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<document id="1BBAD4BC3FFA7D2ED55CD3A32DECA62A" ID-CLB-Dataset="26715" ID-DOI="10.11646/zootaxa.4609.2.12" ID-GBIF-Dataset="6ba5edc7-407c-41e6-81ca-bff60e226cb8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3187565" ID-ZooBank="BD1E8997-9DC5-40A9-967C-A63FB8F5F212" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558593538407" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2019" docId="C82E878592221568C4B3FF21FEED9B69" docLanguage="en" docName="zootaxa.4609.2.12.pdf" docOrigin="Zootaxa 4609 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Prosoglypta Shavrin &amp; Smetana 2019, gen. nov." docType="treatment" docVersion="8" lastPageNumber="391" masterDocId="3417FFFD9223156BC424FFC0FFED9E12" masterDocTitle="Prosoglypta, a new remarkable genus from Vietnam (Coleoptera: Staphylinidae: Omaliinae: Omaliini)" masterLastPageNumber="394" masterPageNumber="388" pageNumber="389" updateTime="1698731780336" updateUser="ExternalLinkService">
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<mods:title id="294F9BE1E04F36DCC8E085133AF37028">Prosoglypta, a new remarkable genus from Vietnam (Coleoptera: Staphylinidae: Omaliinae: Omaliini)</mods:title>
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<mods:name id="76A274EC9E5C46BB150C8F0563D7B70E" type="personal">
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<mods:namePart id="E90DACE54AA8AA39F26C891619BB15D6">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="365C6A169D7C5267C1AC91CBB4A0474B">Smetana, Aleš</mods:namePart>
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<mods:date id="17D31D08C0DBD5F27EC76202F868470D">2019</mods:date>
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<mods:number id="98EC5A2A9F84E0F5D5D24325E31BCFD2">2019-05-23</mods:number>
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<mods:number id="71525255D24B154500C57F12B888DF10">4609</mods:number>
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<mods:identifier id="CF9F64DFBE186DA3D36883ACCD99936C" type="DOI">10.11646/zootaxa.4609.2.12</mods:identifier>
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<treatment id="C82E878592221568C4B3FF21FEED9B69" ID-DOI="http://doi.org/10.5281/zenodo.5935373" ID-GBIF-Taxon="157251132" ID-Zenodo-Dep="5935373" LSID="urn:lsid:plazi:treatment:C82E878592221568C4B3FF21FEED9B69" httpUri="http://treatment.plazi.org/id/C82E878592221568C4B3FF21FEED9B69" lastPageId="3" lastPageNumber="391" pageId="1" pageNumber="389">
<subSubSection id="089D65189222156AC4B3FF21FD4F9EEE" box="[151,674,225,252]" pageId="1" pageNumber="389" type="nomenclature">
<paragraph id="403836939222156AC4B3FF21FD4F9EEE" blockId="1.[151,674,225,288]" box="[151,674,225,252]" pageId="1" pageNumber="389">
<heading id="1B7081FF9222156AC4B3FF21FD4F9EEE" bold="true" box="[151,674,225,252]" fontSize="11" level="1" pageId="1" pageNumber="389" reason="1">
<emphasis id="72F3EA819222156AC4B3FF21FD4F9EEE" bold="true" box="[151,674,225,252]" pageId="1" pageNumber="389">
<taxonomicName id="87874D109222156AC4B3FF21FDCA9EE9" ID-CoL="8XP5H" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[151,551,225,252]" class="Insecta" family="Staphylinidae" genus="Prosoglypta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="389" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="72F3EA819222156AC4B3FF21FECB9EEE" bold="true" box="[151,294,225,252]" italics="true" pageId="1" pageNumber="389">Prosoglypta</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="69C057FA9222156AC610FF22FD4F9EEE" box="[564,674,226,252]" pageId="1" pageNumber="389" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="089D65189222156AC4B3FEC5FECF9F32" box="[151,290,261,288]" pageId="1" pageNumber="389" type="description">
<paragraph id="403836939222156AC4B3FEC5FECF9F32" blockId="1.[151,674,225,288]" box="[151,290,261,288]" pageId="1" pageNumber="389">
(
<figureCitation id="D8BC2A169222156AC4BBFEC5FEF49F32" box="[159,281,261,288]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–6" captionStart-2="FIGURES 7–13" captionStart-3="FIGURES 14–16" captionStartId-0="1.[151,264,1426,1451]" captionStartId-1="2.[151,264,1831,1856]" captionStartId-2="4.[151,264,1200,1225]" captionStartId-3="5.[151,264,1708,1733]" captionTargetBox-0="[178,1378,617,1373]" captionTargetBox-1="[180,1422,765,1796]" captionTargetBox-2="[184,1407,187,1151]" captionTargetBox-3="[164,1412,869,1677]" captionTargetId-0="figure@1.[151,1436,588,1414]" captionTargetId-1="figure@2.[151,1436,737,1820]" captionTargetId-2="figure@4.[151,1436,167,1188]" captionTargetId-3="figure@5.[151,1436,845,1697]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 1–2. Habitus of holotype (Fig. 1) and location of the type locality in northern Vietnam (Fig. 2) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov. Scale bar: 1.0 mm." captionText-1="FIGURES 3–6. Mouthparts (dorsal view) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 3–labrum, 4–mandibles (left and right), 5–labium and mentum, 6–maxilla (right). Scale bars: 0.1 mm." captionText-2="FIGURES 7–13. Male hind leg (Fig. 7) and abdominal segments of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov. (Figs. 8–13): 8–male tergite VIII (dorsal view), 9–male sternite VIII (ventral view), 10–male genital segment, 11–female tergite VIII (dorsal view), 12–female sternite VIII (ventral view), 13–female genital segment and accessory sclerite (dorsal view). Scale bars: 0.1 mm." captionText-3="FIGURES 14–16. Aedeagus of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 14–aedeagus, parameral view (outline of internal sac shown as point line), 15–internal sac, parameral view, 16–aedeagus, lateral view. Scale bars: 0.1 mm." httpUri-0="https://zenodo.org/record/3187567/files/figure.png" httpUri-1="https://zenodo.org/record/3187569/files/figure.png" httpUri-2="https://zenodo.org/record/3187571/files/figure.png" httpUri-3="https://zenodo.org/record/3187573/files/figure.png" pageId="1" pageNumber="389">Figs. 1−15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="089D65189222156AC4B3FE8AFCD89F70" box="[151,821,329,354]" pageId="1" pageNumber="389" type="materials_examined">
<paragraph id="403836939222156AC4B3FE8AFCD89F70" blockId="1.[151,821,329,354]" box="[151,821,329,354]" pageId="1" pageNumber="389">
<emphasis id="72F3EA819222156AC4B3FE8AFECB9F70" bold="true" box="[151,294,329,354]" pageId="1" pageNumber="389">
<typeStatus id="9F3C88319222156AC4B3FE8AFF219F70" box="[151,204,330,354]" pageId="1" pageNumber="389">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="87874D109222156AC508FE8AFD0C9F70" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[300,737,329,354]" class="Insecta" family="Staphylinidae" genus="Prosoglypta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="389" phylum="Arthropoda" rank="species" species="alesenkae" status="sp. nov.">
<emphasis id="72F3EA819222156AC508FE8AFDFC9F70" box="[300,529,330,354]" italics="true" pageId="1" pageNumber="389">Prosoglypta alesenkae</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="69C057FA9222156AC6CEFE8AFCD89F70" box="[746,821,330,354]" pageId="1" pageNumber="389" rank="species">
<emphasis id="72F3EA819222156AC6CEFE8AFCD89F70" bold="true" box="[746,821,330,354]" pageId="1" pageNumber="389">sp.nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="089D651892221568C4B3FE4FFE1B9C2D" lastPageId="3" lastPageNumber="391" pageId="1" pageNumber="389" type="description">
<paragraph id="403836939222156AC4B3FE4FFB8F9C28" blockId="1.[151,1437,399,571]" pageId="1" pageNumber="389">
<emphasis id="72F3EA819222156AC4B3FE4FFEC49FB8" bold="true" box="[151,297,399,426]" pageId="1" pageNumber="389">Description.</emphasis>
Body strongly convex, oblong; reddish-brown (
<figureCitation id="D8BC2A169222156AC76CFE50FC669FB8" box="[840,907,400,426]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1426,1451]" captionTargetBox="[178,1378,617,1373]" captionTargetId="figure@1.[151,1436,588,1414]" captionTargetPageId="1" captionText="FIGURES 1–2. Habitus of holotype (Fig. 1) and location of the type locality in northern Vietnam (Fig. 2) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov. Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/3187567/files/figure.png" pageId="1" pageNumber="389">Fig. 1</figureCitation>
). Punctation of forebody moderately large and dense, markedly sparser and smaller on head and deeper on pronotum. Dorsal surface of body very glossy, without microsculpture between punctures, without pubescence except of several long setae on apical part of clypeus and dorsolateral portions of head between eyes and middle elevation, short setae along lateral and middle portions of pronotum and elytra, and very short and sparse recumbent setae on abdominal tergites.
