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<document id="2E6F252753947DDF636F271B663A04F3" ID-DOI="10.5281/zenodo.193489" ID-GBIF-Dataset="7975c079-5a3e-47f5-83b4-b47524d1298c" ID-ISSN="1175-5326" ID-Zenodo-Dep="193489" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460146994407" checkinUser="plazi" docAuthor="Lin, Tomoyuki Komai Feng-Jiau &amp; Chan, Tin-Yam" docDate="2010" docId="038C87AC5E44FFB943E0CB6654EACCFA" docLanguage="en" docName="zt02352p028.pdf" docOrigin="Zootaxa 2352" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Calastacus formosus Lin &amp; Chan, 2010, n. sp." docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FFB5FFD45E48FFA84377CA2B5178CE6A" masterDocTitle="Five new species of Axiidae (Crustacea: Decapoda: Axiidea) from deep-water off Taiwan, with description of a new genus" masterLastPageNumber="28" masterPageNumber="1" pageNumber="13" updateTime="1698235478488" updateUser="plazi">
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<mods:title id="3740C222C07EDB1AF03E4F13A5F9F830">Five new species of Axiidae (Crustacea: Decapoda: Axiidea) from deep-water off Taiwan, with description of a new genus</mods:title>
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<mods:namePart id="79ACE64F7567F7BD61CDC53647DDF827">Lin, Tomoyuki Komai Feng-Jiau</mods:namePart>
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<mods:namePart id="66A8AD26F176D216611E510DC60A7974">Chan, Tin-Yam</mods:namePart>
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<mods:date id="5C2C62E8FAA57BC700C3BAC4B53B4FFA">2010</mods:date>
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<treatment id="038C87AC5E44FFB943E0CB6654EACCFA" ID-DOI="http://doi.org/10.5281/zenodo.6211755" ID-GBIF-Taxon="124955950" ID-Zenodo-Dep="6211755" LSID="urn:lsid:plazi:treatment:038C87AC5E44FFB943E0CB6654EACCFA" httpUri="http://treatment.plazi.org/id/038C87AC5E44FFB943E0CB6654EACCFA" lastPageId="17" lastPageNumber="18" pageId="12" pageNumber="13">
<subSubSection id="C33F65315E44FFA443E0CB665021CFE0" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B9A36BA5E44FFA443E0CB6650A4CF0D" blockId="12.[151,527,298,394]" box="[151,476,333,359]" pageId="12" pageNumber="13">
<heading id="D0D281D65E44FFA443E0CB6650A4CF0D" bold="true" box="[151,476,333,359]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B951EAA85E44FFA443E0CB6650A4CF0D" bold="true" box="[151,476,333,359]" pageId="12" pageNumber="13">
<taxonomicName id="4C254D395E44FFA443E0CB6650EACF0D" box="[151,402,333,359]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B951EAA85E44FFA443E0CB6650EACF0D" bold="true" box="[151,402,333,359]" italics="true" pageId="12" pageNumber="13">Calastacus formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257D35E44FFA442EECB6650A4CF0D" box="[409,476,333,359]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A36BA5E44FFA443E0CB5B5021CFE0" blockId="12.[151,527,298,394]" box="[151,345,368,394]" pageId="12" pageNumber="13">
(
<figureCitation id="131E2A3F5E44FFA443E8CB5B5068CFE0" box="[159,272,368,394]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="13.[151,255,1807,1831]" captionStartId-1="14.[151,259,1872,1896]" captionStartId-2="16.[151,255,1859,1883]" captionTargetBox-0="[211,1357,184,1758]" captionTargetBox-1="[225,1359,191,1810]" captionTargetBox-2="[227,1346,201,1819]" captionTargetId-0="figure@13.[200,1386,147,1785]" captionTargetId-1="figure@14.[200,1386,155,1849]" captionTargetId-2="figure@16.[200,1386,170,1838]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="16" captionText-0="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." captionText-1="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." captionText-2="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri-0="https://zenodo.org/record/193494/files/figure.png" httpUri-1="https://zenodo.org/record/193495/files/figure.png" httpUri-2="https://zenodo.org/record/193496/files/figure.png" pageId="12" pageNumber="13">Figs. 68</figureCitation>
,
<figureCitation id="131E2A3F5E44FFA4426BCB5B5040CFE0" box="[284,312,368,394]" captionStart="FIGURE 13. A" captionStartId="25.[151,260,1531,1555]" captionTargetBox="[151,1434,293,1507]" captionTargetId="figure@25.[151,1436,291,1507]" captionTargetPageId="25" captionText="FIGURE 13. A, Formosaxius dorsum n. gen., n. sp., holotype, female (cl about 12.0 mm), NTOU A 01087; B, Formosaxius dorsum n. gen., n. sp., paratype, female (cl 12.6 mm), NTOU A 00023; C, Ambiaxius propinquus n. sp., holotype, hermaphrodite (cl 11.8 mm), NTOU A 00110; D, Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083; E, Eiconaxius rubrirostris n. sp., holotype, male (cl 5.4 mm), NTOU A 00114; F, Eiconaxius kensleyi n. sp., holotype, male (cl 3.1 mm), NTOU A 00111." httpUri="https://zenodo.org/record/193501/files/figure.png" pageId="12" pageNumber="13">13</figureCitation>
D)
</paragraph>
</subSubSection>
<subSubSection id="C33F65315E44FFA443E0CB9C5538CC28" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8B9A36BA5E44FFA443E0CB9C53E7CF9D" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
<emphasis id="B951EAA85E44FFA443E0CB9C503BCFBB" bold="true" box="[151,323,439,465]" pageId="12" pageNumber="13">
<typeStatus id="549E88185E44FFA443E0CB9C51AACFBB" box="[151,210,439,465]" pageId="12" pageNumber="13">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="549E88185E44FFA44224CB9350BDCFB8" box="[339,453,440,466]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
: hermaphrodite (cl
<quantity id="4CDD9B5F5E44FFA441D0CB93527DCFB8" box="[679,773,440,466]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="12" pageNumber="13" unit="mm" value="7.1">7.1 mm</quantity>
),
<collectingCountry id="F332762A5E44FFA4406FCB9352F1CFB8" box="[792,905,440,466]" name="Taiwan" pageId="12" pageNumber="13">TAIWAN</collectingCountry>
2001, stn CP 104,
<geoCoordinate id="EE11507D5E44FFA44710CB93558CCFB8" box="[1127,1268,440,466]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="9" value="24.814333">24°48.86N</geoCoordinate>
,
<geoCoordinate id="EE11507D5E44FFA44677CB9354E0CFB8" box="[1280,1432,440,466]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="9" value="122.0885">122°05.31E</geoCoordinate>
,
<quantity id="4CDD9B5F5E44FFA443E0CBF65067CF9D" box="[151,287,477,503]" metricMagnitude="2" metricUnit="m" metricValue="4.06" metricValueMax="4.47" metricValueMin="3.65" pageId="12" pageNumber="13" unit="m" value="406.0" valueMax="447.0" valueMin="365.0">365447 m</quantity>
,
<date id="FF9B107A5E44FFA4425DCBF650BECF9D" box="[298,454,477,503]" pageId="12" pageNumber="13" value="2001-05-19">19 May 2001</date>
(
<collectionCode id="ED34AE7F5E44FFA442A2CBF65350CF9D" box="[469,552,477,503]" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="12" pageNumber="13">NTOU</collectionCode>
<accessionNumber id="9476AB595E44FFA44147CBF653E8CF9D" box="[560,656,477,503]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A00083" pageId="12" pageNumber="13" type="EnaNcbi">A00083</accessionNumber>
).
