBank">D961FDF8-B2FB-4B63-A01A-45A1978FC6D4</mods:identifier>
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<treatment id="03E687CEEA1AFFFBFF0BFBD04B69FA20" ID-DOI="http://doi.org/10.5281/zenodo.6161860" ID-GBIF-Taxon="119582680" ID-Zenodo-Dep="6161860" LSID="urn:lsid:plazi:treatment:03E687CEEA1AFFFBFF0BFBD04B69FA20" httpUri="http://treatment.plazi.org/id/03E687CEEA1AFFFBFF0BFBD04B69FA20" lastPageId="14" lastPageNumber="583" pageId="12" pageNumber="581">
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<subSubSection id="C3556553EA1AFFF9FF0BFBD04B17FB69" box="[151,828,1142,1168]" pageId="12" pageNumber="581" type="nomenclature">
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<paragraph id="8BF036D8EA1AFFF9FF0BFBD04B17FB69" blockId="12.[151,828,1142,1168]" box="[151,828,1142,1168]" pageId="12" pageNumber="581">
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<heading id="D0B881B4EA1AFFF9FF0BFBD04B17FB69" bold="true" box="[151,828,1142,1168]" fontSize="11" level="1" pageId="12" pageNumber="581" reason="1">
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<emphasis id="B93BEACAEA1AFFF9FF0BFBD04B17FB69" bold="true" box="[151,828,1142,1168]" pageId="12" pageNumber="581">
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<taxonomicName id="4C4F4D5BEA1AFFF9FF0BFBD04AFAFB69" authority="Gilka et Zakrzewska" authorityName="Gilka et Zakrzewska" box="[151,721,1142,1168]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="species" species="congregabilis" status="sp. nov.">
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<emphasis id="B93BEACAEA1AFFF9FF0BFBD049EEFB69" bold="true" box="[151,453,1142,1168]" italics="true" pageId="12" pageNumber="581">Tanytarsus congregabilis</emphasis>
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Giłka
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<emphasis id="B93BEACAEA1AFFF9FD86FBD04A04FB69" bold="true" box="[538,559,1142,1168]" italics="true" pageId="12" pageNumber="581">et</emphasis>
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Zakrzewska
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20857B1EA1AFFF9FD42FBD04B17FB69" box="[734,828,1142,1168]" pageId="12" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556553EA1AFFF9FF0BFB1A49C3FAB8" pageId="12" pageNumber="581" type="materials_examined">
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<paragraph id="8BF036D8EA1AFFF9FF0BFB1A49C3FAB8" blockId="12.[151,1437,1212,2002]" pageId="12" pageNumber="581">
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<emphasis id="B93BEACAEA1AFFF9FF0BFB1A4963FB2C" bold="true" box="[151,328,1212,1237]" pageId="12" pageNumber="581">
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<typeStatus id="54F4887AEA1AFFF9FF0BFB1A48F9FB2C" box="[151,210,1212,1237]" pageId="12" pageNumber="581">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54F4887AEA1AFFF9FECEFB1A4995FB2C" box="[338,446,1212,1237]" pageId="12" pageNumber="581" type="holotype">Holotype</typeStatus>
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. Adult male (tarsus of left fore leg missing, tarsi of hind legs broken and separated).
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<typeStatus id="54F4887AEA1AFFF9FF0BFB474923FB00" box="[151,264,1249,1273]" pageId="12" pageNumber="581" type="paratype">Paratypes</typeStatus>
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. Adult male (left hind leg and wings poorly observable); adult male (tarsi of all legs missing and/or unobservable, distal abdominal segments missing). Three syninclusions preserved in 15 x
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<date id="FFF11018EA1AFFF9FB2DFAA34CD5FAE4" box="[1201,1278,1285,1309]" pageId="12" pageNumber="581" value="2010-10-04">
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10 x
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<quantity id="4CB79B3DEA1AFFF9FB6CFAA34CD5FAE4" box="[1264,1278,1285,1309]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="581" unit="mm" value="4.0">4</quantity>
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</date>
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mm piece of amber (SIZ K-8855,
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<figureCitation id="13742A5DEA1AFFF9FE1EFA8E49EFFAB8" box="[386,452,1320,1345]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="12" pageNumber="581">Fig. 7</figureCitation>
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A).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556553EA1AFFF9FF5BFAEA499CFA70" pageId="12" pageNumber="581" type="etymology">
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<paragraph id="8BF036D8EA1AFFF9FF5BFAEA499CFA70" blockId="12.[151,1437,1212,2002]" pageId="12" pageNumber="581">
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<emphasis id="B93BEACAEA1AFFF9FF5BFAEA4988FA9C" bold="true" box="[199,419,1356,1381]" pageId="12" pageNumber="581">Derivatio nominis.</emphasis>
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From Latin, meaning ‘gregarious’. We presume that the specimens examined were trapped in resin during swarming.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556553EA1AFFF9FF5BFA32490DF998" pageId="12" pageNumber="581" type="diagnosis">
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<paragraph id="8BF036D8EA1AFFF9FF5BFA32490DF998" blockId="12.[151,1437,1212,2002]" pageId="12" pageNumber="581">
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<emphasis id="B93BEACAEA1AFFF9FF5BFA32496AFA54" bold="true" box="[199,321,1428,1453]" pageId="12" pageNumber="581">Diagnosis.</emphasis>
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Femur of mid leg longer than femur of hind leg. Gonostylus stout, slightly curved at mid length, as long as gonocoxite, in distal half with several setae placed at median margin and directed anteriorly. Median setae on anal tergite absent. Anal point triangular, tapering to blunt apex; in lateral aspect visible as straight prolongation of anal tergite, with tip bent down. Superior volsella stout, bean-shaped, with broadly rounded apex, directed posteromedially. Stem of median volsella bulb-shaped, with slender leaf-shaped lamellae. Inferior volsella robust, club-shaped.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556553EA1AFFFBFF5BF9CA4AABFC6E" lastPageId="14" lastPageNumber="583" pageId="12" pageNumber="581" type="description">
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<paragraph id="8BF036D8EA1AFFF9FF5BF9CA4CE9F97D" blockId="12.[151,1437,1212,2002]" box="[199,1218,1644,1669]" pageId="12" pageNumber="581">
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<emphasis id="B93BEACAEA1AFFF9FF5BF9CA49CEF97C" bold="true" box="[199,485,1644,1669]" pageId="12" pageNumber="581">Description. Adult male</emphasis>
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(n = 1–3). Total length
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<quantity id="4CB79B3DEA1AFFF9FD73F9CB4B52F97D" box="[751,889,1645,1669]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="12" pageNumber="581" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.7–3.2 mm</quantity>
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; wing length 1620–1680 Μm.
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</paragraph>
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<paragraph id="8BF036D8EA1AFFF9FF5BF9364C58F908" blockId="12.[151,1437,1212,2002]" pageId="12" pageNumber="581">
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<emphasis id="B93BEACAEA1AFFF9FF5BF936492FF950" box="[199,260,1680,1705]" italics="true" pageId="12" pageNumber="581">Head</emphasis>
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(
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<figureCitation id="13742A5DEA1AFFF9FE88F9364973F951" box="[276,344,1680,1705]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="12" pageNumber="581">Fig. 7</figureCitation>
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B, C). Eyes bare, with dorsomedian extensions. Frontal tubercles not observed. Antenna with 13 well discernible flagellomeres, AR 1.05–1.07, plume fully developed (
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<figureCitation id="13742A5DEA1AFFF9FC26F9124BD5F934" box="[954,1022,1716,1741]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="12" pageNumber="581">Fig. 7</figureCitation>
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C). Length of palpomeres 2–5 (Μm): 45–48, 112–141, 97–141, 179–193. Clypeals present but uncountable in lateral position.
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</paragraph>
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<paragraph id="8BF036D8EA1AFFF9FF5BF95A4C8EF8EC" blockId="12.[151,1437,1212,2002]" box="[199,1189,1788,1813]" pageId="12" pageNumber="581">
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<emphasis id="B93BEACAEA1AFFF9FF5BF95A4933F8EC" box="[199,280,1788,1813]" italics="true" pageId="12" pageNumber="581">Thorax</emphasis>
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(
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<figureCitation id="13742A5DEA1AFFF9FEBBF95A4941F8EC" box="[295,362,1788,1813]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="12" pageNumber="581">Fig. 7</figureCitation>
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B). Ac at least 8, Dc 5 on each side, Pa 3 on each side, Scts unobservable.
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</paragraph>
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<paragraph id="8BF036D8EA1AFFF9FF5BF884494BF82B" blockId="12.[151,1437,1212,2002]" pageId="12" pageNumber="581">
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<emphasis id="B93BEACAEA1AFFF9FF5BF88448D4F8C0" box="[199,255,1826,1849]" italics="true" pageId="12" pageNumber="581">Wing</emphasis>
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(
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<figureCitation id="13742A5DEA1AFFF9FE8CF886497FF8C0" box="[272,340,1824,1849]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="12" pageNumber="581">Fig. 7</figureCitation>
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D). Ellipse-shaped, with weak anal lobe, broadest at 2/3 length, width: 410–430 Μm, length/width ratio 3.77–4.10. FCu placed distally of RM; VR Cu 1.16–1.18. Veins ending as in most extant
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<taxonomicName id="4C4F4D5BEA1AFFF9FB3FF8E04D36F8A4" box="[1187,1309,1862,1885]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="genus">
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<emphasis id="B93BEACAEA1AFFF9FB3FF8E04D36F8A4" box="[1187,1309,1862,1885]" italics="true" pageId="12" pageNumber="581">Tanytarsus</emphasis>
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</taxonomicName>
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(from base to tip): An, Sc, Cu1, R1, R2+3, M3+4, R4+5, M1+2; ends of R1, R2+3, R4+5 equidistant (VR C 1.0). Wing membrane densely covered with macrotrichia in distal part of r4+5, m1+2, and at posterior margin of wing. Proximal section of Cu1 slightly deflected.
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</paragraph>
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<caption id="DF306650EA1BFFF8FF0BF8314BFBF812" httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="13" pageNumber="582" targetBox="[242,1339,155,1922]" targetPageId="13">
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<paragraph id="8BF036D8EA1BFFF8FF0BF8314BFBF812" blockId="13.[151,1436,1943,2027]" pageId="13" pageNumber="582">
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<emphasis id="B93BEACAEA1BFFF8FF0BF8314938F854" bold="true" box="[151,275,1943,1965]" pageId="13" pageNumber="582">FIGURE 7.</emphasis>
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<taxonomicName id="4C4F4D5BEA1BFFF8FE86F83F4A32F854" box="[282,537,1943,1966]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="582" phylum="Arthropoda" rank="species" species="congregabilis" status="sp. nov.">
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<emphasis id="B93BEACAEA1BFFF8FE86F83F4A32F854" box="[282,537,1943,1966]" italics="true" pageId="13" pageNumber="582">Tanytarsus congregabilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857B1EA1BFFF8FD83F83E4A5BF854" box="[543,624,1944,1965]" pageId="13" pageNumber="582" rank="species">
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<emphasis id="B93BEACAEA1BFFF8FD83F83E4A5BF854" bold="true" box="[543,624,1944,1965]" pageId="13" pageNumber="582">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, male.
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<emphasis id="B93BEACAEA1BFFF8FD20F8314AC3F855" bold="true" box="[700,744,1943,1964]" pageId="13" pageNumber="582">A—</emphasis>
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type series: holotype and two paratypes (syninclusions);
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<emphasis id="B93BEACAEA1BFFF8FAB5F83E4D78F854" bold="true" box="[1321,1363,1944,1965]" pageId="13" pageNumber="582">B—</emphasis>
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habitus (holotype);
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<emphasis id="B93BEACAEA1BFFF8FE92F8104911F835" bold="true" box="[270,314,1974,1996]" pageId="13" pageNumber="582">C—</emphasis>
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antenna;
|
||
<emphasis id="B93BEACAEA1BFFF8FE0BF81149E8F835" bold="true" box="[407,451,1975,1996]" pageId="13" pageNumber="582">D—</emphasis>
|
||
wing;
|
||
<emphasis id="B93BEACAEA1BFFF8FD9FF8114A68F835" bold="true" box="[515,579,1975,1996]" pageId="13" pageNumber="582">E–I—</emphasis>
|
||
hypopygium:
|
||
<emphasis id="B93BEACAEA1BFFF8FD4EF8114AD7F835" bold="true" box="[722,764,1975,1996]" pageId="13" pageNumber="582">E—</emphasis>
|
||
lateral aspect;
|
||
<emphasis id="B93BEACAEA1BFFF8FC13F8114BB7F835" bold="true" box="[911,924,1975,1996]" pageId="13" pageNumber="582">F</emphasis>
|
||
,
|
||
<emphasis id="B93BEACAEA1BFFF8FC36F8104BFCF835" bold="true" box="[938,983,1974,1996]" pageId="13" pageNumber="582">G—</emphasis>
|
||
ventral aspect;
|
||
<emphasis id="B93BEACAEA1BFFF8FBEEF8114CB4F835" bold="true" box="[1138,1183,1975,1996]" pageId="13" pageNumber="582">H—</emphasis>
|
||
distal part of gonostylus;
|
||
<emphasis id="B93BEACAEA1BFFF8FF0BF8734891F813" bold="true" box="[151,186,2005,2026]" pageId="13" pageNumber="582">I—</emphasis>
|
||
median volsella (
|
||
<emphasis id="B93BEACAEA1BFFF8FEF4F8734957F813" bold="true" box="[360,380,2005,2026]" pageId="13" pageNumber="582">H</emphasis>
|
||
,
|
||
<emphasis id="B93BEACAEA1BFFF8FE15F8734987F813" bold="true" box="[393,428,2005,2026]" pageId="13" pageNumber="582">I—</emphasis>
|
||
magnified three times relative to hypopygium drawn).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF036D8EA18FFFBFF5BFF3F4930FEBA" blockId="14.[151,1436,151,323]" pageId="14" pageNumber="583">
|
||
<emphasis id="B93BEACAEA18FFFBFF5BFF3F48D7FF49" box="[199,252,153,176]" italics="true" pageId="14" pageNumber="583">Legs</emphasis>
|
||
. Tibia of fore leg with single, 32–37 Μm long spur (single comb composed of 6 teeth up to 15 Μm long observed in one specimen is recognized as artefact). Tibial combs of mid and hind legs separated, fan-shaped, teeth up to 15 Μm long (mid leg) and c. 15–20 Μm long (hind leg); each comb with slender spur, 20–30 Μm long (mid leg) and 40–48 Μm long (hind leg). Sensilla chaetica on ta1 of p2 not observed. For lengths of leg segments and leg ratios see
|
||
<tableCitation id="C6CD0363EA18FFFBFF5EFE8C493CFEBA" box="[194,279,298,323]" captionStart="TABLE 6" captionStartId="14.[151,239,370,393]" captionTargetBox="[159,1377,421,578]" captionTargetPageId="14" captionText="TABLE 6. Leg segment lengths (µm) and leg ratios of male Tanytarsus congregabilis sp. nov." httpUri="http://table.plazi.org/id/DF306650EA18FFFBFF0BFED44CADFE70" pageId="14" pageNumber="583" tableUuid="DF306650EA18FFFBFF0BFED44CADFE70">Table 6</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF306650EA18FFFBFF0BFED44CADFE70" ID-Table-UUID="DF306650EA18FFFBFF0BFED44CADFE70" box="[151,1158,370,393]" httpUri="http://table.plazi.org/id/DF306650EA18FFFBFF0BFED44CADFE70" pageId="14" pageNumber="583" targetBox="[159,1377,421,578]" targetIsTable="true" targetPageId="14">
|
||
<paragraph id="8BF036D8EA18FFFBFF0BFED44CADFE70" blockId="14.[151,1158,370,393]" box="[151,1158,370,393]" pageId="14" pageNumber="583">
|
||
<emphasis id="B93BEACAEA18FFFBFF0BFED44929FE70" bold="true" box="[151,258,370,393]" pageId="14" pageNumber="583">TABLE 6</emphasis>
|
||
. Leg segment lengths (µm) and leg ratios of male
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFC81FED44C02FE70" box="[797,1065,370,393]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="congregabilis" status="sp. nov.">
|
||
<emphasis id="B93BEACAEA18FFFBFC81FED44C02FE70" box="[797,1065,370,393]" italics="true" pageId="14" pageNumber="583">Tanytarsus congregabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20857B1EA18FFFBFBACFED44CADFE70" box="[1072,1158,370,393]" pageId="14" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF036D8EA18FFFBFF4BFE034D4AFDC6" pageId="14" pageNumber="583">
|
||
<table id="F94FC478EA18000AFF03FE034D4AFDBB" box="[159,1377,421,578]" gridcols="8" gridrows="4" pageId="14" pageNumber="583">
|
||
<tr id="357F349AEA18000AFF03FE034D4AFE46" box="[159,1377,421,447]" gridrow="0" pageId="14" pageNumber="583">
|
||
<th id="76AE5DE6EA18000AFF03FE034904FE46" box="[159,303,421,447]" gridcol="0" gridrow="0" pageId="14" pageNumber="583">fe</th>
|
||
<th id="76AE5DE6EA18000AFEF2FE0349ECFE46" box="[366,455,421,447]" gridcol="1" gridrow="0" pageId="14" pageNumber="583">ti</th>
|
||
<th id="76AE5DE6EA18000AFD9AFE034A74FE46" box="[518,607,421,447]" gridcol="2" gridrow="0" pageId="14" pageNumber="583">ta1</th>
|
||
<th id="76AE5DE6EA18000AFD01FE034ADDFE46" box="[669,758,421,447]" gridcol="3" gridrow="0" pageId="14" pageNumber="583">ta2</th>
|
||
<th id="76AE5DE6EA18000AFCA9FE034BA5FE46" box="[821,910,421,447]" gridcol="4" gridrow="0" pageId="14" pageNumber="583">ta3</th>
|
||
<th id="76AE5DE6EA18000AFC50FE034C0EFE46" box="[972,1061,421,447]" gridcol="5" gridrow="0" pageId="14" pageNumber="583">ta4</th>
|
||
<th id="76AE5DE6EA18000AFBF8FE034C97FE46" box="[1124,1212,421,447]" gridcol="6" gridrow="0" pageId="14" pageNumber="583">ta5</th>
|
||
<th id="76AE5DE6EA18000AFB67FE034D4AFE46" box="[1275,1377,421,447]" gridcol="7" gridrow="0" pageId="14" pageNumber="583">LR</th>
|
||
</tr>
|
||
<tr id="357F349AEA18000AFF03FE774D4AFE12" box="[159,1377,465,491]" gridrow="1" pageId="14" pageNumber="583">
|
||
<th id="76AE5DE6EA18000AFF03FE774904FE12" box="[159,303,465,491]" gridcol="0" gridrow="1" pageId="14" pageNumber="583">P1 715–805</th>
|
||
<td id="76AE5DE6EA18000AFEF2FE7749ECFE12" box="[366,455,465,491]" gridcol="1" gridrow="1" pageId="14" pageNumber="583">590–660</td>
|
||
<td id="76AE5DE6EA18000AFD9AFE774A74FE12" box="[518,607,465,491]" gridcol="2" gridrow="1" pageId="14" pageNumber="583">775–795</td>
|
||
<td id="76AE5DE6EA18000AFD01FE774ADDFE12" box="[669,758,465,491]" gridcol="3" gridrow="1" pageId="14" pageNumber="583">460–510</td>
|
||
<td id="76AE5DE6EA18000AFCA9FE774BA5FE12" box="[821,910,465,491]" gridcol="4" gridrow="1" pageId="14" pageNumber="583">365–415</td>
|
||
<td id="76AE5DE6EA18000AFC50FE774C0EFE12" box="[972,1061,465,491]" gridcol="5" gridrow="1" pageId="14" pageNumber="583">280–285</td>
|
||
<td id="76AE5DE6EA18000AFBF8FE774C97FE12" box="[1124,1212,465,491]" gridcol="6" gridrow="1" pageId="14" pageNumber="583">115–125</td>
|
||
<td id="76AE5DE6EA18000AFB67FE774D4AFE12" box="[1275,1377,465,491]" gridcol="7" gridrow="1" pageId="14" pageNumber="583">1.28–1.34</td>
|
||
</tr>
|
||
<tr id="357F349AEA18000AFF03FE5B4D4AFDEF" box="[159,1377,509,534]" gridrow="2" pageId="14" pageNumber="583">
|
||
<th id="76AE5DE6EA18000AFF03FE5B4904FDEF" box="[159,303,509,534]" gridcol="0" gridrow="2" pageId="14" pageNumber="583">P2 765–780</th>
|
||
<td id="76AE5DE6EA18000AFEF2FE5B49ECFDEF" box="[366,455,509,534]" gridcol="1" gridrow="2" pageId="14" pageNumber="583">720–780</td>