</paragraph>
<caption id="14F8661B9222156AC4B3FA52FCD59BD8" httpUri="https://zenodo.org/record/3187567/files/figure.png" pageId="1" pageNumber="389" startId="1.[151,264,1426,1451]" targetBox="[178,1378,617,1373]" targetPageId="1">
<paragraph id="403836939222156AC4B3FA52FCD59BD8" blockId="1.[151,1436,1426,1483]" pageId="1" pageNumber="389">
<emphasis id="72F3EA819222156AC4B3FA52FED09BB9" bold="true" box="[151,317,1426,1451]" pageId="1" pageNumber="389">FIGURES 1–2.</emphasis>
Habitus of holotype (Fig. 1) and location of the type locality in northern Vietnam (Fig. 2) of
<taxonomicName id="87874D109222156AC105FA52FE269BD8" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Prosoglypta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="389" phylum="Arthropoda" rank="species" species="alesenkae" status="gen. et sp. nov.">
<emphasis id="72F3EA819222156AC105FA52FF169BD8" italics="true" pageId="1" pageNumber="389">Prosoglypta alesenkae</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="69C057FA9222156AC5F0FA72FD9F9BD8" box="[468,626,1458,1482]" pageId="1" pageNumber="389" rank="species">
<emphasis id="72F3EA819222156AC5F0FA72FD9F9BD8" bold="true" box="[468,626,1458,1482]" pageId="1" pageNumber="389">gen. et sp. nov.</emphasis>
</taxonomicNameLabel>
Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="4038369392221569C4E3F9CBFD309FBD" blockId="1.[151,1437,1547,2042]" lastBlockId="2.[151,1437,153,719]" lastPageId="2" lastPageNumber="390" pageId="1" pageNumber="389">
Head markedly narrower than pronotum, transverse, with sharp nuchal constriction and obtuse postocular ridge, with a pair of ocelli at level of posterior margins of eyes, with very narrow impressions near anterior margins of each ocellus and long furrows beginning from inner lateral margin of ocelli between relatively convex shield-like elevated middle portion, gradually widened apicad. Lateroapical portions of middle elevation with slight and relatively wide impressions at level of antennal insertions. Eyes large and strongly protruded, four to five times as long as surface between basal margin of eye and postocular ridge (if seen laterally); dorsal margins of eyes with thin slightly elevated infraorbital ridge; dorsal surface slightly convex between infraorbital ridge and lateral margin of middle elevation. Antennal insertion concealed under shield-like elevated middle portion of head; antenna slightly shorter than head and pronotum combined, with last five antennomeres forming a loose club, first six antennomeres with scattered long setation, remaining antennomeres with additional dense short setae. Labrum (
<figureCitation id="D8BC2A169222156AC0ECF88FFAE49978" box="[1224,1289,1871,1898]" captionStart="FIGURES 3–6" captionStartId="2.[151,264,1831,1856]" captionTargetBox="[180,1422,765,1796]" captionTargetId="figure@2.[151,1436,737,1820]" captionTargetPageId="2" captionText="FIGURES 3–6. Mouthparts (dorsal view) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 3–labrum, 4–mandibles (left and right), 5–labium and mentum, 6–maxilla (right). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187569/files/figure.png" pageId="1" pageNumber="389">Fig. 3</figureCitation>
) short, transverse, slightly concaved anteriorly, with four pairs of long setae on anterolateral portions and two rows of short setae in middle of apical portion. Mandibles (
<figureCitation id="D8BC2A169222156AC671F857FD4999A0" box="[597,676,1943,1970]" captionStart="FIGURES 3–6" captionStartId="2.[151,264,1831,1856]" captionTargetBox="[180,1422,765,1796]" captionTargetId="figure@2.[151,1436,737,1820]" captionTargetPageId="2" captionText="FIGURES 3–6. Mouthparts (dorsal view) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 3–labrum, 4–mandibles (left and right), 5–labium and mentum, 6–maxilla (right). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187569/files/figure.png" pageId="1" pageNumber="389">Figs. 4</figureCitation>
) moderately broad, each with acute apex; right mandible with more curved apex and small obtuse tooth at distal third of cutting edge; articulating molar lobes with moderately strong acute dents almost along entire lateral margin. Maxilla (
<figureCitation id="D8BC2A169222156AC72EF81FFCA399E8" box="[778,846,2015,2042]" captionStart="FIGURES 3–6" captionStartId="2.[151,264,1831,1856]" captionTargetBox="[180,1422,765,1796]" captionTargetId="figure@2.[151,1436,737,1820]" captionTargetPageId="2" captionText="FIGURES 3–6. Mouthparts (dorsal view) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 3–labrum, 4–mandibles (left and right), 5–labium and mentum, 6–maxilla (right). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187569/files/figure.png" pageId="1" pageNumber="389">Fig. 6</figureCitation>
) narrow; galea oblong, with dense long apical setae; lacinia narrow, straight on outer margin, with row of small thorns on apical third of inner margin, apex with feebly curved tooth; maxillary palpus moderately long, second palpomere about as long as and slightly wider than third palpomere, apical palpomere slightly narrower and longer than penultimate palpomere, from apical third tapering toward apex. Labium (
<figureCitation id="D8BC2A1692211569C5BFFEC5FE0D9F0D" box="[411,480,260,287]" captionStart="FIGURES 3–6" captionStartId="2.[151,264,1831,1856]" captionTargetBox="[180,1422,765,1796]" captionTargetId="figure@2.[151,1436,737,1820]" captionTargetPageId="2" captionText="FIGURES 3–6. Mouthparts (dorsal view) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 3–labrum, 4–mandibles (left and right), 5–labium and mentum, 6–maxilla (right). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187569/files/figure.png" pageId="2" pageNumber="390">Fig. 5</figureCitation>
) distinctly bilobed, palpus three-segmented, narrow, each segment distinctly longer than wide, apical segment narrower than penultimate segment. Mentum (
<figureCitation id="D8BC2A1692211569C7E1FEE9FBEA9F51" box="[965,1031,296,323]" captionStart="FIGURES 3–6" captionStartId="2.[151,264,1831,1856]" captionTargetBox="[180,1422,765,1796]" captionTargetId="figure@2.[151,1436,737,1820]" captionTargetPageId="2" captionText="FIGURES 3–6. Mouthparts (dorsal view) of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 3–labrum, 4–mandibles (left and right), 5–labium and mentum, 6–maxilla (right). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187569/files/figure.png" pageId="2" pageNumber="390">Fig. 5</figureCitation>
) moderately large, trapezoidal, with four long setae on anterior third. Gular sutures widely separated from each other at level of posterior margins of eyes, strongly divergent posteriad. Each gena with distinct, moderately protruded anterolateral extensions, visible in front of anterior margins of eyes, if seen dorsally.
</paragraph>
<paragraph id="4038369392211569C4E3FE79FC2F9CDD" blockId="2.[151,1437,153,719]" pageId="2" pageNumber="390">Pronotum transverse, markedly convex, without longitudinal impressions, lateral margins narrowly and slightly explanate, indistinctly serrate on anterior half; posterolateral portions bearing indistinct, wide lateral pits. Prothorax short, strongly transverse, with moderately short acute intercoxal process reaching about middle of front coxae; surface with sparse and large deep punctures. Mesothorax short, with distinct, long median carina, reaching apex of very long and narrow intercoxal process extending to posterior third of mesocoxae that are narrowly separated from each other. Scutellum wide, triangular. Metathorax markedly transverse, convex, with deep mesocoxal cavities and moderately wide and short intercoxal process, obtuse apically and extending to mesosternal process; surface with sparse, moderately large and deep punctation, denser on lateral portions.</paragraph>
<caption id="14F8661B92211569C4B3F8E7FC46994D" httpUri="https://zenodo.org/record/3187569/files/figure.png" pageId="2" pageNumber="390" startId="2.[151,264,1831,1856]" targetBox="[180,1422,765,1796]" targetPageId="2">
<paragraph id="4038369392211569C4B3F8E7FC46994D" blockId="2.[151,1436,1831,1888]" pageId="2" pageNumber="390">
<emphasis id="72F3EA8192211569C4B3F8E7FED49952" bold="true" box="[151,313,1831,1856]" pageId="2" pageNumber="390">FIGURES 3–6.</emphasis>
Mouthparts (dorsal view) of
<taxonomicName id="87874D1092211569C67CF8E8FBE99952" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[600,1028,1831,1856]" class="Insecta" family="Staphylinidae" genus="Prosoglypta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="390" phylum="Arthropoda" rank="species" species="alesenkae" status="gen. et sp. nov.">
<emphasis id="72F3EA8192211569C67CF8E8FCD69952" box="[600,827,1832,1856]" italics="true" pageId="2" pageNumber="390">Prosoglypta alesenkae</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="69C057FA92211569C028F8E8FB4E9952" box="[1036,1187,1832,1856]" pageId="2" pageNumber="390" rank="species">
<emphasis id="72F3EA8192211569C028F8E8FB4E9952" bold="true" box="[1036,1187,1832,1856]" pageId="2" pageNumber="390">gen. et sp. nov.</emphasis>
</taxonomicNameLabel>
: 3–labrum, 4–mandibles (left and right), 5–labium and mentum, 6–maxilla (right). Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="4038369392211568C4E3F862FCC79EC5" blockId="2.[151,1436,1953,2016]" lastBlockId="3.[151,1437,153,1403]" lastPageId="3" lastPageNumber="391" pageId="2" pageNumber="390">Legs short; apical half of pro- and mesotibiae with spines on inner and outer margins, markedly denser on protibiae; metatibiae without spines, covered with long setae; shape of hind trochanter, and shape and setation of protarsomeres 1−4 sexually dimorphic (for details see the species description); tarsi short, apical tarsomere equal to or slightly longer than previous four tarsomeres combined.</paragraph>
<paragraph id="4038369392201568C4E3FF21FC1B9F51" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="391">Elytra markedly convex, moderately short and wide, extending to apical margin of abdominal tergite IV or basal margin of abdominal tergite V, weakly widened posteriad, with moderately broad epipleura and well-developed rounded humeri, with slightly serrate lateral margins. Wings fully developed.</paragraph>
<paragraph id="4038369392201568C4E3FE8DFC9D9FBD" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="391">Abdomen convex, distinctly narrower than elytra, with a pair of moderately wide tomentose spots (wing-folding patches) on middle of abdominal tergite IV; apical margins of tergites IV and V with brickwall sculpture, apical margin of tergite VII with very narrow, indistinct palisade fringe.</paragraph>
<paragraph id="4038369392201568C4E3FE79FAFF9FC1" blockId="3.[151,1437,153,1403]" box="[199,1298,441,468]" pageId="3" pageNumber="391">
Male. Apical abdominal (genital) segment as in
<figureCitation id="D8BC2A1692201568C6C5FE79FCD99FC1" box="[737,820,441,467]" captionStart="FIGURES 7–13" captionStartId="4.[151,264,1200,1225]" captionTargetBox="[184,1407,187,1151]" captionTargetId="figure@4.[151,1436,167,1188]" captionTargetPageId="4" captionText="FIGURES 7–13. Male hind leg (Fig. 7) and abdominal segments of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov. (Figs. 8–13): 8–male tergite VIII (dorsal view), 9–male sternite VIII (ventral view), 10–male genital segment, 11–female tergite VIII (dorsal view), 12–female sternite VIII (ventral view), 13–female genital segment and accessory sclerite (dorsal view). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187571/files/figure.png" pageId="3" pageNumber="391">Fig. 10</figureCitation>
. Aedeagus with fused parameres (
<figureCitation id="D8BC2A1692201568C095FE79FAE89FC1" box="[1201,1285,441,467]" captionStart="FIGURES 14–16" captionStartId="5.[151,264,1708,1733]" captionTargetBox="[164,1412,869,1677]" captionTargetId="figure@5.[151,1436,845,1697]" captionTargetPageId="5" captionText="FIGURES 14–16. Aedeagus of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov.: 14–aedeagus, parameral view (outline of internal sac shown as point line), 15–internal sac, parameral view, 16–aedeagus, lateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187573/files/figure.png" pageId="3" pageNumber="391">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="4038369392201568C4E3FE1DFCFE9C09" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="391">
Female. Accessory abdominal sternite (
<figureCitation id="D8BC2A1692201568C6A6FE1DFD3E9FE5" box="[642,723,477,503]" captionStart="FIGURES 7–13" captionStartId="4.[151,264,1200,1225]" captionTargetBox="[184,1407,187,1151]" captionTargetId="figure@4.[151,1436,167,1188]" captionTargetPageId="4" captionText="FIGURES 7–13. Male hind leg (Fig. 7) and abdominal segments of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov. (Figs. 8–13): 8–male tergite VIII (dorsal view), 9–male sternite VIII (ventral view), 10–male genital segment, 11–female tergite VIII (dorsal view), 12–female sternite VIII (ventral view), 13–female genital segment and accessory sclerite (dorsal view). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187571/files/figure.png" pageId="3" pageNumber="391">Fig. 13</figureCitation>
) wide and oval. Sclerotized spermatheca missing. Gonocoxites moderately wide and long, with thin short styli (
<figureCitation id="D8BC2A1692201568C697FDC1FCEB9C09" box="[691,774,513,539]" captionStart="FIGURES 7–13" captionStartId="4.[151,264,1200,1225]" captionTargetBox="[184,1407,187,1151]" captionTargetId="figure@4.[151,1436,167,1188]" captionTargetPageId="4" captionText="FIGURES 7–13. Male hind leg (Fig. 7) and abdominal segments of Prosoglypta alesenkae Shavrin &amp; Smetana, gen. et sp. nov. (Figs. 8–13): 8–male tergite VIII (dorsal view), 9–male sternite VIII (ventral view), 10–male genital segment, 11–female tergite VIII (dorsal view), 12–female sternite VIII (ventral view), 13–female genital segment and accessory sclerite (dorsal view). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3187571/files/figure.png" pageId="3" pageNumber="391">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="4038369392201568C4E3FDE5FE1B9C2D" blockId="3.[151,1437,153,1403]" box="[199,502,549,575]" pageId="3" pageNumber="391">Immature stages unknown.</paragraph>
</subSubSection>
<subSubSection id="089D651892201568C4E3FD88FBF29B21" pageId="3" pageNumber="391" type="discussion">
<paragraph id="4038369392201568C4E3FD88FC839A6D" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="391">
<emphasis id="72F3EA8192201568C4E3FD88FEA19C71" bold="true" box="[199,332,584,611]" pageId="3" pageNumber="391">Discussion.</emphasis>
Based on the shape of the strongly convex and short body, with elevated middle portion of head separated from each lateral side by a distinct, long furrow, as well as on the presence of conspicuous process of anterolateral portion of each gena, visible dorsally, the new genus is similar to two genera of
<taxonomicName id="87874D1092201568C0BBFD51FAEA9CB9" authorityName="MacLeay" authorityYear="1825" box="[1183,1287,657,683]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="tribe" tribe="Omaliini">Omaliini</taxonomicName>
:
<taxonomicName id="87874D1092201568C134FD51FA719CB9" authorityName="Thomson" authorityYear="1858" box="[1296,1436,657,683]" class="Insecta" family="Staphylinidae" genus="Pycnoglypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C134FD51FA719CB9" box="[1296,1436,657,683]" italics="true" pageId="3" pageNumber="391">Pycnoglypta</emphasis>
</taxonomicName>
and
<taxonomicName id="87874D1092201568C4ECFD76FEBC9CDD" authorityName="Smetana" authorityYear="1987" box="[200,337,694,719]" class="Insecta" family="Staphylinidae" genus="Prosopaspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C4ECFD76FEBC9CDD" box="[200,337,694,719]" italics="true" pageId="3" pageNumber="391">Prosopaspis</emphasis>
</taxonomicName>
. Despite the similar habitus with
<taxonomicName id="87874D1092201568C6F4FD75FCB19CDD" authorityName="Thomson" authorityYear="1858" box="[720,860,693,719]" class="Insecta" family="Staphylinidae" genus="Pycnoglypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C6F4FD75FCB19CDD" box="[720,860,693,719]" italics="true" pageId="3" pageNumber="391">Pycnoglypta</emphasis>
</taxonomicName>
, as well as in the presence of well-developed neck constriction, shapes of apical portions of mandibles and transverse metathorax, absence of median carina on the prothorax, contiguous position of the mesocoxae, general shape of the aedeagus with the presence of the median lobe, and well defined female accessory sclerite (
<bibRefCitation id="24164B6292201568C65BFCE1FCF69D2E" author="Shavrin, A. V." box="[639,795,801,828]" pageId="3" pageNumber="391" pagination="1 - 94" refId="ref2991" refString="Shavrin, A. V. (2016) A review of the genus Pycnoglypta Thomson, 1858 (Staphylinidae, Omaliinae, Omaliini) with notes on related taxa. Zootaxa, 4077 (1), 1 - 94. https: // doi. org / 10.11646 / zootaxa. 4077.1.1" type="journal article" year="2016">Shavrin 2016</bibRefCitation>
),
<taxonomicName id="87874D1092201568C70AFCE1FC5B9D29" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[814,950,801,827]" class="Insecta" family="Staphylinidae" genus="Prosoglypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="72F3EA8192201568C70AFCE1FC5B9D29" box="[814,950,801,827]" italics="true" pageId="3" pageNumber="391">Prosoglypta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69C057FA92201568C799FCE1FBCB9D29" box="[957,1062,801,827]" pageId="3" pageNumber="391" rank="genus">
<emphasis id="72F3EA8192201568C799FCE1FBCB9D29" bold="true" box="[957,1062,801,827]" pageId="3" pageNumber="391">gen. nov.</emphasis>
</taxonomicNameLabel>
differs from it by the presence of ocelli, significantly protruded eyes, and by the absence of sclerotized spermatheca. Based on the shapes of eyes and gonocoxites, presence of the ocelli, modified male adhesive setae on protarsomeres 1−4, a pair of very long spines in basal portion of the internal sac, and the absence of the sclerotized spermatheca (
<bibRefCitation id="24164B6292201568C07CFC4DFB129DB5" author="Smetana, A." box="[1112,1279,909,935]" pageId="3" pageNumber="391" pagination="367 - 380" refId="ref3045" refString="Smetana, A. (1987) Prosopaspis, a new omaliine genus with two new species from the Himalayas (Coleoptera: Staphylinidae). (132 nd contribution to the knowledge of Staphylinidae). The Coleopterists Bulletin, 40, 367 - 380. [1986]" type="journal article" year="1987">Smetana 1987</bibRefCitation>
),
<taxonomicName id="87874D1092201568C130FC4DFA719DB5" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[1300,1436,909,935]" class="Insecta" family="Staphylinidae" genus="Prosoglypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="72F3EA8192201568C130FC4DFA719DB5" box="[1300,1436,909,935]" italics="true" pageId="3" pageNumber="391">Prosoglypta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69C057FA92201568C4B3FC71FEEC9DD9" box="[151,257,945,971]" pageId="3" pageNumber="391" rank="genus">
<emphasis id="72F3EA8192201568C4B3FC71FEEC9DD9" bold="true" box="[151,257,945,971]" pageId="3" pageNumber="391">gen. nov.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="87874D1092201568C5B2FC72FDF29DD9" authorityName="Smetana" authorityYear="1987" box="[406,543,946,971]" class="Insecta" family="Staphylinidae" genus="Prosopaspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C5B2FC72FDF29DD9" box="[406,543,946,971]" italics="true" pageId="3" pageNumber="391">Prosopaspis</emphasis>
</taxonomicName>
, from which it differs by the narrower elevation of the middle portion of head, shorter conspicuous process of anteriolateral portions of each gena, shapes and structure of gular sutures (more modified and coriaceous in
<taxonomicName id="87874D1092201568C5F5FC3AFDB79A01" authorityName="Smetana" authorityYear="1987" box="[465,602,1018,1043]" class="Insecta" family="Staphylinidae" genus="Prosopaspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C5F5FC3AFDB79A01" box="[465,602,1018,1043]" italics="true" pageId="3" pageNumber="391">Prosopaspis</emphasis>
</taxonomicName>
), presence of the preapical tooth in left mandible and nuchal constriction, significantly shorter elytra, long median carina on mesothorax (
<taxonomicName id="87874D1092201568C75EFBDEFBEE9A25" authorityName="Smetana" authorityYear="1987" box="[890,1027,1054,1079]" class="Insecta" family="Staphylinidae" genus="Prosopaspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C75EFBDEFBEE9A25" box="[890,1027,1054,1079]" italics="true" pageId="3" pageNumber="391">Prosopaspis</emphasis>
</taxonomicName>
has two moderately strong median ridges on it), significantly narrowly separated mesocoxae, absence of the median carina on the prothorax, as well as by presence of the median lobe and basal pieces of the aedeagus.
</paragraph>
<paragraph id="4038369392201568C4E3FB49FBF29B21" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="391">
From both these genera,
<taxonomicName id="87874D1092201568C5C4FB49FD859AB1" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[480,616,1161,1187]" class="Insecta" family="Staphylinidae" genus="Prosoglypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="72F3EA8192201568C5C4FB49FD859AB1" box="[480,616,1161,1187]" italics="true" pageId="3" pageNumber="391">Prosoglypta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69C057FA92201568C654FB49FD379AB1" box="[624,730,1161,1187]" pageId="3" pageNumber="391" rank="genus">
<emphasis id="72F3EA8192201568C654FB49FD379AB1" bold="true" box="[624,730,1161,1187]" pageId="3" pageNumber="391">gen. nov.</emphasis>
</taxonomicNameLabel>
differs by the following characters: 1) dorsal and ventral margins of eyes with narrowly elevated infraorbital ridge; 2) antenomeres 7−11 forming a loose club; 3) lateroapical margins of the pronotum indistinctly serrate; 4) middle portion of mesothorax with distinct long carina; 5) hind trochanter of male with very long projection, as that in some western Palaearctic species of
<taxonomicName id="87874D1092201568C0B9FB35FE979B21" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" class="Insecta" family="Staphylinidae" genus="Hapalaraea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C0B9FB35FAC99B1D" box="[1181,1316,1269,1295]" italics="true" pageId="3" pageNumber="391">Hapalaraea</emphasis>
Thomson, 1858 (
<bibRefCitation id="24164B6292201568C4FAFAD9FE9F9B21" author="Zanetti, A." box="[222,370,1305,1331]" pageId="3" pageNumber="391" pagination="461 - 467" refId="ref3131" refString="Zanetti, A. (1983) Descrizione di Hapalaraea settei n. sp. di Corsica, con considerazioni sui generi Hapalaraea Thoms. e Phyllodrepa Thoms. (Coleoptera, Staphylinidae, Omaliinae). Bollettino del Museo Civico di Storia Naturale di Verona, 9, 461 - 467. [1982]" type="journal article" year="1983">Zanetti 1983</bibRefCitation>
)
</taxonomicName>
; 6) unusual structure of the aedeagus, with fused parameres.
</paragraph>
</subSubSection>
<subSubSection id="089D651892201568C4E3FAFDFEED9B69" pageId="3" pageNumber="391" type="etymology">
<paragraph id="4038369392201568C4E3FAFDFEED9B69" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="391">
<emphasis id="72F3EA8192201568C4E3FAFDFEA39B45" bold="true" box="[199,334,1341,1367]" pageId="3" pageNumber="391">Etymology.</emphasis>
The feminine generic name is a combination of parts of the existing names
<taxonomicName id="87874D1092201568C08CFAFEFADC9B45" authorityName="Smetana" authorityYear="1987" box="[1192,1329,1342,1367]" class="Insecta" family="Staphylinidae" genus="Prosopaspis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C08CFAFEFADC9B45" box="[1192,1329,1342,1367]" italics="true" pageId="3" pageNumber="391">Prosopaspis</emphasis>
</taxonomicName>
and
<taxonomicName id="87874D1092201568C14DFAFEFF149B69" authorityName="Thomson" authorityYear="1858" class="Insecta" family="Staphylinidae" genus="Pycnoglypta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="72F3EA8192201568C14DFAFEFF149B69" italics="true" pageId="3" pageNumber="391">Pycnoglypta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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