</paragraph>
<paragraph id="8B9A36BA5E44FFA443B1C8295538CC28" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
<typeStatus id="549E88185E44FFA443B1C8295045CC76" box="[198,317,514,540]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
: 2 hermaphrodites (cl 6.0,
<quantity id="4CDD9B5F5E44FFA44106C82953B2CC76" box="[625,714,514,540]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="13" unit="mm" value="6.5">6.5 mm</quantity>
), 1 ovigerous hermaphrodite (cl 9.0 mm),
<collectingCountry id="F332762A5E44FFA447CEC8295452CC76" box="[1209,1322,514,540]" name="Taiwan" pageId="12" pageNumber="13">TAIWAN</collectingCountry>
2001, stn CP 102,
<geoCoordinate id="EE11507D5E44FFA4438AC80350F1CC28" box="[253,393,552,578]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="9" value="24.806334">24°48.38N</geoCoordinate>
, 122°07.97E,
<quantity id="4CDD9B5F5E44FFA4414EC80353B9CC28" box="[569,705,552,578]" metricMagnitude="2" metricUnit="m" metricValue="3.285" metricValueMax="3.31" metricValueMin="3.26" pageId="12" pageNumber="13" unit="m" value="328.5" valueMax="331.0" valueMin="326.0">326331 m</quantity>
,
<date id="FF9B107A5E44FFA441BBC8035211CC28" box="[716,873,552,578]" pageId="12" pageNumber="13" value="2001-05-19">19 May 2001</date>
(
<collectionCode id="ED34AE7F5E44FFA44000C80352B2CC28" box="[887,970,552,578]" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="12" pageNumber="13">NTOU</collectionCode>
A00110).
</paragraph>
</subSubSection>
<subSubSection id="C33F65315E44FFA743B1C866525FC8B8" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="8B9A36BA5E44FFA443B1C86650C0CD2D" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
<emphasis id="B951EAA85E44FFA443B1C8665089CC0D" bold="true" box="[198,497,589,615]" pageId="12" pageNumber="13">
Description of
<typeStatus id="549E88185E44FFA442F1C8665089CC0D" box="[390,497,589,615]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
</emphasis>
. Rostrum (
<figureCitation id="131E2A3F5E44FFA4410AC86653BECC0D" box="[637,710,589,615]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
AD) 0.2 times as long as carapace and about 0.4 times of distance between rostral base and cervical groove, spike-like with acute tip, unarmed anterior to supraocular spine, not continuous with lateral gastric carinae on carapace. Carapace (
<figureCitation id="131E2A3F5E44FFA44088C8B3553DCCD8" box="[1023,1093,664,690]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
AD) smooth; gastric region convex, higher than rostral base; supraocular spines prominent; lateral and submedian gastric carinae absent; median gastric carina unarmed, extending over midway between rostral base and cervical groove; cervical groove distinct, extending nearly to pterygostomial part of carapace; no postcervical carina; pterygostomial margin with small spine.
</paragraph>
<paragraph id="8B9A36BA5E44FFA443B1C9795278CDF8" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
Thoracic sternum (
<figureCitation id="131E2A3F5E44FFA442DFC9795096CD06" box="[424,494,850,876]" captionStart="FIGURE 7" captionStartId="14.[151,259,1872,1896]" captionTargetBox="[225,1359,191,1810]" captionTargetId="figure@14.[200,1386,155,1849]" captionTargetPageId="14" captionText="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
A) with poorly delimited shield on seventh somite, medially separated by deep groove; sixth somite with spade-shaped prominence.
</paragraph>
<paragraph id="8B9A36BA5E44FFA443B1C9B65273CA26" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
First abdominal pleuron (
<figureCitation id="131E2A3F5E44FFA4428EC9B65339CDDD" box="[505,577,925,951]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
E) short, produced ventrally as acute projection; second pleuron broad, anteroventrally rounded; third to fifth pleura rounded; sixth pleura also rounded. Telson (
<figureCitation id="131E2A3F5E44FFA447B3C9E95474CDB6" box="[1220,1292,962,988]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
F) 1.6 times longer than wide, widest proximally, then approximately parallel-sided, lateral margin unarmed, posterior margin strongly convex without posteromedian spine, posterolateral region with 1 minute spiniform seta; dorsal face without spines on obsolete oblique ridges.
</paragraph>
<paragraph id="8B9A36BA5E44FFA443B1CE735349CBA8" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
Eyestalks (
<figureCitation id="131E2A3F5E44FFA4423BCE7350EECA18" box="[332,406,1112,1138]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
C, D) subglobose, 0.4 length of rostrum, immovably attached to cephalothorax, not contiguous; cornea unpigmented, unfaceted, division between cornea and eyestalk unclear. Antennular peduncle (
<figureCitation id="131E2A3F5E44FFA44264CE895023CAD6" box="[275,347,1186,1212]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
C) reaching distal margin of fourth segment of antennal peduncle; first segment with small submarginal spine on statocyst lobe dorsodistally; flagella missing. Antennal peduncle (
<figureCitation id="131E2A3F5E44FFA447D6CEE3559FCA88" box="[1185,1255,1224,1250]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
C, D) with first segment bearing 2 spines on ventrodistal margin (lateral spine longer than mesial spine); second segment with dorsolateral distal spine slender, slightly curved inward in dorsal view, slightly directed dorsally in lateral view, reaching to midlength of fourth segment; scaphocerite slender, directed slightly upwards, slightly falling short of distal margin of fourth segment; third segment with sharp spine on distomesial angle; fourth segment slightly longer than second segment (excluding dorsolateral distal spine); fifth segment about half length of fourth segment; flagellum missing.
</paragraph>
<paragraph id="8B9A36BA5E44FFA443B1CFE65564C858" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
Third maxilliped (
<figureCitation id="131E2A3F5E44FFA442D3CFE65093CB8D" box="[420,491,1485,1511]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
A) moderately slender; coxa with 1 small ventromesial spine; basis also with small ventromesial spine; ischium unarmed on ventral margin; crista dentata with about 20 corneous-tipped teeth; merus with 2 long subdistal spines on ventral margin; carpus unarmed.
</paragraph>
<paragraph id="8B9A36BA5E44FFA443B1CC1654E4C998" blockId="12.[151,1437,439,2034]" pageId="12" pageNumber="13">
Chelipeds (
<figureCitation id="131E2A3F5E44FFA44239CC1650ECC83D" box="[334,404,1597,1623]" captionStart="FIGURE 7" captionStartId="14.[151,259,1872,1896]" captionTargetBox="[225,1359,191,1810]" captionTargetId="figure@14.[200,1386,155,1849]" captionTargetPageId="14" captionText="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
BE) subequal, similar, fairly elongate. Right cheliped (
<figureCitation id="131E2A3F5E44FFA44744CC165503C83D" box="[1075,1147,1597,1623]" captionStart="FIGURE 7" captionStartId="14.[151,259,1872,1896]" captionTargetBox="[225,1359,191,1810]" captionTargetId="figure@14.[200,1386,155,1849]" captionTargetPageId="14" captionText="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
D, E) with coxa bearing spine on ventrodistal margin. Basis also with small spine ventrodistally. Ischium with row of 3 slender spines on ventral margin. Merus with 1 subdistal spine (slightly hooked) and row of 5 slender spines increasing in size distally on ventral margin. Carpus unarmed. Chela 0.9 times as long as carapace and 3.6 times longer than high, ventral margin slightly curving. Palm slightly becoming higher distally, 1.8 times longer than high, with 1 subdistal spine on faintly carinate dorsal margin and 1 spine on lateral surface adjacent to base of fixed finger; lateral surface slightly convex, with few tufts of long setae and row of low tubercles bearing tufts of setae adjacent to ventral margin, extending onto fixed finger; mesial face also with spine adjacent to base of fixed finger. Fixed finger with row of small tubercles and tufts of long setae on mesial face along ventral margin, latter extending onto palm; cutting edge with row of rounded teeth. Dactylus with 3 longitudinal rows of long stiff setae on lateral surface; mesial face medially carinate, also with rows of long stiff setae; cutting edge with broad concavity proximally, forming narrow hiatus when closed, tip closing with tip of fixed finger.
</paragraph>
<caption id="DF5A66325E45FFA543E0CD245314C9AC" httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="13" pageNumber="14" targetBox="[211,1357,184,1758]" targetPageId="13">
<paragraph id="8B9A36BA5E45FFA543E0CD245314C9AC" blockId="13.[151,1436,1807,1990]" pageId="13" pageNumber="14">
<emphasis id="B951EAA85E45FFA543E0CD245061C94D" bold="true" box="[151,281,1807,1831]" pageId="13" pageNumber="14">FIGURE 6.</emphasis>
<taxonomicName id="4C254D395E45FFA54268CD245086C94C" box="[287,510,1807,1830]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B951EAA85E45FFA54268CD245086C94C" box="[287,510,1807,1830]" italics="true" pageId="13" pageNumber="14">Calastacus formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257D35E45FFA54173CD245345C94C" box="[516,573,1807,1830]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
, holotype, hermaphrodite (cl 7.6 mm), NTOU
<accessionNumber id="9476AB595E45FFA54758CD2455FDC94C" box="[1071,1157,1807,1830]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A00083" pageId="13" pageNumber="14" type="EnaNcbi">A00083</accessionNumber>
. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm.
</paragraph>
</caption>
<caption id="DF5A66325E46FFA643E0CD7B52CFC99E" httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="14" pageNumber="15" targetBox="[225,1359,191,1810]" targetPageId="14">
<paragraph id="8B9A36BA5E46FFA643E0CD7B52CFC99E" blockId="14.[151,1436,1872,2036]" pageId="14" pageNumber="15">
<emphasis id="B951EAA85E46FFA643E0CD7B5058C902" bold="true" box="[151,288,1872,1896]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
<taxonomicName id="4C254D395E46FFA64250CD7A536BC902" box="[295,531,1873,1896]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B951EAA85E46FFA64250CD7A536BC902" box="[295,531,1873,1896]" italics="true" pageId="14" pageNumber="15">Calastacus formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257D35E46FFA6416CCD7B5320C90D" box="[539,600,1872,1895]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
, holotype, hermaphrodite (cl 7.6 mm), NTOU
<accessionNumber id="9476AB595E46FFA6471ACD7B55BFC90D" box="[1133,1223,1872,1895]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A00083" pageId="14" pageNumber="15" type="EnaNcbi">A00083</accessionNumber>
. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8B9A36BA5E47FFA743B1CAB355C8CE96" blockId="15.[151,1437,152,2007]" pageId="15" pageNumber="16">
Left cheliped (
<figureCitation id="131E2A3F5E47FFA74204CAB350C1CED8" box="[371,441,152,178]" captionStart="FIGURE 7" captionStartId="14.[151,259,1872,1896]" captionTargetBox="[225,1359,191,1810]" captionTargetId="figure@14.[200,1386,155,1849]" captionTargetPageId="14" captionText="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
A, B) only slightly shorter than major cheliped, otherwise generally similar. Ischium with 4 spines on ventral margin. Merus with 1 subdistal spine on dorsal margin and 7 spines on ventral margin. Armament and setation of other segments identical with those on major cheliped.
</paragraph>
<paragraph id="8B9A36BA5E47FFA743B1CB2353DDCC88" blockId="15.[151,1437,152,2007]" pageId="15" pageNumber="16">
Second pereopod (
<figureCitation id="131E2A3F5E47FFA742D7CB23509DCF48" box="[416,485,264,290]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
B) slender, unarmed on ischium to carpus; carpus about 0.6 length of chela; chela with tufts of long setae on margins, fingers slightly shorter than palm, each with row of minute corneous spinules on cutting edge. Ambulatory legs (third to fifth pereopods) somewhat elongate. Third pereopod (
<figureCitation id="131E2A3F5E47FFA7461BCB7951DDCFF8" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
C) unarmed on ischium to carpus; propodus with row of spiniform setae on lateral surface distally (
<figureCitation id="131E2A3F5E47FFA74658CB53540CCFF8" box="[1327,1396,376,402]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
D); dactylus (
<figureCitation id="131E2A3F5E47FFA7427FCBB65035CFDD" box="[264,333,413,439]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
D, E) subconical, slender, about 0.3 times as long as propodus, slightly twisted, setose. Fourth pereopod (
<figureCitation id="131E2A3F5E47FFA74262CBE95022CFB6" box="[277,346,450,476]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
F) unarmed on ischium to carpus; propodus distally with cluster consisting of stiff setae and longer, stout setulose setae, possibly representing grooming apparatus, and with sets of 1 or 2 spiniform setae and longer stiff setae on lateral surface distally (
<figureCitation id="131E2A3F5E47FFA741ABC826525BCC4D" box="[732,803,525,551]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
G); dactylus (
<figureCitation id="131E2A3F5E47FFA740BFC8265576CC4D" box="[968,1038,525,551]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
G, H) subconical, setose, slightly twisted (tip broken, thus proportion against propodus unknown). Fifth pereopod (
<figureCitation id="131E2A3F5E47FFA74709C81955BFCC26" box="[1150,1223,562,588]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">
Fig.
<date id="FF9B107A5E47FFA747CFC81955BFCC26" box="[1208,1223,562,588]" pageId="15" pageNumber="16">8</date>
</figureCitation>
I) not subchelate, unarmed on ischium to carpus; propodus distally with grooming apparatus consisting of cluster of short to long setae, extending onto lateral and mesial faces (
<figureCitation id="131E2A3F5E47FFA74077C856523FCCFD" box="[768,839,637,663]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
J, K); dactylus (
<figureCitation id="131E2A3F5E47FFA74772C8565535CCFD" box="[1029,1101,637,663]" captionStart="FIGURE 8" captionStartId="16.[151,255,1859,1883]" captionTargetBox="[227,1346,201,1819]" captionTargetId="figure@16.[200,1386,170,1838]" captionTargetPageId="16" captionText="FIGURE 8. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
J, K) very slender, elongate, about 0.4 times as long as propodus, flexor surface excavated near base, proximomesial margin expanded, dorsal surface with row of tufts of stiff setae.
</paragraph>
<paragraph id="8B9A36BA5E47FFA743B1C8C652A0CD6D" blockId="15.[151,1437,152,2007]" box="[198,984,749,775]" pageId="15" pageNumber="16">
Gonopores present on coxae of third and fifth pereopods (
<figureCitation id="131E2A3F5E47FFA7401BC8C652CACD6D" box="[876,946,749,775]" captionStart="FIGURE 7" captionStartId="14.[151,259,1872,1896]" captionTargetBox="[225,1359,191,1810]" captionTargetId="figure@14.[200,1386,155,1849]" captionTargetPageId="14" captionText="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
A).
</paragraph>
<paragraph id="8B9A36BA5E47FFA743B1C939533ACD1D" blockId="15.[151,1437,152,2007]" pageId="15" pageNumber="16">Pleurobranchs absent; podobranchs and arthrobranchs present on second maxilliped through fourth pereopods, podobranchs bearing papillae representing rudimentary gill filaments; epipods present on first maxilliped through fourth pereopod.</paragraph>
<paragraph id="8B9A36BA5E47FFA743B1C9A95469CAAD" blockId="15.[151,1437,152,2007]" pageId="15" pageNumber="16">
First pleopod (
<figureCitation id="131E2A3F5E47FFA7420CC9A950BDCDF6" box="[379,453,898,924]" captionStart="FIGURE 7" captionStartId="14.[151,259,1872,1896]" captionTargetBox="[225,1359,191,1810]" captionTargetId="figure@14.[200,1386,155,1849]" captionTargetPageId="14" captionText="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
F) with first segment (protopod) strongly flattened; second segment (ramus) 0.7 length of first segment, leaf-like, deeply concave on mesial margin proximally, proximomesial protrusion representing appendix interna conspicuous on broad triangular anterior fold, lateral fold forming definite shoulder on lateral margin. Second pleopod (
<figureCitation id="131E2A3F5E47FFA741D3C9D95390CA66" box="[676,744,1010,1036]" captionStart="FIGURE 7" captionStartId="14.[151,259,1872,1896]" captionTargetBox="[225,1359,191,1810]" captionTargetId="figure@14.[200,1386,155,1849]" captionTargetPageId="14" captionText="FIGURE 7. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, sixth to eighth thoracic sternites and coxae of third to fifth pereopods, ventral view (setae omitted); B, chela and carpus of left cheliped, lateral view; C, merus and ischium of left cheliped, lateral view; D, chela and carpus of right cheliped, lateral view; E, merus and ischium of right cheliped, lateral view; F, left first pleopod, lateral view; G, left second pleopod, lateral view; H, distal segments of right second pleopod, mesial view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/193495/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
G, H) with inner ramus consisting of 2-segmented endopod plus distally articulated appendix masculina, all components more or less in line with peduncle, with thumblike appendix interna at base of second segment of endopod; second segment of endopod with slightly concave dorsal (or anterior) margin bearing numerous stiff, curving setae; appendix masculina tapering distally with row of sparse stiff setae laterally and mesially, and with terminal tuft of stiff setae; exopod very slender, half length of endopod. Third to fifth pleopods very slender, without appendices internae.
</paragraph>
<paragraph id="8B9A36BA5E47FFA743B1CEF955BFCB5D" blockId="15.[151,1437,152,2007]" pageId="15" pageNumber="16">
Uropodal endopod (
<figureCitation id="131E2A3F5E47FFA742C7CEF9508CCA86" box="[432,500,1234,1260]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
G) about 2.0 times as long as wide, without lateral spines, dorsal ridge unarmed, distal margin subtruncate; exopod (
<figureCitation id="131E2A3F5E47FFA74143CED35302CB78" box="[564,634,1272,1298]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
G) without lateral serration, posterolateral angle with 1 spine, but no spiniform setae apparent; transverse suture with about 10 slender spiniform setae (
<figureCitation id="131E2A3F5E47FFA7472CCF3655D9CB5D" box="[1115,1185,1309,1335]" captionStart="FIGURE 6" captionStartId="13.[151,255,1807,1831]" captionTargetBox="[211,1357,184,1758]" captionTargetId="figure@13.[200,1386,147,1785]" captionTargetPageId="13" captionText="FIGURE 6. Calastacus formosus n. sp., holotype, hermaphrodite (cl 7.6 mm), NTOU A 00083. A, carapace and cephalic appendages, dorsal view; B, same, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, same, lateral view; E, posterior part of thorax, abdomen and telson, lateral view; F, telson, dorsal view (setae partially omitted); G, left uropod, dorsal view (perpendicular to horizontal plane of uropod; setae partially omitted; lateral margin of exopod damaged proximally); H. close up of distal part of uropodal exopod (setae partially omitted). Scale bars: A, B, E = 2 mm; C, D, F, G = 1 mm; H = 0.5 mm." httpUri="https://zenodo.org/record/193494/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
H).
</paragraph>
<paragraph id="8B9A36BA5E47FFA743B1CF695462C8ED" blockId="15.[151,1437,152,2007]" pageId="15" pageNumber="16">
<emphasis id="B951EAA85E47FFA743B1CF6950CCCB36" bold="true" box="[198,436,1346,1372]" pageId="15" pageNumber="16">
Note on
<typeStatus id="549E88185E47FFA74246CF6950D7CB36" box="[305,431,1346,1372]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
.
</emphasis>
One
<typeStatus id="549E88185E47FFA74280CF695324CB36" box="[503,604,1346,1372]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
lacks right first and fifth pereopods; other two missing first chelipeds and several pereopods. Antennal peduncle with first segment bearing 2 spines on ventrodistal margin (lateral spine longer than mesial spine, and mesial spine small to distinct). First chela 0.7 times as long as carapace and 3.3 times longer than high, ventral margin slightly curving; palm slightly becoming higher distally, 1.8 times longer than high, with 1 subdistal spine on dorsal margin and 1 spine on lateral surface near base of fixed finger; carpus unarmed; merus with 1 subdistal spine on dorsal margin and 7 spines on ventral margin; ischium with 3 spines on ventral margin. Uropodal endopod about 2.0 times as long as wide, without lateral spines, dorsal ridge unarmed, distal margin subtruncate; exopod without lateral serration, posterolateral angle with 1 spine and 1 small spiniform seta; transverse suture with 10 or more slender spiniform setae.
</paragraph>
<paragraph id="8B9A36BA5E47FFA743B1CCB9525FC8B8" blockId="15.[151,1437,152,2007]" pageId="15" pageNumber="16">
<emphasis id="B951EAA85E47FFA743B1CCB9502DC8C6" bold="true" box="[198,341,1682,1708]" pageId="15" pageNumber="16">Coloration.</emphasis>
Body and appendages entirely pale yellowish to pale yellowish pink; cornea of eye opaque. Inner margins of fingers of second chelae dark purplish.
</paragraph>
</subSubSection>
<subSubSection id="C33F65315E47FFA743B1CCF65519C89D" box="[198,1121,1757,1783]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8B9A36BA5E47FFA743B1CCF65519C89D" blockId="15.[151,1437,152,2007]" box="[198,1121,1757,1783]" pageId="15" pageNumber="16">
<emphasis id="B951EAA85E47FFA743B1CCF65027C89D" bold="true" box="[198,351,1757,1783]" pageId="15" pageNumber="16">Distribution</emphasis>
. Known only from northeastern
<collectingCountry id="F332762A5E47FFA741A8CCF65240C89D" box="[735,824,1757,1783]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
; at depths of
<quantity id="4CDD9B5F5E47FFA740A3CCF65524C89D" box="[980,1116,1757,1783]" metricMagnitude="2" metricUnit="m" metricValue="3.8649999999999998" metricValueMax="4.470000000000001" metricValueMin="3.2600000000000002" pageId="15" pageNumber="16" unit="m" value="386.5" valueMax="447.0" valueMin="326.0">
326
<specimenCount id="9D23FD335E47FFA74778CCF65524C89D" box="[1039,1116,1757,1783]" pageId="15" pageNumber="16" type="generic">447 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F65315E47FFB943B1CD295016CC01" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8B9A36BA5E47FFB943B1CD29526FCFB2" blockId="15.[151,1437,152,2007]" lastBlockId="17.[151,1437,152,656]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<emphasis id="B951EAA85E47FFA743B1CD295040C976" bold="true" box="[198,312,1794,1820]" pageId="15" pageNumber="16">Remarks</emphasis>
. The present new species agrees well with the diagnosis of
<taxonomicName id="4C254D395E47FFA74779CD2855E8C976" box="[1038,1168,1795,1820]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAA85E47FFA74779CD2855E8C976" box="[1038,1168,1795,1820]" italics="true" pageId="15" pageNumber="16">Calastacus</emphasis>
</taxonomicName>
, provided by de
<bibRefCitation id="EFB44B4B5E47FFA74617CD295032C928" pageId="15" pageNumber="27" refString="Saint Laurent, M. de (1972) Un thalassinide nouveau du golfe de Gascogne, Calastacus laevis sp. nov. Remarques sur le genre Calastacus Faxon (Crustacea Decapoda Axiidae). Bulletin du Museum national d'Histoire naturelle, Serie 3, Zoology, 29, 347 - 356." type="journal article">Saint Laurent (1972)</bibRefCitation>
,
<bibRefCitation id="EFB44B4B5E47FFA74222CD035375C928" author="Kensley" box="[341,525,1832,1858]" pageId="15" pageNumber="27" refString="Kensley, B. (1989) New genera in the thalassinidean families Calocarididae and Axiidae (Crustacea: Decapoda). Proceedings of the Biological Society of Washington, 102, 960 - 967." type="journal article" year="1989">Kensley (1989)</bibRefCitation>
,
<bibRefCitation id="EFB44B4B5E47FFA7416FCD0353C9C928" author="Poore" box="[536,689,1832,1858]" pageId="15" pageNumber="27" refString="Poore, G. C. B. (1994) A phylogeny of the families of Thalassinidea (Crustacea: Decapoda) with keys to the families and genera. Memoirs of the Museum of Victoria, 54, 79 - 120." type="journal article" year="1994">Poore (1994)</bibRefCitation>
and
<bibRefCitation id="EFB44B4B5E47FFA7419DCD035578C928" author="Poore" box="[746,1024,1832,1858]" pageId="15" pageNumber="27" refString="Poore, G. C. B. &amp; Collins, D. J. (2009) Australian Axiidae. Memoirs of Museum Victoria, 66, 221 - 287" type="journal article" year="2009">Poore &amp; Collins (2009)</bibRefCitation>
. The genus is currently represented by seven species worldwide,
<taxonomicName id="4C254D395E47FFA74287CD66525EC90D" authority="Faxon, 1893" authorityName="Faxon" authorityYear="1893" box="[496,806,1869,1895]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="stilirostris">
<emphasis id="B951EAA85E47FFA74287CD6653F2C90C" box="[496,650,1869,1894]" italics="true" pageId="15" pageNumber="16">C. stilirostris</emphasis>
Faxon, 1893
</taxonomicName>
(
<typeStatus id="549E88185E47FFA74041CD665212C90D" box="[822,874,1869,1895]" pageId="15" pageNumber="16">type</typeStatus>
species) from the eastern Pacific off
<collectingCountry id="F332762A5E47FFA74653CD665406C90D" box="[1316,1406,1869,1895]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
to
<collectingCountry id="F332762A5E47FFA743E0CD5951A8C9E6" box="[151,208,1906,1932]" name="Peru" pageId="15" pageNumber="16">Peru</collectingCountry>
,
<taxonomicName id="4C254D395E47FFA743ABCD58503BC9E6" box="[220,323,1907,1932]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B951EAA85E47FFA743ABCD58503BC9E6" box="[220,323,1907,1932]" italics="true" pageId="15" pageNumber="16">C. laevis</emphasis>
</taxonomicName>
de
<bibRefCitation id="EFB44B4B5E47FFA74218CD595321C9E6" author="Saint" box="[367,601,1906,1932]" pageId="15" pageNumber="27" refString="Saint Laurent, M. de (1972) Un thalassinide nouveau du golfe de Gascogne, Calastacus laevis sp. nov. Remarques sur le genre Calastacus Faxon (Crustacea Decapoda Axiidae). Bulletin du Museum national d'Histoire naturelle, Serie 3, Zoology, 29, 347 - 356." type="journal article" year="1972">Saint Laurent, 1972</bibRefCitation>
from the eastern Atlantic and Mediterranean,
<taxonomicName id="4C254D395E47FFA7470BCD5854E4C9E6" authority="Kensley, 1996" authorityName="Kensley" authorityYear="1996" box="[1148,1436,1906,1932]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="colpos">
<emphasis id="B951EAA85E47FFA7470BCD585594C9E6" box="[1148,1260,1907,1932]" italics="true" pageId="15" pageNumber="16">C. colpos</emphasis>
Kensley, 1996
</taxonomicName>
from the northwestern Gulf of
<collectingCountry id="F332762A5E47FFA7417CCDB35311C9D8" box="[523,617,1944,1970]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
,
<taxonomicName id="4C254D395E47FFA74102CDB352B3C9D8" authority="Kensley, 1996" authorityName="Kensley" authorityYear="1996" box="[629,971,1944,1970]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B951EAA85E47FFA74102CDB3526FC9DB" box="[629,791,1944,1969]" italics="true" pageId="15" pageNumber="16">C. mexicanus</emphasis>
Kensley, 1996
</taxonomicName>
from the Gulf of
<collectingCountry id="F332762A5E47FFA747D4CDB35479C9D8" box="[1187,1281,1944,1970]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
,
<taxonomicName id="4C254D395E47FFA7467ACDB350DFC9BD" authority="Kensley &amp; Chan, 1998" authorityName="Kensley &amp; Chan" authorityYear="1998" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="crosnieri">
<emphasis id="B951EAA85E47FFA7467ACDB354E3C9DB" box="[1293,1435,1944,1969]" italics="true" pageId="15" pageNumber="16">C. crosnieri</emphasis>
<bibRefCitation id="EFB44B4B5E47FFA743E0CD9650DFC9BD" author="Kensley" box="[151,423,1981,2007]" pageId="15" pageNumber="27" refString="Kensley, B. &amp; Chan, T. Y. (1998) Three new species of thalassinidean shrimps (Crustacea, Axiidae and Calocarididae) from Taiwan. Zoosystema, 20, 255 - 264." type="journal article" year="1998">Kensley &amp; Chan, 1998</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F332762A5E47FFA74299CD965331C9BD" box="[494,585,1981,2007]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
,
<taxonomicName id="4C254D395E47FFA74123CD965576C9BD" authority="Komai, Lin &amp; Chan, 2009" authorityName="Komai, Lin &amp; Chan" authorityYear="2009" box="[596,1038,1981,2007]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="B951EAA85E47FFA74123CD9653A8C9BC" box="[596,720,1981,2006]" italics="true" pageId="15" pageNumber="16">C. inflatus</emphasis>
<bibRefCitation id="EFB44B4B5E47FFA741A0CD965576C9BD" author="Komai" box="[727,1038,1981,2007]" pageId="15" pageNumber="27" refString="Komai, T., Lin, F. J. &amp; Chan, T. Y. (2009) A new mud shrimp species of Calastacus (Crustacea: Decapoda: Thalassinidea) from the South China Sea. Zootaxa, 2088, 24 - 30." type="journal article" year="2009">Komai, Lin &amp; Chan, 2009</bibRefCitation>
</taxonomicName>
from the South
<collectingCountry id="F332762A5E47FFA747B9CD96546DC9BD" box="[1230,1301,1981,2007]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
Sea off the Pratas Islands, and
<taxonomicName id="4C254D395E59FFB9420CCAB3527FCED8" authority="Poore &amp; Collins, 2009" authorityName="Poore &amp; Collins" authorityYear="2009" box="[379,775,152,178]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="myalup">
<emphasis id="B951EAA85E59FFB9420CCAB3508CCEDB" box="[379,500,152,177]" italics="true" pageId="17" pageNumber="18">C. myalup</emphasis>
<bibRefCitation id="EFB44B4B5E59FFB9428BCAB3527FCED8" author="Poore" box="[508,775,152,178]" pageId="17" pageNumber="27" refString="Poore, G. C. B. &amp; Collins, D. J. (2009) Australian Axiidae. Memoirs of Museum Victoria, 66, 221 - 287" type="journal article" year="2009">Poore &amp; Collins, 2009</bibRefCitation>
</taxonomicName>
from south Western
<collectingCountry id="F332762A5E59FFB94088CAB35512CED8" box="[1023,1130,152,178]" name="Australia" pageId="17" pageNumber="18">Australia</collectingCountry>
(
<bibRefCitation id="EFB44B4B5E59FFB9470DCAB35457CED8" author="Kensley" box="[1146,1327,152,178]" pageId="17" pageNumber="27" refString="Kensley, B. (1996 a) New species of Calocarididae from the Caribbean Sea and Gulf of Mexico (Crustacea: Decapoda: Thalassinidea). Bulletin of Marine Science, 59, 158 - 168." type="journal article" year="1996" yearSuffix="a">Kensley 1996a</bibRefCitation>
;
<bibRefCitation id="EFB44B4B5E59FFB9464BCAB35041CEBC" author="Kensley" pageId="17" pageNumber="27" refString="Kensley, B. &amp; Chan, T. Y. (1998) Three new species of thalassinidean shrimps (Crustacea, Axiidae and Calocarididae) from Taiwan. Zoosystema, 20, 255 - 264." type="journal article" year="1998">Kensley &amp; Chan 1998</bibRefCitation>
;
<bibRefCitation id="EFB44B4B5E59FFB94232CA975080CEBC" author="Ngoc-Ho" box="[325,504,188,214]" pageId="17" pageNumber="27" refString="Ngoc-Ho, N. (2003) European and Mediterranean Thalassinidea (Crustacea, Decapoda). Zoosystema, 25, 439 - 555." type="journal article" year="2003">Ngoc-Ho 2003</bibRefCitation>
;
<bibRefCitation id="EFB44B4B5E59FFB94173CA9753A4CEBC" author="Komai" box="[516,732,188,214]" pageId="17" pageNumber="27" refString="Komai, T., Lin, F. J. &amp; Chan, T. Y. (2009) A new mud shrimp species of Calastacus (Crustacea: Decapoda: Thalassinidea) from the South China Sea. Zootaxa, 2088, 24 - 30." type="journal article" year="2009">
Komai
<emphasis id="B951EAA85E59FFB9412BCA9653EFCEBC" box="[604,663,189,214]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2009
</bibRefCitation>
;
<bibRefCitation id="EFB44B4B5E59FFB9419ECA975297CEBC" author="Poore" box="[745,1007,188,214]" pageId="17" pageNumber="27" refString="Poore, G. C. B. &amp; Collins, D. J. (2009) Australian Axiidae. Memoirs of Museum Victoria, 66, 221 - 287" type="journal article" year="2009">Poore &amp; Collins 2009</bibRefCitation>
). The new species appears closest to
<taxonomicName id="4C254D395E59FFB943E0CACA5069CE90" box="[151,273,225,250]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="myalup">
<emphasis id="B951EAA85E59FFB943E0CACA5069CE90" box="[151,273,225,250]" italics="true" pageId="17" pageNumber="18">C. myalup</emphasis>
</taxonomicName>
in the presence of a pterygostomial spine on the carapace and the suture of the uropodal exopod, which bears 10 or more slender spiniform setae. Nevertheless,
<taxonomicName id="4C254D395E59FFB94006CB2D5286CF75" box="[881,1022,262,287]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B951EAA85E59FFB94006CB2D5286CF75" box="[881,1022,262,287]" italics="true" pageId="17" pageNumber="18">C. formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257D35E59FFB94772CB2D553BCF4A" box="[1029,1091,262,288]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C254D395E59FFB947ABCB2D542CCF75" box="[1244,1364,262,287]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="myalup">
<emphasis id="B951EAA85E59FFB947ABCB2D542CCF75" box="[1244,1364,262,287]" italics="true" pageId="17" pageNumber="18">C. myalup</emphasis>
</taxonomicName>
in the shape of the uropodal endopod. In
<taxonomicName id="4C254D395E59FFB94144CB0053B9CF2E" box="[563,705,299,324]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B951EAA85E59FFB94144CB0053B9CF2E" box="[563,705,299,324]" italics="true" pageId="17" pageNumber="18">C. formosus</emphasis>
</taxonomicName>
, the posterior margin of the uropodal endopod is subtruncate, but it is rounded in
<taxonomicName id="4C254D395E59FFB942FDCB7B537FCF03" box="[394,519,336,361]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="myalup">
<emphasis id="B951EAA85E59FFB942FDCB7B537FCF03" box="[394,519,336,361]" italics="true" pageId="17" pageNumber="18">C. myalup</emphasis>
</taxonomicName>
. Furthermore, the supraorbital spine on the carapace only just exceeds the orbital margin of the carapace in
<taxonomicName id="4C254D395E59FFB94147CB5E53BBCFE4" box="[560,707,373,398]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B951EAA85E59FFB94147CB5E53BBCFE4" box="[560,707,373,398]" italics="true" pageId="17" pageNumber="18">C. formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257D35E59FFB941BBCB5F5269CFE4" box="[716,785,372,398]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
, rather than overreaching the middle of the eye in
<taxonomicName id="4C254D395E59FFB946F7CB5E5195CFD8" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="myalup">
<emphasis id="B951EAA85E59FFB946F7CB5E5195CFD8" italics="true" pageId="17" pageNumber="18">C. myalup</emphasis>
</taxonomicName>
. The ornamentation of the appendix masculina of the second pleopod consists only of stiff setae in
<taxonomicName id="4C254D395E59FFB946F6CBB2507BCFBD" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B951EAA85E59FFB946F6CBB2507BCFBD" italics="true" pageId="17" pageNumber="18">C. formosus</emphasis>
</taxonomicName>
, but it includes spiniform setae in
<taxonomicName id="4C254D395E59FFB941E0CB955268CFBD" box="[663,784,446,471]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="myalup">
<emphasis id="B951EAA85E59FFB941E0CB955268CFBD" box="[663,784,446,471]" italics="true" pageId="17" pageNumber="18">C. myalup</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF5A66325E58FFB843E0CD6855D9C98C" httpUri="https://zenodo.org/record/193496/files/figure.png" pageId="16" pageNumber="17" targetBox="[227,1346,201,1819]" targetPageId="16">
<paragraph id="8B9A36BA5E58FFB843E0CD6855D9C98C" blockId="16.[151,1436,1859,2022]" pageId="16" pageNumber="17">
<emphasis id="B951EAA85E58FFB843E0CD685061C931" bold="true" box="[151,281,1859,1883]" pageId="16" pageNumber="17">FIGURE 8.</emphasis>
<taxonomicName id="4C254D395E58FFB84256CD685379C930" box="[289,513,1859,1882]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B951EAA85E58FFB84256CD685379C930" box="[289,513,1859,1882]" italics="true" pageId="16" pageNumber="17">Calastacus formosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257D35E58FFB8417FCD685339C930" box="[520,577,1859,1882]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
, holotype, hermaphrodite (cl 7.6 mm), NTOU
<accessionNumber id="9476AB595E58FFB8474DCD6855E8C930" box="[1082,1168,1859,1882]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A00083" pageId="16" pageNumber="17" type="EnaNcbi">A00083</accessionNumber>
. A, left third maxilliped, lateral view; B, left second pereopod, lateral view; C, right third pereopod, lateral view; D, same, distal part of propodus and dactylus, lateral view; E, same, mesial view; F, left fourth pereopod, lateral view (distal part of dactylus damaged); G, same, distal part of propodus and dactylus, lateral view; H, same, mesial view; I, right fifth pereopod, lateral view; J, same, distal part of propodus and dactylus, lateral view; K, same, mesial view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8B9A36BA5E59FFB943B1CBC85016CC01" blockId="17.[151,1437,152,656]" pageId="17" pageNumber="18">
This is the second species of
<taxonomicName id="4C254D395E59FFB94156CBC853DCCF96" box="[545,676,483,508]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAA85E59FFB94156CBC853DCCF96" box="[545,676,483,508]" italics="true" pageId="17" pageNumber="18">Calastacus</emphasis>
</taxonomicName>
known from
<collectingCountry id="F332762A5E59FFB94032CBC852E7CF97" box="[837,927,483,509]" name="Taiwan" pageId="17" pageNumber="18">Taiwan</collectingCountry>
. The other species is
<taxonomicName id="4C254D395E59FFB947EACBC85451CF96" box="[1181,1321,483,508]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crosnieri">
<emphasis id="B951EAA85E59FFB947EACBC85451CF96" box="[1181,1321,483,508]" italics="true" pageId="17" pageNumber="18">C. crosnieri</emphasis>
</taxonomicName>
, which is golden yellow in color and very different from the pale yellowish body of
<taxonomicName id="4C254D395E59FFB94744C82355BFCC4B" box="[1075,1223,520,545]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B951EAA85E59FFB94744C82355BFCC4B" box="[1075,1223,520,545]" italics="true" pageId="17" pageNumber="18">C. formosus</emphasis>
</taxonomicName>
. Considering the general scarcity of species of the genus around the world, the presence of two species from the same region is rather remarkable.
</paragraph>
</subSubSection>
<subSubSection id="C33F65315E59FFB943B1C85D54EACCFA" box="[198,1426,630,656]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8B9A36BA5E59FFB943B1C85D54EACCFA" blockId="17.[151,1437,152,656]" box="[198,1426,630,656]" pageId="17" pageNumber="18">
<emphasis id="B951EAA85E59FFB943B1C85D5033CCFA" bold="true" box="[198,331,630,656]" pageId="17" pageNumber="18">Etymology</emphasis>
. The species is named after its
<typeStatus id="549E88185E59FFB941CCC85D5396CCFA" box="[699,750,630,656]" pageId="17" pageNumber="18">type</typeStatus>
locality
<collectingCountry id="F332762A5E59FFB94023C85D52D2CCFA" box="[852,938,630,656]" name="Taiwan" pageId="17" pageNumber="18">Taiwan</collectingCountry>
derived from a former name, “
<taxonomicName id="4C254D395E59FFB94663C85D54FCCCFA" box="[1300,1412,630,656]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="1" phylum="Arthropoda" rank="species" species="formosa">
<collectingCountry id="F332762A5E59FFB94663C85D54FCCCFA" box="[1300,1412,630,656]" name="Taiwan" pageId="17" pageNumber="18">Formosa</collectingCountry>
</taxonomicName>
”.
</paragraph>
</subSubSection>
</treatment>
</document>