|
||
<td id="76AE5DE6EA18000AFD9AFE5B4A74FDEF" box="[518,607,509,534]" gridcol="2" gridrow="2" pageId="14" pageNumber="583">425–460</td>
|
||
<td id="76AE5DE6EA18000AFD01FE5B4ADDFDEF" box="[669,758,509,534]" gridcol="3" gridrow="2" pageId="14" pageNumber="583">245–290</td>
|
||
<td id="76AE5DE6EA18000AFCA9FE5B4BA5FDEF" box="[821,910,509,534]" gridcol="4" gridrow="2" pageId="14" pageNumber="583">180–215</td>
|
||
<td id="76AE5DE6EA18000AFC50FE5B4C0EFDEF" box="[972,1061,509,534]" gridcol="5" gridrow="2" pageId="14" pageNumber="583">110–140</td>
|
||
<td id="76AE5DE6EA18000AFBF8FE5B4C97FDEF" box="[1124,1212,509,534]" gridcol="6" gridrow="2" pageId="14" pageNumber="583">75–90</td>
|
||
<td id="76AE5DE6EA18000AFB67FE5B4D4AFDEF" box="[1275,1377,509,534]" gridcol="7" gridrow="2" pageId="14" pageNumber="583">0.55–0.60</td>
|
||
</tr>
|
||
<tr id="357F349AEA18000AFF03FD8F4D4AFDBB" box="[159,1377,553,578]" gridrow="3" pageId="14" pageNumber="583">
|
||
<th id="76AE5DE6EA18000AFF03FD8F4904FDBB" box="[159,303,553,578]" gridcol="0" gridrow="3" pageId="14" pageNumber="583">P3 700–715</th>
|
||
<td id="76AE5DE6EA18000AFEF2FD8F49ECFDBB" box="[366,455,553,578]" gridcol="1" gridrow="3" pageId="14" pageNumber="583">910–915</td>
|
||
<td id="76AE5DE6EA18000AFD9AFD8F4A74FDBB" box="[518,607,553,578]" gridcol="2" gridrow="3" pageId="14" pageNumber="583">515–555</td>
|
||
<td id="76AE5DE6EA18000AFD01FD8F4ADDFDBB" box="[669,758,553,578]" gridcol="3" gridrow="3" pageId="14" pageNumber="583">335–365</td>
|
||
<td id="76AE5DE6EA18000AFCA9FD8F4BA5FDBB" box="[821,910,553,578]" gridcol="4" gridrow="3" pageId="14" pageNumber="583">260–290</td>
|
||
<td id="76AE5DE6EA18000AFC50FD8F4C0EFDBB" box="[972,1061,553,578]" gridcol="5" gridrow="3" pageId="14" pageNumber="583">150–170</td>
|
||
<td id="76AE5DE6EA18000AFBF8FD8F4C97FDBB" box="[1124,1212,553,578]" gridcol="6" gridrow="3" pageId="14" pageNumber="583">90–110</td>
|
||
<td id="76AE5DE6EA18000AFB67FD8F4D4AFDBB" box="[1275,1377,553,578]" gridcol="7" gridrow="3" pageId="14" pageNumber="583">0.57–0.61</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF036D8EA18FFFBFF5BFD254AABFC6E" blockId="14.[151,1437,642,1498]" pageId="14" pageNumber="583">
|
||
<emphasis id="B93BEACAEA18FFFBFF5BFD25497DFD63" box="[199,342,643,666]" italics="true" pageId="14" pageNumber="583">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="13742A5DEA18FFFBFEFAFD244981FD63" box="[358,426,642,667]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">Fig. 7</figureCitation>
|
||
E–I). Gonostylus stout, slightly curved at mid length, boomerang-like, c. 115–125 Μm, as long as gonocoxite, in distal half with several setae placed at median margin and directed anteriorly (
|
||
<figureCitation id="13742A5DEA18FFFBFA8AFD004D71FD46" box="[1302,1370,678,703]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">Fig. 7</figureCitation>
|
||
F–H). Median setae on anal tergite absent. Anal point triangular, tapering to blunt apex; in lateral aspect visible as straight prolongation of anal tergite, tip bent down (
|
||
<figureCitation id="13742A5DEA18FFFBFD12FD484AF8FCFE" box="[654,723,750,775]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">Fig. 7</figureCitation>
|
||
E); spinulae, crests and posterolateral setae unobservable from ventral/lateral side. Superior volsella stout, bean-shaped, with broadly rounded apex, directed posteromedially. Digitus not observed (
|
||
<figureCitation id="13742A5DEA18FFFBFE05FC9049F6FCB7" box="[409,477,822,847]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">Fig. 7</figureCitation>
|
||
F, G). Stem of median volsella c. 25 Μm long, bulb-shaped, with 5–6 slender leafshaped lamellae (
|
||
<figureCitation id="13742A5DEA18FFFBFEC1FCFC4988FC8B" box="[349,419,858,883]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">
|
||
Fig.
|
||
<date id="FFF11018EA18FFFBFE0FFCFC4988FC8B" box="[403,419,858,882]" pageId="14" pageNumber="583">7</date>
|
||
</figureCitation>
|
||
I). Inferior volsella reaching 1/3 length of gonostylus at most, robust, club-shaped, armed with several stout curved setae (
|
||
<figureCitation id="13742A5DEA18FFFBFE62FCD84A14FC6F" box="[510,575,894,919]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">Fig. 7</figureCitation>
|
||
F, G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556553EA18FFFBFF5BFC044B69FA20" pageId="14" pageNumber="583" type="discussion">
|
||
<paragraph id="8BF036D8EA18FFFBFF5BFC044DB7FB4E" blockId="14.[151,1437,642,1498]" pageId="14" pageNumber="583">
|
||
<emphasis id="B93BEACAEA18FFFBFF5BFC044910FC42" bold="true" box="[199,315,930,955]" pageId="14" pageNumber="583">Remarks.</emphasis>
|
||
The structure of male hypopygium of
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFC98FC054C0AFC42" box="[772,1057,930,955]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="congregabilis">
|
||
<emphasis id="B93BEACAEA18FFFBFC98FC054C0AFC42" box="[772,1057,930,955]" italics="true" pageId="14" pageNumber="583">Tanytarsus congregabilis</emphasis>
|
||
</taxonomicName>
|
||
suggests inclusion in the
|
||
<emphasis id="B93BEACAEA18FFFBFACFFC044DB7FC42" box="[1363,1436,930,955]" italics="true" pageId="14" pageNumber="583">lugens</emphasis>
|
||
species group.
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFEDBFC614A2BFC26" box="[327,512,966,991]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="congregabilis">
|
||
<emphasis id="B93BEACAEA18FFFBFEDBFC614A2BFC26" box="[327,512,966,991]" italics="true" pageId="14" pageNumber="583">T. congregabilis</emphasis>
|
||
</taxonomicName>
|
||
fits the group diagnosis (Reiss & Fittkau 1971, Giłka & Paasivirta 2007) and displays the following character combination: gonostylus stout; median setae on anal tergite absent; superior volsella bean-shaped, with broadly rounded apex, directed posteromedially; digitus not observed; stem of median volsella bulb-shaped, bearing leaf-shaped lamellae (
|
||
<figureCitation id="13742A5DEA18FFFBFC99FB944B7BFBB3" box="[773,848,1074,1099]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">Fig. 7</figureCitation>
|
||
E–I). Unfortunately, the specimens examined are positioned such as to render observation of the dorsal aspect of hypopygium impossible, but the shape of the anal point in lateral view is similar to that known from
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFD50FBDD4C03FB6A" authority="Kieffer" authorityName="Kieffer" box="[716,1064,1146,1171]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="bathophilus">
|
||
<emphasis id="B93BEACAEA18FFFBFD50FBDD4BF9FB6A" box="[716,978,1146,1171]" italics="true" pageId="14" pageNumber="583">Tanytarsus bathophilus</emphasis>
|
||
Kieffer
|
||
</taxonomicName>
|
||
(Reiss & Fittkau 1971, fig. 36a). The peculiar, boomerang-shaped gonostylus is the best character for rapid identification of the species (
|
||
<figureCitation id="13742A5DEA18FFFBFA84FB384D71FB4E" box="[1304,1370,1182,1207]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1965]" captionTargetBox="[242,1339,155,1922]" captionTargetId="figure@13.[242,1340,155,1922]" captionTargetPageId="13" captionText="FIGURE 7. Tanytarsus congregabilis sp. nov., male. A— type series: holotype and two paratypes (syninclusions); B— habitus (holotype); C— antenna; D— wing; E – I— hypopygium: E— lateral aspect; F, G— ventral aspect; H— distal part of gonostylus; I— median volsella (H, I— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249106/files/figure.png" pageId="14" pageNumber="583">Fig. 7</figureCitation>
|
||
F–H).
|
||
</paragraph>
|
||
<paragraph id="8BF036D8EA18FFFBFF5BFB644B69FA20" blockId="14.[151,1437,642,1498]" pageId="14" pageNumber="583">
|
||
The large size (the largest among the known Eocene
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFCBCFB644BB4FB22" box="[800,927,1218,1243]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="tribe" tribe="Tanytarsini">Tanytarsini</taxonomicName>
|
||
) of adult
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFB92FB654D02FB22" box="[1038,1321,1218,1243]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="congregabilis">
|
||
<emphasis id="B93BEACAEA18FFFBFB92FB654D02FB22" box="[1038,1321,1218,1243]" italics="true" pageId="14" pageNumber="583">Tanytarsus congregabilis</emphasis>
|
||
</taxonomicName>
|
||
males are also consistent with the
|
||
<emphasis id="B93BEACAEA18FFFBFE3AFB4049C4FB06" box="[422,495,1254,1279]" italics="true" pageId="14" pageNumber="583">lugens</emphasis>
|
||
group diagnosis, although
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFCBEFB414BFCFB06" box="[802,983,1254,1279]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="congregabilis">
|
||
<emphasis id="B93BEACAEA18FFFBFCBEFB414BFCFB06" box="[802,983,1254,1279]" italics="true" pageId="14" pageNumber="583">T. congregabilis</emphasis>
|
||
</taxonomicName>
|
||
is one of the smallest representatives of the group. We observed a similar pattern in other families of dipterans of the suborder Nematocera which, in the Baltic amber, are represented by specimens smaller than those belonging to related extant taxa (
|
||
<emphasis id="B93BEACAEA18FFFBFB77FA884D2CFABF" box="[1259,1287,1326,1350]" italics="true" pageId="14" pageNumber="583">cf.</emphasis>
|
||
Szadziewski 1988). The nature of the resin trap, enclosing primarily small insects, does not seem to particularly affect the
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFF0BFAD0493CFA76" box="[151,279,1398,1423]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="tribe" tribe="Tanytarsini">Tanytarsini</taxonomicName>
|
||
, as neither the fossil nor extant representatives of the tribe are larger than
|
||
<quantity id="4CB79B3DEA18FFFBFBC2FAD04C89FA77" box="[1118,1186,1398,1423]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="583" unit="mm" value="6.0">6 mm</quantity>
|
||
. Our hypothesis gains an additional support from the fact that the three
|
||
<taxonomicName id="4C4F4D5BEA18FFFBFC99FA3D4BE9FA4A" box="[773,962,1434,1459]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="congregabilis">
|
||
<emphasis id="B93BEACAEA18FFFBFC99FA3D4BE9FA4A" box="[773,962,1434,1459]" italics="true" pageId="14" pageNumber="583">T. congregabilis</emphasis>
|
||
</taxonomicName>
|
||
specimens examined in this study are syninclusions in an amber chunk slightly larger than
|
||
<quantity id="4CB79B3DEA18FFFBFD7BFA674B18FA20" box="[743,819,1473,1498]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="583" unit="cm" value="0.5">0.5 cm</quantity>
|
||
3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |