169 lines
19 KiB
XML
169 lines
19 KiB
XML
<document id="82F7DBF761A76BC05C4973EF2BB3F787" ID-CLB-Dataset="300555" ID-DOI="10.5281/zenodo.13175056" ID-GBIF-Dataset="347efc47-0aab-4629-98e4-be1034c9fc21" ID-ISSN="1732-2421" ID-Zenodo-Dep="13175056" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1721349230000" checkinUser="felipe" docAuthor="Feist, Raimund" docDate="2002" docId="03D887C9FF88FF85FCDCFCB38FB3FE62" docLanguage="en" docName="ActaPalaeontolPol.47.2.203-210.pdf" docOrigin="Acta Palaeontologica Polonica 47 (2)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Pteroparia ziegleri subsp. maroccanica Feist, 2002, subsp. nov." docType="treatment" docVersion="3" lastPageNumber="206" masterDocId="FFE1FFB1FF8BFF80FFF0FFE28D62FF99" masterDocTitle="Trilobites from the latest Frasnian Kellwasser Crisis in North Africa (Mrirt, central Moroccan Meseta)" masterLastPageNumber="210" masterPageNumber="203" pageNumber="206" updateTime="1722623223228" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="EB6567E19004883567E6E0FDCCBA6490">
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<mods:title id="51DC85509BE45A8E27A0E1D92B0648DD">Trilobites from the latest Frasnian Kellwasser Crisis in North Africa (Mrirt, central Moroccan Meseta)</mods:title>
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<mods:name id="B116345DB35CEEAF5136F9FA9720A749" type="personal">
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<mods:roleTerm id="09D2C6F86C2F1A7F26F74B1E982A2EFD">Author</mods:roleTerm>
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<mods:namePart id="4013DB201FD88ACB009C79A73DC4EB4D">Feist, Raimund</mods:namePart>
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<mods:typeOfResource id="094B2E07CE15AABFDF0CECFBA6801F1F">text</mods:typeOfResource>
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<mods:title id="F45F313A667A13F36CACAF82C722771D">Acta Palaeontologica Polonica</mods:title>
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</mods:titleInfo>
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<mods:part id="652D5DF712A02E3C5E39EDEDCDE37BAD">
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<mods:date id="1D5C8EB5DA5F4749BD1D97A7F2255826">2002</mods:date>
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<mods:number id="099F6D52EDDD2D0CAF6B5B3A85C06E2E">2002-12-31</mods:number>
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<mods:detail id="263440BED26AFF2C9F7724F148DF2EDD" type="volume">
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<mods:number id="7DB1D7FDB892BCEA2FE562FEB554287D">47</mods:number>
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<mods:detail id="1C8D466183012FDC9FD0C0B0330387AF" type="issue">
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<mods:number id="CBD46042C93A68FD0CDC5152FADE7A83">2</mods:number>
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<mods:start id="F22FF3C22EC2A81B82E52E57424661D8">203</mods:start>
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<mods:end id="2C832413A507C5164C794A6246C2972D">210</mods:end>
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<mods:classification id="D8B9503030C905C9A11F25CAD65F371A">journal article</mods:classification>
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<mods:identifier id="27F19AB2587144B48E7DAB8EC0802696" type="DOI">10.5281/zenodo.13175056</mods:identifier>
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<mods:identifier id="714FDFBDAAAB60C9F91C1BEB8BCA6DCB" type="ISSN">1732-2421</mods:identifier>
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<mods:identifier id="902104AAD3BDB318E024CF91AD8E30A9" type="Zenodo-Dep">13175056</mods:identifier>
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<treatment id="03D887C9FF88FF85FCDCFCB38FB3FE62" ID-DOI="http://doi.org/10.5281/zenodo.13175074" ID-Zenodo-Dep="13175074" LSID="urn:lsid:plazi:treatment:03D887C9FF88FF85FCDCFCB38FB3FE62" httpUri="http://treatment.plazi.org/id/03D887C9FF88FF85FCDCFCB38FB3FE62" lastPageId="5" lastPageNumber="206" pageId="3" pageNumber="206">
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<subSubSection id="C36B6554FF88FF83FCDCFCB38806FCF5" box="[812,1380,848,877]" pageId="3" pageNumber="206" type="nomenclature">
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<paragraph id="8BCE36DFFF88FF83FCDCFCB38806FCF5" blockId="3.[812,1380,664,912]" box="[812,1380,848,877]" pageId="3" pageNumber="206">
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<heading id="D08681B3FF88FF83FCDCFCB38806FCF5" box="[812,1380,848,877]" fontSize="36" level="2" pageId="3" pageNumber="206" reason="7">
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<taxonomicName id="4C714D5CFF88FF83FCDCFCB389AFFCF4" authorityName="Feist" authorityYear="2002" box="[812,1229,848,877]" class="Trilobita" family="Tropidocoryphidae" genus="Pteroparia" kingdom="Animalia" order="Proetida" pageId="3" pageNumber="206" phylum="Arthropoda" rank="subSpecies" species="ziegleri" status="subsp. nov." subSpecies="maroccanica">
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<emphasis id="B905EACDFF88FF83FCDCFCB389AFFCF4" box="[812,1229,848,877]" italics="true" pageId="3" pageNumber="206">Pteroparia ziegleri maroccanica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23657B6FF88FF83FB25FCB28806FCF5" box="[1237,1380,848,877]" pageId="3" pageNumber="206" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6554FF88FF83FCDCFC9889E8FC2E" pageId="3" pageNumber="206" type="description">
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<paragraph id="8BCE36DFFF88FF83FCDCFC988EF2FC09" blockId="3.[812,1380,664,912]" box="[812,912,890,912]" pageId="3" pageNumber="206">
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<figureCitation id="134A2A5AFF88FF83FCDCFC988EEFFC09" box="[812,909,890,912]" captionStart="Fig" captionStartId="5.[102,135,1862,1881]" captionTargetId="figure-8@6.[102,1485,233,1913]" captionTargetPageId="6" captionText="Fig. 3. A–I. Acuticryphops acuticeps (Kayser, 1889). A, H, I. Cephalon from section “2”, upper trilobite association, UM2−RF 220, partially external mould, in dorsal view (A), × 6.9; in lateral view (H), eye lobe with more than 6 lenses, × 6.5; in frontal view (I), slightly tilted backward showing vincular furrow and ventral doublure, × 7.1. B, C. Cephalon from section “1”, upper trilobite association, UM2−RF 221, external mould; in lateral view (B), eye lobe with 4 lenses, × 7.8; in dorsal view (C), × 8.2. D–F. Pygidium from section “2”, upper trilobite association, UM2−RF 222, external mould; lateral view (D), × 14.5; in poste− rior view (E), × 10.7; in dorsal view (F), × 12.2. J–K. Otarion stigmatophthalmus (Richter, 1914), cranidium from section „2“, upper trilobite association, UM2−RF 248, internal mould in dorsal view (J), × 14.1; in lateral view (K), × 14.8. L–Q. Pteroparia ziegleri maroccanica subsp. nov.from section “1”, lower +" figureDoi="http://doi.org/10.5281/zenodo.13175064" httpUri="https://zenodo.org/record/13175064/files/figure.png" pageId="3" pageNumber="206">Fig. 3L–P</figureCitation>
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.
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</paragraph>
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<paragraph id="8BCE36DFFF88FF83FCDCFC4089E8FC2E" blockId="3.[812,1485,929,1150]" box="[812,1162,929,951]" pageId="3" pageNumber="206">
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<emphasis id="B905EACDFF88FF83FCDCFC408E85FC2E" box="[812,999,929,951]" italics="true" pageId="3" pageNumber="206">Derivation of name</emphasis>
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: After
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<collectingCountry id="F366764FFF88FF83FBDCFC4389E4FC2E" box="[1068,1158,929,951]" name="Morocco" pageId="3" pageNumber="206">Morocco</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6554FF88FF83FCDCFC5C89BCFA9E" pageId="3" pageNumber="206" type="materials_examined">
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<paragraph id="8BCE36DFFF88FF83FCDCFC5C8870FC4D" blockId="3.[812,1485,929,1150]" box="[812,1298,958,980]" pageId="3" pageNumber="206">
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<materialsCitation id="3B193C82FF88FF83FCDCFC5C8870FC4D" box="[812,1298,958,980]" collectionCode="UM" country="Russia" location="RF" pageId="3" pageNumber="206" specimenCode="UM2" specimenCount="1" typeStatus="holotype">
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<emphasis id="B905EACDFF88FF83FCDCFC5C8EE6FC4D" box="[812,900,958,980]" italics="true" pageId="3" pageNumber="206">
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<typeStatus id="54CA887DFF88FF83FCDCFC5C8EE6FC4D" box="[812,900,958,980]" pageId="3" pageNumber="206" type="holotype">Holotype</typeStatus>
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</emphasis>
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: Cranidium
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<specimenCode id="DBD79EA4FF88FF83FC0DFC5C894FFC4D" box="[1021,1069,958,980]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="3" pageNumber="206">UM2</specimenCode>
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−
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<collectingCountry id="F366764FFF88FF83FBC9FC5C8937FC4D" box="[1081,1109,958,980]" name="Russia" pageId="3" pageNumber="206">RF</collectingCountry>
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252, fig. 3L, M, Q.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCE36DFFF88FF83FCDCFC3E897AFBB0" blockId="3.[812,1485,929,1150]" pageId="3" pageNumber="206">
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<emphasis id="B905EACDFF88FF83FCDCFC3E8ECAFC68" box="[812,936,987,1009]" italics="true" pageId="3" pageNumber="206">Type locality</emphasis>
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: Section 1, lower trilobite association at Bou Ounabdou (section II in
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<bibRefCitation id="EFE04B2EFF88FF83FC20FC158928FB94" author="Lazreq, N." box="[976,1098,1015,1037]" pageId="3" pageNumber="206" pagination="107 - 123" refId="ref6244" refString="Lazreq, N. 1992. The Upper Devonian of M'rirt (Morocco). Courier Forschungs-Institut Senckenberg 154: 107 - 123." type="journal article" year="1992">Lazreq 1992</bibRefCitation>
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), north of track, Sof Al Gara hill (see location map
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<figureCitation id="134A2A5AFF88FF83FC32FBF18969FBB0" box="[962,1035,1043,1065]" captionStart="Fig" captionStartId="1.[977,1010,1123,1142]" captionTargetBox="[102,937,226,1392]" captionTargetId="figure-267@1.[102,938,226,1392]" captionTargetPageId="1" captionText="Fig. 1. A. Devonian outcrops in Morocco with location of the Mrirt area in the central Meseta east of Rabat. B. Measured section “1” showing the Lower (LKW) and Upper (UKW) Kellwasser horizons and the intervening limestone beds. The levels of the lower and upper trilobite associations are marked with a star. C. Location of the studied sections “1” and “2” at Bou Ounabdou, 4 km to the SE of Mrirt township (after Becker and House 1999 and Walliser et al. 1999). Section “1” is the same as Bou Ounabdou, section II of Lasreq (1992)." figureDoi="http://doi.org/10.5281/zenodo.13175058" httpUri="https://zenodo.org/record/13175058/files/figure.png" pageId="3" pageNumber="206">Fig. 1C</figureCitation>
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).
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</paragraph>
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<paragraph id="8BCE36DFFF88FF83FCDCFBD289F0FBFB" blockId="3.[812,1485,929,1150]" pageId="3" pageNumber="206">
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<emphasis id="B905EACDFF88FF83FCDCFBD28EC5FBDF" box="[812,935,1072,1094]" italics="true" pageId="3" pageNumber="206">Type horizon</emphasis>
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: Grey−beige limestone bed immediately above prominent dark grey Lower Kellwasser bed.
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</paragraph>
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<paragraph id="8BCE36DFFF88FF83FCDCFB8A88D8FBE7" blockId="3.[812,1485,929,1150]" box="[812,1466,1128,1150]" pageId="3" pageNumber="206">
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<emphasis id="B905EACDFF88FF83FCDCFB8A8E33FBE7" box="[812,849,1128,1150]" italics="true" pageId="3" pageNumber="206">Age</emphasis>
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: According to
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<bibRefCitation id="EFE04B2EFF88FF83FC12FB8A890CFBE7" author="Lazreq, N." box="[994,1134,1128,1150]" pageId="3" pageNumber="206" pagination="107 - 123" refId="ref6244" refString="Lazreq, N. 1992. The Upper Devonian of M'rirt (Morocco). Courier Forschungs-Institut Senckenberg 154: 107 - 123." type="journal article" year="1992">Lazreq (1992)</bibRefCitation>
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: Late
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<taxonomicName id="4C714D5CFF88FF83FB58FB8A88D8FBE7" authority="Zone." authorityName="Zone." box="[1192,1466,1128,1150]" class="Conodonta" family="Palmatolepidae" genus="Palmatolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="3" pageNumber="206" phylum="Chordata" rank="species" species="rhenana">
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<emphasis id="B905EACDFF88FF83FB58FB8A8819FBE7" box="[1192,1403,1128,1150]" italics="true" pageId="3" pageNumber="206">Palmatolepis rhenana</emphasis>
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Zone.
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</taxonomicName>
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</paragraph>
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<paragraph id="8BCE36DFFF88FF83FCDCFB6D89BCFA9E" blockId="3.[812,1485,1167,1287]" pageId="3" pageNumber="206">
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<emphasis id="B905EACDFF88FF83FCDCFB6D8EECFB3E" box="[812,910,1167,1191]" italics="true" pageId="3" pageNumber="206">Material</emphasis>
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.—
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<materialsCitation id="3B193C82FF88FF83FC5FFB6D88AFFB3E" box="[943,1485,1167,1191]" collectionCode="UM" country="Russia" location="RF" pageId="3" pageNumber="206" specimenCode="UM2" specimenCount="2">
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Figured specimens: 1 cranidium (
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<specimenCode id="DBD79EA4FF88FF83FAD4FB6D883FFB3E" box="[1316,1373,1167,1191]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="3" pageNumber="206">UM2</specimenCode>
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−
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<collectingCountry id="F366764FFF88FF83FA9BFB6D88EFFB3E" box="[1387,1421,1167,1191]" name="Russia" pageId="3" pageNumber="206">RF</collectingCountry>
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252),
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</materialsCitation>
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<materialsCitation id="3B193C82FF88FF83FCDCFB4D8935FB5E" box="[812,1111,1199,1223]" collectionCode="UM" country="Russia" location="RF" pageId="3" pageNumber="206" specimenCode="UM2" specimenCount="2">
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1 librigena (
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<specimenCode id="DBD79EA4FF88FF83FC5FFB4D8E8AFB5E" box="[943,1000,1199,1223]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="3" pageNumber="206">UM2</specimenCode>
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−
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<collectingCountry id="F366764FFF88FF83FC06FB4D897AFB5E" box="[1014,1048,1199,1223]" name="Russia" pageId="3" pageNumber="206">RF</collectingCountry>
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253),
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</materialsCitation>
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<materialsCitation id="3B193C82FF88FF83FBACFB4D88ECFB5E" box="[1116,1422,1199,1223]" collectionCode="UM" country="Russia" location="RF" pageId="3" pageNumber="206" specimenCode="UM2" specimenCount="2">
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1 pygidium (
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<specimenCode id="DBD79EA4FF88FF83FB16FB4D887DFB5E" box="[1254,1311,1199,1223]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="3" pageNumber="206">UM2</specimenCode>
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−
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<collectingCountry id="F366764FFF88FF83FADDFB4D882DFB5E" box="[1325,1359,1199,1223]" name="Russia" pageId="3" pageNumber="206">RF</collectingCountry>
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254);
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</materialsCitation>
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<materialsCitation id="3B193C82FF88FF83FA63FB4D8EFCFA9E" collectionCode="UM" country="Russia" location="RF" pageId="3" pageNumber="206" specimenCode="UM2" specimenCount="2">
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additional material all from type locality: 3 cranidia (
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<specimenCode id="DBD79EA4FF88FF83FA94FB2D88FFFB7E" box="[1380,1437,1231,1255]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="3" pageNumber="206">UM2</specimenCode>
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−
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<collectingCountry id="F366764FFF88FF83FA5BFB2D88AFFB7E" box="[1451,1485,1231,1255]" name="Russia" pageId="3" pageNumber="206">RF</collectingCountry>
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255–257),
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</materialsCitation>
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<materialsCitation id="3B193C82FF88FF83FC55FB0D89B8FA9E" box="[933,1242,1263,1287]" collectionCode="UM" country="Russia" location="RF" pageId="3" pageNumber="206" specimenCode="UM2" specimenCount="2">
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1 pygidium (
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<specimenCode id="DBD79EA4FF88FF83FBC4FB0D890FFA9E" box="[1076,1133,1263,1287]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="3" pageNumber="206">UM2</specimenCode>
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−
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<collectingCountry id="F366764FFF88FF83FB8BFB0D89FFFA9E" box="[1147,1181,1263,1287]" name="Russia" pageId="3" pageNumber="206">RF</collectingCountry>
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258)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6554FF88FF83FCDCFAF888E6FA08" pageId="3" pageNumber="206" type="diagnosis">
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<paragraph id="8BCE36DFFF88FF83FCDCFAF888E6FA08" blockId="3.[812,1485,1305,1425]" pageId="3" pageNumber="206">
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<emphasis id="B905EACDFF88FF83FCDCFAF88EF9FAA8" box="[812,923,1306,1329]" italics="true" pageId="3" pageNumber="206">Diagnosis</emphasis>
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.—Anterior outline of glabella semicircular; fixigenae without ocular ridges or swellings. Librigena with rather broad, stout genal spine. Axis of pygidium narrower than pleural field; postaxial area shorter than half length of axis.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6554FF88FF85FCDCFA468FB3FE62" lastPageId="5" lastPageNumber="208" pageId="3" pageNumber="206" type="discussion">
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<paragraph id="8BCE36DFFF88FF85FCDCFA468FB3FE62" blockId="3.[812,1485,1443,1819]" lastBlockId="5.[102,775,227,507]" lastPageId="5" lastPageNumber="208" pageId="3" pageNumber="206">
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<emphasis id="B905EACDFF88FF83FCDCFA468ED0FA22" box="[812,946,1444,1467]" italics="true" pageId="3" pageNumber="206">Comparison</emphasis>
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.—The Moroccan material is very near to
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<taxonomicName id="4C714D5CFF88FF83FA79FA468EDFFA42" authorityName="Feist" authorityYear="1994" class="Trilobita" family="Tropidocoryphidae" genus="Pteroparia" kingdom="Animalia" order="Proetida" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="ziegleri">
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<emphasis id="B905EACDFF88FF83FA79FA468EDFFA42" italics="true" pageId="3" pageNumber="206">Pteroparia ziegleri</emphasis>
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</taxonomicName>
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from Sessacker in the general outline and vault of the cephalon, the presence of a small, flat ocular field with lenses and the dense granulation of the exoskeleton. There are a few morphological differences that distinguish the new material that is considered here as a contemporaneous geographic subspecies of
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<taxonomicName id="4C714D5CFF88FF83FC34F986894DF9E2" authorityName="Feist" authorityYear="1994" box="[964,1071,1635,1659]" class="Trilobita" family="Tropidocoryphidae" genus="Pteroparia" kingdom="Animalia" order="Proetida" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="ziegleri">
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<emphasis id="B905EACDFF88FF83FC34F986894DF9E2" box="[964,1071,1635,1659]" italics="true" pageId="3" pageNumber="206">P. ziegleri</emphasis>
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</taxonomicName>
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. The most important feature is the absence of ocular ridges on the fixigenae opposite to the eye: this area remains virtually flush with the general curvature of the anterior enlarged fixigenal field whereas, in the nominate subspecies, it is differentiated from the latter by the swollen palpebral area. Though subject to considerable variation, the anterior outline of the glabella is broader in
|
||
<taxonomicName id="4C714D5CFF8EFF85FE16FF018F8BFF63" authorityName="Feist" authorityYear="2002" box="[486,745,227,251]" class="Trilobita" family="Tropidocoryphidae" genus="Pteroparia" kingdom="Animalia" order="Proetida" pageId="5" pageNumber="208" phylum="Arthropoda" rank="subSpecies" species="ziegleri" subSpecies="maroccanica">
|
||
<emphasis id="B905EACDFF8EFF85FE16FF018F8BFF63" box="[486,745,227,251]" italics="true" pageId="5" pageNumber="208">P. ziegleri maroccanica</emphasis>
|
||
</taxonomicName>
|
||
: it is semicircular with a slight incurvature of the antero−lateral dorsal furrows. By contrast, in the nominate subspecies, the front of the glabella is more conical with straight diverging dorsal furrows. In
|
||
<taxonomicName id="4C714D5CFF8EFF85FEF6FE818F6AFEE3" authorityName="Feist" authorityYear="2002" box="[262,520,355,379]" class="Trilobita" family="Tropidocoryphidae" genus="Pteroparia" kingdom="Animalia" order="Proetida" pageId="5" pageNumber="208" phylum="Arthropoda" rank="subSpecies" species="ziegleri" subSpecies="maroccanica">
|
||
<emphasis id="B905EACDFF8EFF85FEF6FE818F6AFEE3" box="[262,520,355,379]" italics="true" pageId="5" pageNumber="208">P. ziegleri maroccanica</emphasis>
|
||
</taxonomicName>
|
||
the pygidial axis is narrower; however, the dimensional relations between axis and pleural field varies with ontogenetic growth and additional material of various growth stages must be recovered in order to confirm whether the two subspecies differ in this feature.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |