treatments-xml/data/4D/64/9C/4D649C12E0B357F397E75E598849649E.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1180.109298" ID-Pensoft-Pub="1313-2970-1180-27" ID-Pensoft-UUID="F1B73EB596495DC9AA147719790994F7" ID-ZooBank="6E385B2EBC934E1AB84AB2E5CD9ECBB3" ModsDocID="1313-2970-1180-27" checkinTime="1694791467365" checkinUser="pensoft" docAuthor="El Yaagoubi, Sara, Vuataz, Laurent, El Alami, Majida &amp; Gattolliat, Jean-Luc" docDate="2023" docId="4D649C12E0B357F397E75E598849649E" docLanguage="en" docName="ZooKeys 1180: 27-50" docOrigin="ZooKeys 1180" docPubDate="2023-09-15" docSource="http://dx.doi.org/10.3897/zookeys.1180.109298" docTitle="Baetis rifensis El Yaagoubi, Vuataz &amp; Gattolliat 2023, sp. nov." docType="treatment" docUuid="39BCB7F2-5C63-43AB-8C01-8A111337EE67" docUuidSource="ZooBank" docVersion="1" id="F1B73EB596495DC9AA147719790994F7" lastPageNumber="27" masterDocId="F1B73EB596495DC9AA147719790994F7" masterDocTitle="A new species of the Baetis fuscatus group (Ephemeroptera, Baetidae) from Morocco" masterLastPageNumber="50" masterPageNumber="27" pageNumber="27" updateTime="1694791467365" updateUser="pensoft">
<mods:mods id="EF6602EE9F40C69A2F0089C33697AA80" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="056F9DA9B81AE42ECAB253F5A7FD59F2">A new species of the Baetis fuscatus group (Ephemeroptera, Baetidae) from Morocco</mods:title>
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<mods:namePart id="0B85F6D1FE8037CC9FD82573B137AFBB">El Yaagoubi, Sara</mods:namePart>
<mods:nameIdentifier id="F594BBEEAE897A4A453EB4BB7FD3BC28" type="ORCID">https://orcid.org/0000-0003-1860-6433</mods:nameIdentifier>
<mods:affiliation id="E197737880354CE0917D570BB4CF8E64">Laboratoire Ecologie, Systematique, Conservation de la Biodiversite (LESCB), Unite de Recherche Labellisee CNRST N ° 18, Universite Abdelmalek Essaadi, Faculte des Sciences, Departement de Biologie, B. P. 2121 93002 Tetouan, Morocco</mods:affiliation>
<mods:nameIdentifier id="C251A7B8EF844E49E9E1B57850FF472E" type="email">sara1996.sey@gmail.com</mods:nameIdentifier>
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<mods:namePart id="87B905865EC6DDE176DD4E9984AF09F2">Vuataz, Laurent</mods:namePart>
<mods:nameIdentifier id="0D36C02CD27E2686B8F26F7842DA9272" type="ORCID">https://orcid.org/0000-0001-9193-8683</mods:nameIdentifier>
<mods:affiliation id="31F7E25C43A689896B8FB65764D58D4F">Museum Cantonal des Sciences Naturelles, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:namePart id="4C44AB967251FE51BFE9D3444BC0E863">El Alami, Majida</mods:namePart>
<mods:nameIdentifier id="307523F4B7E33239526B857788A23EB1" type="ORCID">https://orcid.org/0000-0003-2664-646X</mods:nameIdentifier>
<mods:affiliation id="3D1BA2259B19497643A32B46C30A5884">Laboratoire Ecologie, Systematique, Conservation de la Biodiversite (LESCB), Unite de Recherche Labellisee CNRST N ° 18, Universite Abdelmalek Essaadi, Faculte des Sciences, Departement de Biologie, B. P. 2121 93002 Tetouan, Morocco</mods:affiliation>
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<mods:namePart id="24B44B4BFEA25F2C1656588F564F3EA9">Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier id="F72792F19C4787CDAB30991C9B6EBA69" type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation id="3E8001E23F95391645B565C97D648979">Museum Cantonal des Sciences Naturelles, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:date id="AD71C07AC44BA4856E60BE4C77181054">2023</mods:date>
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<treatment id="4D649C12E0B357F397E75E598849649E" LSID="urn:lsid:zoobank.org:act:39BCB7F2-5C63-43AB-8C01-8A111337EE67" httpUri="http://treatment.plazi.org/id/4D649C12E0B357F397E75E598849649E" lastPageNumber="27" pageId="0" pageNumber="27">
<subSubSection id="92E72E8EF0B24E262207CB960E5A9657" pageId="0" pageNumber="27" type="nomenclature">
<paragraph id="EA8DE17FDBA64BE4480885A1BE0E3983" pageId="0" pageNumber="27">
<taxonomicName id="C6BD983C2203AC931303BBCB8D661741" LSID="https://zoobank.org/39BCB7F2-5C63-43AB-8C01-8A111337EE67" authority="El Yaagoubi, Vuataz &amp; Gattolliat" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">Baetis rifensis El Yaagoubi, Vuataz &amp; Gattolliat</taxonomicName>
<taxonomicNameLabel id="FC8B27135FFAA85FEF5C20C588C7CC8C" pageId="0" pageNumber="27">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="1A5210CD2B375D948499C606CA1EC43E" pageId="0" pageNumber="27" type="description">
<paragraph id="E065BE6A603E96D06F70ABD7B4387D59" pageId="0" pageNumber="27">
<figureCitation id="6725E9E4D46C76B22868AAF5B06F41EC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Baetis rifensis sp. nov., larva habitus a female, dorsal view b male, dorsal view c male, lateral view." figureDoi="10.3897/zookeys.1180.109298.figure2" httpUri="https://binary.pensoft.net/fig/905319" pageId="0" pageNumber="27">Figs 2</figureCitation>
<figureCitation id="96F34E6C949B4CD1AB2BF86AA77A2F1C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baetis rifensis sp. nov., larva morphology a labrum dorsal face b labrum ventral face." figureDoi="10.3897/zookeys.1180.109298.figure3" httpUri="https://binary.pensoft.net/fig/905320" pageId="0" pageNumber="27">, 3</figureCitation>
<figureCitation id="7873E277608CFC3DBD44941F0C2E8E10" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Baetis rifensis sp. nov., larva morphology a, b right mandible c, d left mandible." figureDoi="10.3897/zookeys.1180.109298.figure4" httpUri="https://binary.pensoft.net/fig/905321" pageId="0" pageNumber="27">, 4</figureCitation>
<figureCitation id="9C5899F80C52B621E6E5F7048049A371" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Baetis rifensis sp. nov., larva morphology a maxilla b apex of maxilla, c apex of maxillary palp, d hypopharynx." figureDoi="10.3897/zookeys.1180.109298.figure5" httpUri="https://binary.pensoft.net/fig/905322" pageId="0" pageNumber="27">, 5</figureCitation>
<figureCitation id="35BEE27055E0AF5B14EFC35DF7CCABE2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Baetis rifensis sp. nov., larva morphology a labial palp b paraglossae." figureDoi="10.3897/zookeys.1180.109298.figure6" httpUri="https://binary.pensoft.net/fig/905323" pageId="0" pageNumber="27">, 6</figureCitation>
<figureCitation id="37862D3D4EBF914B07A97C16365B35ED" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Baetis rifensis sp. nov., larva morphology a leg b femur c femur dorsal margin d tibia ventral margin e claw." figureDoi="10.3897/zookeys.1180.109298.figure7" httpUri="https://binary.pensoft.net/fig/905324" pageId="0" pageNumber="27">, 7</figureCitation>
<figureCitation id="B699A87ADB1452B1F873C1F354EF0E11" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Baetis rifensis sp. nov., larva morphology abdominal terga V." figureDoi="10.3897/zookeys.1180.109298.figure8" httpUri="https://binary.pensoft.net/fig/905325" pageId="0" pageNumber="27">, 8</figureCitation>
<figureCitation id="E53AEBD8C28548029BBCA687D27BE7DC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Baetis rifensis sp. nov., larva morphology a paraprocts b protogonostylus c gill." figureDoi="10.3897/zookeys.1180.109298.figure9" httpUri="https://binary.pensoft.net/fig/905326" pageId="0" pageNumber="27">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8FE3FDE21DB80BC43CCDF820A467945F" pageId="0" pageNumber="27" type="type material">
<paragraph id="E0545D86C17498C8113EE50AB605A361" pageId="0" pageNumber="27">Type materials.</paragraph>
<paragraph id="4FD5AAE4A69E916A0A26F7DD485CC215" pageId="0" pageNumber="27">
<emphasis id="1DC621AD349B3822595A39B7ADEF0750" bold="true" pageId="0" pageNumber="27">
<emphasis id="541056A8C1DA0F09994A3A93F063A277" italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
.
</emphasis>
Morocco • 1 nymph; Tetouan Province, S1 Oued El Hamma, Loc. Jbel Laalam;
<geoCoordinate id="B568FD8D69CA52C2F65F763178FF7091" degrees="35" direction="north" minutes="23" orientation="latitude" precision="1" seconds="23.2" value="35.38978">35°23'23.2&quot;N</geoCoordinate>
,
<geoCoordinate id="9FA0BDFEEC12CCBAE34963EC184CD212" degrees="5" direction="west" minutes="29" orientation="longitude" precision="1" seconds="57.2" value="-5.499222">5°29'57.2&quot;W</geoCoordinate>
; alt. 200 m; 20.II.2022; El Yaagoubi leg.; DNA; GBIFCH01144399; MZL.
</paragraph>
<paragraph id="82B3A7FA2B6C99AC8065F6C67071085C" pageId="0" pageNumber="27">
<emphasis id="03F682CD9E60265D1D0877CC0C59E722" bold="true" pageId="0" pageNumber="27">
<emphasis id="AF713BB1718553EBE805F2B183BA15D2" italics="true" pageId="0" pageNumber="27">Paratypes</emphasis>
.
</emphasis>
Morocco • 38 larvae; same data as holotype; 25.III.2023; El Yaagoubi leg.; 3 on slide; 35 larvae on alcohol; LESCB • 19 nymphs; same data as holotype; 20.IV.2021; El Yaagoubi leg.; 2 on slide; LESCB and 1 larva for DNA; GBIFCH01144184; MZL • 6 nymphs; Tetouan Province, S2 Oued Taida, Loc. Beni idder;
<geoCoordinate id="049AEEEC33086457AFE0F4A687540D1F" degrees="35" direction="north" minutes="22" orientation="latitude" precision="1" seconds="6.10" value="35.36836">35°22'6.10&quot;N</geoCoordinate>
,
<geoCoordinate id="C4CCE2000DB09B2A7ECDA2B509121379" degrees="5" direction="west" minutes="32" orientation="longitude" precision="1" seconds="16.99" value="-5.5380526">5°32'16.99&quot;W</geoCoordinate>
; alt. 507 m; 28.IV.2017; El Alami leg.; DNA; GBIFCH01144398; MZL • 1 larva; Tetouan Province, S2 Oued Taida, Loc. Beni idder;
<geoCoordinate id="ECFAB5874E651958CA6610247C76529D" degrees="35" direction="north" minutes="22" orientation="latitude" precision="1" seconds="6.10" value="35.36836">35°22'6.10&quot;N</geoCoordinate>
,
<geoCoordinate id="94EC1F9DF8CA98DFD39E842DF3D8A505" degrees="5" direction="west" minutes="32" orientation="longitude" precision="1" seconds="16.99" value="-5.5380526">5°32'16.99&quot;W</geoCoordinate>
; alt. 507 m; 04.VI.2014; El Bazi leg.; LESCB • 8 nymphs; Chefchaouen Province, S3 Oued Kelaa, Loc. Akchour;
<geoCoordinate id="B3702765DAE40906D8B176D571F7C977" degrees="35" direction="north" minutes="14" orientation="latitude" precision="1" seconds="25.6" value="35.240444">35°14'25.6&quot;N</geoCoordinate>
,
<geoCoordinate id="A28ED15DB2AF5F284B9636657FD9CD2D" degrees="5" direction="west" minutes="9" orientation="longitude" precision="1" seconds="46.7" value="-5.1629725">5°9'46.7&quot;W</geoCoordinate>
; alt. 460 m; 13.III.2021; El Yaagoubi leg.; 1 on slide; LESCB; DNA; GBIFCH01144199; MZL • 7 larvae; Chefchaouen Province, S3 Oued Kelaa, Loc. Akchour;
<geoCoordinate id="BB339B10279D53147093BD9C993FF2C8" degrees="35" direction="north" minutes="14" orientation="latitude" precision="1" seconds="25.6" value="35.240444">35°14'25.6&quot;N</geoCoordinate>
, 5°9'46.7&quot; W; alt. 460 m; 20.IX.2014; Khadri leg.; LESCB • 6 larvae; Chefchaouen Province, S4 Oued El Kannar I, Loc. Souk Lhad;
<geoCoordinate id="28CF769C9F07B201527F736FDCFEF391" degrees="35" direction="north" minutes="13" orientation="latitude" precision="1" seconds="1.7" value="35.21714">35°13'1.7&quot;N</geoCoordinate>
,
<geoCoordinate id="FD68731F137EACA08D609A5005766607" degrees="5" direction="west" minutes="1" orientation="longitude" precision="1" seconds="24.02" value="-5.0233393">5°1'24.02&quot;W</geoCoordinate>
; alt. 105 m; 17.V.2014; Khadri leg.; LESCB • 4 larvae; Chefchaouen Province, S5 Oued El Kannar II, Loc. Assoul;
<geoCoordinate id="88A7D5EC4B5A19CBC8CF0B2E0A05BD8A" degrees="35" direction="north" minutes="13" orientation="latitude" precision="1" seconds="17.0" value="35.22139">35°13'17.0&quot;N</geoCoordinate>
,
<geoCoordinate id="7F8411AD073B29D4410CF40149814FC8" degrees="5" direction="west" minutes="0" orientation="longitude" precision="1" seconds="47.2" value="-5.013111">5°0'47.2&quot;W</geoCoordinate>
; alt. 68 m; 18.VI.2014; Khadri leg.; LESCB • 2 larvae; Chefchaouen Province, S6 Oued Bouhiya, Loc. Silloufene;
<geoCoordinate id="62007D2085219FEBB7E120EA20854278" degrees="35" direction="north" minutes="17" orientation="latitude" precision="1" seconds="53.4" value="35.298164">35°17'53.4&quot;N</geoCoordinate>
,
<geoCoordinate id="993E039B34C02AED39F92A3DDE5FDC40" degrees="4" direction="west" minutes="58" orientation="longitude" precision="1" seconds="26.3" value="-4.9739723">4°58'26.3&quot;W</geoCoordinate>
; alt. 35 m; 17.V.2014; Khadri leg.; LESCB • 1 larva; Chefchaouen Province, S7 Oued Amter I, Loc. Amter;
<geoCoordinate id="DF8D8C3B9CE6A9B317E4DF9FAEAF7F2C" degrees="35" direction="north" minutes="13" orientation="latitude" precision="1" seconds="44.3" value="35.228973">35°13'44.3&quot;N</geoCoordinate>
,
<geoCoordinate id="0FE50D59F99A746F5555745D79722607" degrees="4" direction="west" minutes="48" orientation="longitude" precision="1" seconds="42.9" value="-4.811917">4°48'42.9&quot;W</geoCoordinate>
; alt. 40 m; 17.III.2014; Khadri leg.; LESCB • 5 larvae; Chefchaouen Province, S8 Oued Amter II, Loc. Amter;
<geoCoordinate id="F683A8FD91099DF4D1798C6455B86B00" degrees="35" direction="north" minutes="14" orientation="latitude" precision="1" seconds="39.6" value="35.244335">35°14'39.6&quot;N</geoCoordinate>
,
<geoCoordinate id="80943E68BB75E10333660C8BCD58B352" degrees="4" direction="west" minutes="48" orientation="longitude" precision="1" seconds="12.6" value="-4.8035">4°48'12.6&quot;W</geoCoordinate>
; alt. 10 m; 17.III.2014; Khadri leg.; LESCB • 4 larvae; Chefchaouen Province, S9 Oued Ouringa I, Loc. Jebha;
<geoCoordinate id="B52C93F520B596FD360D7C0FDA4518FC" degrees="35" direction="north" minutes="8" orientation="latitude" precision="1" seconds="03.4" value="35.13428">35°8'03.4&quot;N</geoCoordinate>
,
<geoCoordinate id="35E89BE1BBFEA6824727D87A386B10B1" degrees="4" direction="west" minutes="42" orientation="longitude" precision="1" seconds="9.4" value="-4.702611">4°42'9.4&quot;W</geoCoordinate>
; alt. 95 m; 17.III.2014 • 1 larva; same data as holotype; 17.V.2014; Khadri leg.; LESCB • 3 larvae; Chefchaouen Province, S10 Oued Ouringa II, Loc. Jebha;
<geoCoordinate id="A2035DC1CCFDD1F28D01F84B61359781" degrees="35" direction="north" minutes="10" orientation="latitude" precision="1" seconds="14.2" value="35.170612">35°10'14.2&quot;N</geoCoordinate>
,
<geoCoordinate id="1ABCBDAFFB7C2ECE4F31FC1C4818E8F2" degrees="4" direction="west" minutes="41" orientation="longitude" precision="1" seconds="47.6" value="-4.6965556">4°41'47.6&quot;W</geoCoordinate>
; alt. 60 m; 17.V.2014; Khadri leg.; LESCB • 7 larvae; Chefchaouen Province, S11 Oued Ouringa III, Loc. Jebha;
<geoCoordinate id="5450B181991FD86D6E8BD923B9D25B8C" degrees="35" direction="north" minutes="11" orientation="latitude" precision="1" seconds="11.2" value="35.186447">35°11'11.2&quot;N</geoCoordinate>
,
<geoCoordinate id="12EA73811AF7218610E0BB42122A0705" degrees="4" direction="west" minutes="41" orientation="longitude" precision="1" seconds="13.5" value="-4.6870832">4°41'13.5&quot;W</geoCoordinate>
; alt. 25 m; 17.III.2014; Khadri leg.; LESCB • 1 larva; same data as holotype; 17.V.2014; Khadri leg.; LESCB • 3 larvae; Chefchaouen Province, S12 Oued Mlilah, Loc. Derdara;
<geoCoordinate id="AF4AE9889AE3A17DE564D803D66A4491" degrees="35" direction="north" minutes="2" orientation="latitude" precision="1" seconds="35.0" value="35.043056">35°2'35.0&quot;N</geoCoordinate>
,
<geoCoordinate id="430AC66F6F167B886EC14F3BA4E9EF09" degrees="5" direction="west" minutes="21" orientation="longitude" precision="1" seconds="13.0" value="-5.353611">5°21'13.0&quot;W</geoCoordinate>
; alt. 198 m; 9.V.2015; El Bazi leg.; LESCB • 6 nymphs; Chefchaouen Province, S13 Oued Mansoura, Loc. Tanaqoub;
<geoCoordinate id="68BCE334AC2826F59035E12BD44485E6" degrees="35" direction="north" minutes="5" orientation="latitude" precision="1" seconds="16.0" value="35.087776">35°5'16.0&quot;N</geoCoordinate>
,
<geoCoordinate id="1042EC50B496D14D3342955E0F2EFB55" degrees="5" direction="west" minutes="30" orientation="longitude" precision="1" seconds="37.0" value="-5.5102777">5°30'37.0&quot;W</geoCoordinate>
; alt. 124 m; 1.IV.2021; El Yaagoubi leg.; 1 on slide; LESCB; DNA; GBIFCH01144168; MZL • 6 larvae; Larache Province, S14 Oued Sghir, Loc.
<normalizedToken id="8869DBDD814C0EA1CDE883214426D2EF" originalValue="Béni">Beni</normalizedToken>
Arouss;
<geoCoordinate id="C42FC706CE8D2C0A3BCAE89633FE02C1" degrees="35" direction="north" minutes="16" orientation="latitude" precision="1" seconds="1.63" value="35.26712">35°16'1.63&quot;N</geoCoordinate>
,
<geoCoordinate id="E10701C91768191428A7BB8BC14B2AAF" degrees="5" direction="west" minutes="37" orientation="longitude" precision="1" seconds="10.9" value="-5.6196947">5°37'10.9&quot;W</geoCoordinate>
; alt. 200 m; 02.V.2022; El Yaagoubi leg.; LESCB.
</paragraph>
</subSubSection>
<subSubSection id="8128F30D8F9A0D23B3A1E71FDA47FC4C" pageId="0" pageNumber="27" type="description">
<paragraph id="497DC5D3FBCC653627A49EAC1E6A0D1D" pageId="0" pageNumber="27">Description.</paragraph>
<paragraph id="C73F697D94228EE705D68D76B55A7DE3" pageId="0" pageNumber="27">
<emphasis id="D218D4CF1FF68E4C1DD5A634DC67B749" bold="true" pageId="0" pageNumber="27">Larva</emphasis>
(Figs
<figureCitation id="CCA0385EE6EAB3D86536A8C47DFE8CC0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Baetis rifensis sp. nov., larva habitus a female, dorsal view b male, dorsal view c male, lateral view." figureDoi="10.3897/zookeys.1180.109298.figure2" httpUri="https://binary.pensoft.net/fig/905319" pageId="0" pageNumber="27">2</figureCitation>
-
<figureCitation id="96DED44ED992C63A2B8C229A2B179613" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Baetis rifensis sp. nov., larva morphology a paraprocts b protogonostylus c gill." figureDoi="10.3897/zookeys.1180.109298.figure9" httpUri="https://binary.pensoft.net/fig/905326" pageId="0" pageNumber="27">9</figureCitation>
). Female body length 6.0-7.8 mm; cerci 2.0-2.8 mm; median caudal filament ca 2/3 of cerci. Male body length 6.0-7.2 mm; cerci 2.2-2.8 mm; median caudal filament ca 2/3 of cerci.
</paragraph>
<paragraph id="87E731F9FBA9EB832B923E46CD3FA665" pageId="0" pageNumber="27">
<emphasis id="CAD9EEB5D468C90726E6DA0D5BCAB371" bold="true" italics="true" pageId="0" pageNumber="27">Colouration</emphasis>
(Fig.
<figureCitation id="5BEFC34D9150FA5A351EDAA1975594D1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Baetis rifensis sp. nov., larva habitus a female, dorsal view b male, dorsal view c male, lateral view." figureDoi="10.3897/zookeys.1180.109298.figure2" httpUri="https://binary.pensoft.net/fig/905319" pageId="0" pageNumber="27">2a-c</figureCitation>
): general colouration yellowish brown.
<emphasis id="EE5920ED38C3D62DD38B9203A51E7123" bold="true" italics="true" pageId="0" pageNumber="27">Head</emphasis>
uniformly brown with vermiform, yellow marking along epicranial suture, sometimes diffused and inconspicuous.
<emphasis id="616CBF9EFEE2AD1A187D38DBE45AAB64" bold="true" italics="true" pageId="0" pageNumber="27">Pronotum</emphasis>
brown with dispersed swirling yellowish marks.
<emphasis id="09D8F2EB6BA7716F7DC5BCFFB6F7894C" bold="true" italics="true" pageId="0" pageNumber="27">Mesonotum</emphasis>
brown with yellowish spots.
<emphasis id="D9D4C18246958A0E57054169DB25442F" bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
pale yellow, with a yellowish-brown patch in centre of femur and a submarginal darker stripe near outer margin of femur.
<emphasis id="47207A9A38524F8F00EFB0136B22CF90" bold="true" italics="true" pageId="0" pageNumber="27">Abdominal tergites</emphasis>
medium brown and pale yellow with two central, darker brown spots: tergite I yellow; tergites II-IV brown with lateral yellow spots; tergite V yellow with lateral brown spots; tergites VI-VIII brown, yellowish distolaterally; tergites IX-X yellow; tergite X brownish apically.
<emphasis id="0B643A086D77DA13DE0A4CB8611AC12B" bold="true" italics="true" pageId="0" pageNumber="27">Cerci and paracercus</emphasis>
yellowish or whitish yellow, with a conspicuous, dark transversal band near middle and darker apically. Basal segments sometimes also darker.
</paragraph>
<caption id="51085796095F47AE69BEF3C9F7348D4E" doi="10.3897/zookeys.1180.109298.figure2" httpUri="https://binary.pensoft.net/fig/905319" pageId="0" pageNumber="27" start="Figure 2" startId="F2">
<paragraph id="FB95D102D20A24E956BFC38E19F3981D" pageId="0" pageNumber="27">
<emphasis id="BF5AA8965693A50535AD98F757F65012" bold="true" pageId="0" pageNumber="27">Figure 2.</emphasis>
<taxonomicName id="717972D6EFE0AF4EF93A1F1C865D6D96" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="2975E3745E0BEDE7CD8305618A6B36AD" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva habitus
<emphasis id="70E4F2DC74D72E27610C1DD45A46F83E" bold="true" pageId="0" pageNumber="27">a</emphasis>
female, dorsal view
<emphasis id="F519CABD2A345257510F5EB05190D912" bold="true" pageId="0" pageNumber="27">b</emphasis>
male, dorsal view
<emphasis id="6ECE40C931D623B241CF28601FAC3A03" bold="true" pageId="0" pageNumber="27">c</emphasis>
male, lateral view.
</paragraph>
</caption>
<paragraph id="11C2160B90C9EEEF60FF5B6CD20B2E22" pageId="0" pageNumber="27">
<emphasis id="6C9913C9B20E6B334755EFD2C81BF01D" bold="true" italics="true" pageId="0" pageNumber="27">Labrum</emphasis>
(Fig.
<figureCitation id="6E398A44D2BD0FDDF260EA8B5F816282" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baetis rifensis sp. nov., larva morphology a labrum dorsal face b labrum ventral face." figureDoi="10.3897/zookeys.1180.109298.figure3" httpUri="https://binary.pensoft.net/fig/905320" pageId="0" pageNumber="27">3a, b</figureCitation>
) with 1 + (5-6) long setae, almost symmetric, evenly rounded laterally, and about 1/3 broader than long. Dorsal surface (Fig.
<figureCitation id="7B1C03D1AC204AFDF40DD3A864D4A0CA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baetis rifensis sp. nov., larva morphology a labrum dorsal face b labrum ventral face." figureDoi="10.3897/zookeys.1180.109298.figure3" httpUri="https://binary.pensoft.net/fig/905320" pageId="0" pageNumber="27">3a</figureCitation>
) scattered with long, pointed setae; ventral surface with apicolateral patch of long, bifid setae near margin (Fig.
<figureCitation id="79EB3EDCB35B11DD9AF72309306BE051" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Baetis rifensis sp. nov., larva morphology a labrum dorsal face b labrum ventral face." figureDoi="10.3897/zookeys.1180.109298.figure3" httpUri="https://binary.pensoft.net/fig/905320" pageId="0" pageNumber="27">3b</figureCitation>
).
</paragraph>
<caption id="6295750BB0E2E260519BAE19F1BF3BFB" doi="10.3897/zookeys.1180.109298.figure3" httpUri="https://binary.pensoft.net/fig/905320" pageId="0" pageNumber="27" start="Figure 3" startId="F3">
<paragraph id="B73B25E1CC604AD979A6C79B65266175" pageId="0" pageNumber="27">
<emphasis id="3B132DED9D3088DB171E6EB172E6B04D" bold="true" pageId="0" pageNumber="27">Figure 3.</emphasis>
<taxonomicName id="1D6076CDD30C09AB93E39079F5DCD685" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="36B4E9F9033F4101A937547C14DBEA08" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis id="A63C2F5CB45E30A1341D6427E9980C53" bold="true" pageId="0" pageNumber="27">a</emphasis>
labrum dorsal face
<emphasis id="01EE28BCCDDFDF884A1371412C4644DC" bold="true" pageId="0" pageNumber="27">b</emphasis>
labrum ventral face.
</paragraph>
</caption>
<paragraph id="61F510B1446B67ED218FCDFBFAC250D9" pageId="0" pageNumber="27">
<emphasis id="A402451A52E904A16B5CD7B9B750D49F" bold="true" italics="true" pageId="0" pageNumber="27">Right mandible</emphasis>
(Fig.
<figureCitation id="A4664D72915A554A77F24C02D254946A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Baetis rifensis sp. nov., larva morphology a, b right mandible c, d left mandible." figureDoi="10.3897/zookeys.1180.109298.figure4" httpUri="https://binary.pensoft.net/fig/905321" pageId="0" pageNumber="27">4a, b</figureCitation>
): incisor and kinetodontium fused. Incisor with three apically rounded or pointed teeth subequal in size; outer tooth slightly broader. Kinetodontium with four teeth; inner margin of innermost tooth denticulate. Prostheca slender, apically pectinate.
</paragraph>
<caption id="1ECA51C6B37270D6B7120492F1ED78CB" doi="10.3897/zookeys.1180.109298.figure4" httpUri="https://binary.pensoft.net/fig/905321" pageId="0" pageNumber="27" start="Figure 4" startId="F4">
<paragraph id="120F8CFA411F3FBE9D05BD85498D65C7" pageId="0" pageNumber="27">
<emphasis id="AA94ACA6CFEBACB233350A12B0A1A7AF" bold="true" pageId="0" pageNumber="27">Figure 4.</emphasis>
<taxonomicName id="EF2A637D843609AB44C18431B98E1034" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="AA821362A1571CB4E2C95590B2FEC46A" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis id="346B4AFA6D899FCA1EC8656DDCEBD660" bold="true" pageId="0" pageNumber="27">a, b</emphasis>
right mandible
<emphasis id="65FEF7AEEC4DAFD2A62F81C049207EF2" bold="true" pageId="0" pageNumber="27">c, d</emphasis>
left mandible.
</paragraph>
</caption>
<paragraph id="EE733BE7DA210604018D522AAEB2A314" pageId="0" pageNumber="27">
<emphasis id="69049D634016B5DDE54B7AC2FD08AB38" bold="true" italics="true" pageId="0" pageNumber="27">Left mandible</emphasis>
(Fig.
<figureCitation id="B75411E38A28B14FDB2E754B8ABD02D9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Baetis rifensis sp. nov., larva morphology a, b right mandible c, d left mandible." figureDoi="10.3897/zookeys.1180.109298.figure4" httpUri="https://binary.pensoft.net/fig/905321" pageId="0" pageNumber="27">4c, d</figureCitation>
): incisor and kinetodontium fused. Incisor with three apically rounded or pointed teeth subequal in size; outer tooth slightly broader. Kinetodontium with three teeth; inner margin of innermost tooth denticulate. Prostheca robust, apically with stout denticles and a comb-like structure.
</paragraph>
<paragraph id="F66AB094640DA09262F34996B7F113F7" pageId="0" pageNumber="27">
<emphasis id="3E41795EECC7A730E353AC8F7135ED31" bold="true" italics="true" pageId="0" pageNumber="27">Maxilla</emphasis>
(Fig.
<figureCitation id="627B9DDBBD72ACD0E99F5A708BE16905" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Baetis rifensis sp. nov., larva morphology a maxilla b apex of maxilla, c apex of maxillary palp, d hypopharynx." figureDoi="10.3897/zookeys.1180.109298.figure5" httpUri="https://binary.pensoft.net/fig/905322" pageId="0" pageNumber="27">5a, b</figureCitation>
): galea-lacinia ventrally with two simple setae under teeth. Inner dorsal row of setae with three denti-setae; distal denti-setae tooth-like; middle and proximal denti-setae slender, bifid, and pectinate.
</paragraph>
<caption id="FF3077DC79469CC39005BE822F44E463" doi="10.3897/zookeys.1180.109298.figure5" httpUri="https://binary.pensoft.net/fig/905322" pageId="0" pageNumber="27" start="Figure 5" startId="F5">
<paragraph id="9633B768885D9C705BED0129C7CAD233" pageId="0" pageNumber="27">
<emphasis id="1ADA18C4DCFB0D6F4EADE8474F445BC9" bold="true" pageId="0" pageNumber="27">Figure 5.</emphasis>
<taxonomicName id="42E9F6154BB7BD9C33213EC8E470C0E5" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="B0B67646DA82403153FB237BD4EBFA0A" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis id="E1B4BD52D91ABD165AE4D08F02039E34" bold="true" pageId="0" pageNumber="27">a</emphasis>
maxilla
<emphasis id="CB5228F4BA441D4CF0655D82E4FB1CB3" bold="true" pageId="0" pageNumber="27">b</emphasis>
apex of maxilla,
<emphasis id="14941339C26D261563E025306CFC3C8D" bold="true" pageId="0" pageNumber="27">c</emphasis>
apex of maxillary palp,
<emphasis id="404AD7CE1F6316FF315F64B3898F02E4" bold="true" pageId="0" pageNumber="27">d</emphasis>
hypopharynx.
</paragraph>
</caption>
<paragraph id="78FA42368C2732866FB663D474F7A6BE" pageId="0" pageNumber="27">
<emphasis id="2193EFD095B20CC3CF9ADCFBBD8E724C" bold="true" italics="true" pageId="0" pageNumber="27">Maxillary palp</emphasis>
(Fig.
<figureCitation id="385927993AA9BB0C58763F30547E92E4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Baetis rifensis sp. nov., larva morphology a maxilla b apex of maxilla, c apex of maxillary palp, d hypopharynx." figureDoi="10.3897/zookeys.1180.109298.figure5" httpUri="https://binary.pensoft.net/fig/905322" pageId="0" pageNumber="27">5c</figureCitation>
) slightly longer than length of galea-lacinia; palp segment II 1.2
<normalizedToken id="CFF72A2F52CED4D6B69400CEEFCE65AD" originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface; apex of last segment rounded.
</paragraph>
<paragraph id="2C5C73B7E28A4B84581A329CA01FBC0A" pageId="0" pageNumber="27">
<emphasis id="E5E2C7AD2C396D328479EDD3847659DB" bold="true" italics="true" pageId="0" pageNumber="27">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation id="6B7579BCE8BEAB932FDC124D55688336" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Baetis rifensis sp. nov., larva morphology a maxilla b apex of maxilla, c apex of maxillary palp, d hypopharynx." figureDoi="10.3897/zookeys.1180.109298.figure5" httpUri="https://binary.pensoft.net/fig/905322" pageId="0" pageNumber="27">5d</figureCitation>
): Lingua trilobed, apically covered with thin setae; superlingua subequal to lingua. Lingua longer than broad; medial tuft of stout setae well developed, broad; distal half laterally expanded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph id="F36C5FB5B0D482A9829613906A4170E0" pageId="0" pageNumber="27">
<emphasis id="3239B198D89BFC8DCF637B18B04B7788" bold="true" italics="true" pageId="0" pageNumber="27">Labium</emphasis>
(Fig.
<figureCitation id="1C2B84E2B279154BFA699FE2CF294ABC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Baetis rifensis sp. nov., larva morphology a labial palp b paraglossae." figureDoi="10.3897/zookeys.1180.109298.figure6" httpUri="https://binary.pensoft.net/fig/905323" pageId="0" pageNumber="27">6a, b</figureCitation>
): glossa shorter than paraglossae; distal and lateral margins with long stout simple setae. Paraglossa (Fig.
<figureCitation id="B7EB3FB8840A2ABC0C23E9DA2B70FAA5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Baetis rifensis sp. nov., larva morphology a labial palp b paraglossae." figureDoi="10.3897/zookeys.1180.109298.figure6" httpUri="https://binary.pensoft.net/fig/905323" pageId="0" pageNumber="27">6b</figureCitation>
) apically with three rows of long, simple, stout setae; dorsal surface with a row of five long, stout setae. Labial palps with a distinctly curved outer margin with segment I 0.7
<normalizedToken id="1D56CC57359713224D4272CE7610598E" originalValue="×">x</normalizedToken>
length of segments II and III combined, segment II 1.2
<normalizedToken id="0D5058E6C1ED6CABE9E828AB6CD8F1D3" originalValue="×">x</normalizedToken>
length of segment I; fine, simple setae scattered over surface of segments I and II; segment II with a small, digitiform, apical projection; segment III slightly asymmetric, broadly rounded and about 1.5
<normalizedToken id="158B1C215AAEED632F1004C05F990091" originalValue="×">x</normalizedToken>
broader than long; covered stout medium setae (Fig.
<figureCitation id="2705066CAB261766E33B30A50F1540EF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Baetis rifensis sp. nov., larva morphology a labial palp b paraglossae." figureDoi="10.3897/zookeys.1180.109298.figure6" httpUri="https://binary.pensoft.net/fig/905323" pageId="0" pageNumber="27">6a</figureCitation>
).
</paragraph>
<caption id="EA367A49A220B859D6B9D4AD64FE8667" doi="10.3897/zookeys.1180.109298.figure6" httpUri="https://binary.pensoft.net/fig/905323" pageId="0" pageNumber="27" start="Figure 6" startId="F6">
<paragraph id="A18875180EB42A964CACA03064FCE646" pageId="0" pageNumber="27">
<emphasis id="273D1273FD67CC52F6B90D4BC2ED8E94" bold="true" pageId="0" pageNumber="27">Figure 6.</emphasis>
<taxonomicName id="F0B0B6E18F9A12D6CCF1C8D9389D9EAD" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="B8DAF79BBBB86FCA1C2FFE96F66C0796" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis id="3F253CFFA4BD3916879199AB2C9DC86F" bold="true" pageId="0" pageNumber="27">a</emphasis>
labial palp
<emphasis id="5971A98796C2E0BE5D11095F82D09747" bold="true" pageId="0" pageNumber="27">b</emphasis>
paraglossae.
</paragraph>
</caption>
<paragraph id="E54966731376093DF6E525E2387A3F27" pageId="0" pageNumber="27">
<emphasis id="5DC2590FB21BAFDD0E665E43C8215B6F" bold="true" italics="true" pageId="0" pageNumber="27">Foreleg</emphasis>
(Fig.
<figureCitation id="CA27DCBEDF6F61ACB6DC9E499B9F7E9A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Baetis rifensis sp. nov., larva morphology a leg b femur c femur dorsal margin d tibia ventral margin e claw." figureDoi="10.3897/zookeys.1180.109298.figure7" httpUri="https://binary.pensoft.net/fig/905324" pageId="0" pageNumber="27">7a-e</figureCitation>
): ratio of foreleg segments 1.2:0.8:0.6:0.4.
<emphasis id="386C473348644D9E21BD648B5D2AD240" bold="true" italics="true" pageId="0" pageNumber="27">Femur</emphasis>
(Fig.
<figureCitation id="143C9170F7B4AC98866ECF0557FEC55B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Baetis rifensis sp. nov., larva morphology a leg b femur c femur dorsal margin d tibia ventral margin e claw." figureDoi="10.3897/zookeys.1180.109298.figure7" httpUri="https://binary.pensoft.net/fig/905324" pageId="0" pageNumber="27">7b</figureCitation>
) length 3.5
<normalizedToken id="D0F7CDD4BE77E7575979A7B039994440" originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with spatulate setae (Fig.
<figureCitation id="6DC0DD30292115C3320249B6EBB874AA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Baetis rifensis sp. nov., larva morphology a leg b femur c femur dorsal margin d tibia ventral margin e claw." figureDoi="10.3897/zookeys.1180.109298.figure7" httpUri="https://binary.pensoft.net/fig/905324" pageId="0" pageNumber="27">7c</figureCitation>
), very abundant proximally, scarce distally; ventral and lateral with scattered short stout setae.
<emphasis id="368195C2C3C2C4209FF423BA42415879" bold="true" italics="true" pageId="0" pageNumber="27">Tibia</emphasis>
: dorsal margin almost bare; ventral margin with short, stout, apically pointed setae (Fig.
<figureCitation id="92604237B46434D1DE9203C85D7465A0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Baetis rifensis sp. nov., larva morphology a leg b femur c femur dorsal margin d tibia ventral margin e claw." figureDoi="10.3897/zookeys.1180.109298.figure7" httpUri="https://binary.pensoft.net/fig/905324" pageId="0" pageNumber="27">7d</figureCitation>
), and some fine, simple setae.
<emphasis id="41AE78802BD67C0487278E12A2D4BEBC" bold="true" italics="true" pageId="0" pageNumber="27">Tarsus</emphasis>
: dorsal margin almost bare; ventral margin with row of curved, short to medium-length spine-like setae.
<emphasis id="B15F627135C8BFFE2CFFBAF70B9F834B" bold="true" italics="true" pageId="0" pageNumber="27">Claw</emphasis>
(Fig.
<figureCitation id="8CD5CE59D54D736C83F585B44716E006" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Baetis rifensis sp. nov., larva morphology a leg b femur c femur dorsal margin d tibia ventral margin e claw." figureDoi="10.3897/zookeys.1180.109298.figure7" httpUri="https://binary.pensoft.net/fig/905324" pageId="0" pageNumber="27">7e</figureCitation>
) with one row of 12-14 denticles, distally pointed; subapical setae absent.
</paragraph>
<caption id="B8744A8B3F2BF0ACB7BE3796ED3FF7C3" doi="10.3897/zookeys.1180.109298.figure7" httpUri="https://binary.pensoft.net/fig/905324" pageId="0" pageNumber="27" start="Figure 7" startId="F7">
<paragraph id="7F102384995D241E5A58DCFC4B3124E8" pageId="0" pageNumber="27">
<emphasis id="BA5FDC1DDDFDB6EA2141E062481D60E5" bold="true" pageId="0" pageNumber="27">Figure 7.</emphasis>
<taxonomicName id="E6E6E2E87C95439376819B1A785FE6B8" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="B784AE734F483107302DD7949D6F9773" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis id="0462C3A44ECE4B11E3BA48EA6B004256" bold="true" pageId="0" pageNumber="27">a</emphasis>
leg
<emphasis id="7DC6280FBA3674CEA1A4DEAA16E20DCF" bold="true" pageId="0" pageNumber="27">b</emphasis>
femur
<emphasis id="4A408496FB15A30D5B0136DFD4BFCAF3" bold="true" pageId="0" pageNumber="27">c</emphasis>
femur dorsal margin
<emphasis id="8F0AF758705E73951FA6D2A74231800A" bold="true" pageId="0" pageNumber="27">d</emphasis>
tibia ventral margin
<emphasis id="AA600BE22C6D590DCDC26BE28F792426" bold="true" pageId="0" pageNumber="27">e</emphasis>
claw.
</paragraph>
</caption>
<paragraph id="70D11371D6375B8D5DBF74B1CDCD7DA9" pageId="0" pageNumber="27">
<emphasis id="AF9879B85509870AFDD5A6C8D4DA41CA" bold="true" italics="true" pageId="0" pageNumber="27">Abdominal terga</emphasis>
(Fig.
<figureCitation id="CFAE1D53457BA8DD6CB8EB3EF87F1339" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Baetis rifensis sp. nov., larva morphology abdominal terga V." figureDoi="10.3897/zookeys.1180.109298.figure8" httpUri="https://binary.pensoft.net/fig/905325" pageId="0" pageNumber="27">8</figureCitation>
): surface covered by scale bases and a few setae. Distal margins of tergites with triangular, pointed spines, longer than wide.
</paragraph>
<caption id="B680F35476BD2008097BA5C21073C33B" doi="10.3897/zookeys.1180.109298.figure8" httpUri="https://binary.pensoft.net/fig/905325" pageId="0" pageNumber="27" start="Figure 8" startId="F8">
<paragraph id="A54B31FFAC114D7D9ECB014247E74296" pageId="0" pageNumber="27">
<emphasis id="46FEE488B9CCBBD33AA5BF87552D74DD" bold="true" pageId="0" pageNumber="27">Figure 8.</emphasis>
<taxonomicName id="E0C8866DDF420596040B1364EA4BE0B7" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="E5ABB856265A88E07771F304EE0A0A7F" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva morphology abdominal terga V.
</paragraph>
</caption>
<paragraph id="E5D52689FDB8A70AB730362837CB2037" pageId="0" pageNumber="27">
<emphasis id="F266FF7AB4CB0C73226175FE840DA7F3" bold="true" italics="true" pageId="0" pageNumber="27">Paraproct</emphasis>
(Fig.
<figureCitation id="62811426FAB25869A35389FCD0D59448" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Baetis rifensis sp. nov., larva morphology a paraprocts b protogonostylus c gill." figureDoi="10.3897/zookeys.1180.109298.figure9" httpUri="https://binary.pensoft.net/fig/905326" pageId="0" pageNumber="27">9a</figureCitation>
): surface with few scale bases and broad setae; margin with ca 20 long, slender spines. Cercotractor, margin with ca 15 medium-length spines.
</paragraph>
<caption id="5CF8DB3BD5B555AC5F9E3DC176D98CF5" doi="10.3897/zookeys.1180.109298.figure9" httpUri="https://binary.pensoft.net/fig/905326" pageId="0" pageNumber="27" start="Figure 9" startId="F9">
<paragraph id="EFC1E6D9D8D304FE52BD11339B8C61AF" pageId="0" pageNumber="27">
<emphasis id="3D77A2CD0F423247556A951D63F5B494" bold="true" pageId="0" pageNumber="27">Figure 9.</emphasis>
<taxonomicName id="C806268728BA01260A7849F2349213FA" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="78EEDB2F2D5A160A9806F7D6045B6DDA" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis id="D1C8DFC095E2952BA538304B3F795591" bold="true" pageId="0" pageNumber="27">a</emphasis>
paraprocts
<emphasis id="B8358734108CF4D2E564EC1D23FFD9AC" bold="true" pageId="0" pageNumber="27">b</emphasis>
protogonostylus
<emphasis id="4F080EA48006B7A5FDB63EF30E1573A4" bold="true" pageId="0" pageNumber="27">c</emphasis>
gill.
</paragraph>
</caption>
<paragraph id="8F6B082A4CC409916C36332A2B9C0C2D" pageId="0" pageNumber="27">
<emphasis id="E35AEAFC166E08166D28C4B3C130B116" bold="true" italics="true" pageId="0" pageNumber="27">Protogonostylus</emphasis>
(Fig.
<figureCitation id="6886E2C1E4353CB3E2C6E5EE5E907D40" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Baetis rifensis sp. nov., larva morphology a paraprocts b protogonostylus c gill." figureDoi="10.3897/zookeys.1180.109298.figure9" httpUri="https://binary.pensoft.net/fig/905326" pageId="0" pageNumber="27">9b</figureCitation>
): subimaginal gonostyli developing under cuticle of last instar larvae folded in the following way: segment II bent towards middle, last segment bent laterally.
</paragraph>
<paragraph id="1054D9A590F3451436D51BC90EF45AF7" pageId="0" pageNumber="27">
<emphasis id="5B33933E80D28C32308729449A34119F" bold="true" italics="true" pageId="0" pageNumber="27">Gills</emphasis>
(Fig.
<figureCitation id="DA5E54142D5E8A6E9CCD13FF00740973" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Baetis rifensis sp. nov., larva morphology a paraprocts b protogonostylus c gill." figureDoi="10.3897/zookeys.1180.109298.figure9" httpUri="https://binary.pensoft.net/fig/905326" pageId="0" pageNumber="27">9c</figureCitation>
): tracheae extending from main trunk to inner and outer margins. Serrated costal margin without spine-like setae.
</paragraph>
<paragraph id="81311546EABC2E89CE3C404BE147461E" pageId="0" pageNumber="27">
<emphasis id="BCDDB9E5A190A5D46646057A26085369" bold="true" pageId="0" pageNumber="27">Imagos.</emphasis>
Unknown.
</paragraph>
<paragraph id="BD2C3BF98D5E116F41D51EFBCFFC97B7" pageId="0" pageNumber="27">
<emphasis id="45E665BA394C22A9531DF67D1EEB2A5D" bold="true" pageId="0" pageNumber="27">Subimago.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="ABF68A3E45B8A09776196488753289BE" pageId="0" pageNumber="27" type="etymology">
<paragraph id="418DB61DF3E0B122A4680183BD280233" pageId="0" pageNumber="27">Etymology.</paragraph>
<paragraph id="260373D0C8AC2F88D64BD5936F03933E" pageId="0" pageNumber="27">The new species is named after the Moroccan Rif region, from where it was collected for the first time.</paragraph>
</subSubSection>
<subSubSection id="925B2EE2F5DFFB5106E1D905588895BC" pageId="0" pageNumber="27" type="distribution">
<paragraph id="27ACD82A1F1FB41CC1F282225246431B" pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph id="78D4ACAA7EAAC7E559D790A6202482B9" pageId="0" pageNumber="27">
Morocco: Rif (Fig.
<figureCitation id="0524E7CB63AE128B64546230D5741001" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Baetis rifensis sp. nov. (yellow star: type locality)." figureDoi="10.3897/zookeys.1180.109298.figure1" httpUri="https://binary.pensoft.net/fig/905318" pageId="0" pageNumber="27">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="17952842DD602C90C8FE9598B5678EE2" pageId="0" pageNumber="27" type="biology_ecology">
<paragraph id="BDEA0C9E9586C1C99FB674ADBEED31D7" pageId="0" pageNumber="27">Biology.</paragraph>
<paragraph id="2A5094B15A909C69D93138B01BC6812C" pageId="0" pageNumber="27">The specimens were captured from rocky bottoms of moderate- to slow-velocity streams and from rhithral to potamal portions of rivers.</paragraph>
</subSubSection>
<subSubSection id="58C50A9E1160853B855CE5FC37AB11D7" pageId="0" pageNumber="27" type="molecular analyses">
<paragraph id="709DCB8A3AE1212E2BAD76811F3BC1C9" pageId="0" pageNumber="27">Molecular analyses</paragraph>
<paragraph id="7141CDAE81A138BA135302D7634C1E2D" pageId="0" pageNumber="27">
The COI data set was&gt;95% complete and included 31% of parsimony-informative sites. Pairwise COI distances across all sequences ranged from 0 to 21.8%. The overall mean
<emphasis id="F863698460EAC95334A1D65CE63024AD" italics="true" pageId="0" pageNumber="27">p</emphasis>
-distance within MOTUs was 0.8% (mean range 0-2.1%), while the overall mean
<emphasis id="5FB199A1569FEBF430F6FCB55E8D0E30" italics="true" pageId="0" pageNumber="27">p</emphasis>
-distance between MOTUs was 15.4% (mean range 3.2-21.7%). The maximum
<emphasis id="EE17235E2A03BAE69A3B91473C2294D7" italics="true" pageId="0" pageNumber="27">p</emphasis>
-distance within MOTUs varied from 0 (
<taxonomicName id="43DF096883CC167D6AA95EBDB50487C8" authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="A4B1917B1807CA122344C4960522E42D" italics="true" pageId="0" pageNumber="27">Baetis</emphasis>
</taxonomicName>
sp. 2 and
<taxonomicName id="E41A62B993F2699376256651DCADA3F3" lsidName="B. fuscatus" pageId="0" pageNumber="27" rank="species" species="fuscatus">
<emphasis id="0B3D98F0673257C17AEE8E8BE013680F" italics="true" pageId="0" pageNumber="27">B. fuscatus</emphasis>
</taxonomicName>
5) to 4.3% (
<taxonomicName id="CDAEB23D6B02A33B2144C907028D96E2" lsidName="B. fuscatus" pageId="0" pageNumber="27" rank="species" species="fuscatus">
<emphasis id="82ADB220DB4ECB71370BEB00E1184AEF" italics="true" pageId="0" pageNumber="27">B. fuscatus</emphasis>
</taxonomicName>
1). The minimum
<emphasis id="0B13219FEF54DADD7EA1AA156D6FA371" italics="true" pageId="0" pageNumber="27">p</emphasis>
-distance between MOTUs ranged from 3.2% (
<taxonomicName id="D164D92E337AA5F8800C01CCD959E94C" lsidName="B. rifensis" pageId="0" pageNumber="27" rank="species" species="rifensis">
<emphasis id="9C65B5838A01C5D928357F7D76A209DF" italics="true" pageId="0" pageNumber="27">B. rifensis</emphasis>
</taxonomicName>
sp. nov.-
<taxonomicName id="15165CE02C13CC9261707836C67CE928" genus="B." lsidName="B." pageId="0" pageNumber="27" rank="genus">
<emphasis id="D36AD23A1A9F62AD2D6128663FF1FC8A" italics="true" pageId="0" pageNumber="27">B.</emphasis>
</taxonomicName>
sp. 1 and
<taxonomicName id="8A9496D0523BA1FDD39A258B2584E579" lsidName="B. rifensis" pageId="0" pageNumber="27" rank="species" species="rifensis">
<emphasis id="E1EF250B91C4A1AFB544A62DE4C81E80" italics="true" pageId="0" pageNumber="27">B. rifensis</emphasis>
</taxonomicName>
sp. nov.-
<taxonomicName id="DBD1BFF8C9A2ECB450AD2FFA804E1B71" genus="B." lsidName="B." pageId="0" pageNumber="27" rank="genus">
<emphasis id="FFD16DF95357631A36371B14FCF0C02F" italics="true" pageId="0" pageNumber="27">B.</emphasis>
</taxonomicName>
sp. 2) to 12.2% (
<taxonomicName id="C0E1F0CDD73079FEAC14613AF2F4A2B9" lsidName="B. rifensis" pageId="0" pageNumber="27" rank="species" species="rifensis">
<emphasis id="65D6C17E7B01C2C4353793BF95F49BE9" italics="true" pageId="0" pageNumber="27">B. rifensis</emphasis>
</taxonomicName>
sp. nov.-
<taxonomicName id="180B1C931B871C95C99EA990AE8A4622" lsidName="B. scambus" pageId="0" pageNumber="27" rank="species" species="scambus">
<emphasis id="2D769F40FC492A057EAC75B79015A604" italics="true" pageId="0" pageNumber="27">B. scambus</emphasis>
</taxonomicName>
). The five sequences from
<taxonomicName id="CE74F9E5D2C06369594EA8425860E7C7" lsidName="B. rifensis" pageId="0" pageNumber="27" rank="species" species="rifensis">
<emphasis id="C0B69EF397F94D377205135CCC960DCF" italics="true" pageId="0" pageNumber="27">B. rifensis</emphasis>
</taxonomicName>
sp. nov. formed a strongly supported monophyletic clade, identified as a distinct MOTU in both the ASAP and mPTP species-delimitation analyses (Fig.
<figureCitation id="BE336D925AE52949702A439F9BDA848B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Bayesian (BEAST) maximum clade credibility COI tree of the Baetis fuscatus group in the Western Palaearctic. Coloured vertical boxes indicate species-delimitation hypothesis (i. e., MOTUs) according to the mPTP and ASAP methods. For each ASAP-based MOTU, the corresponding species names (where available) and the country / region of origin are provided, with the newly described species specified in bold and the associated GBIF codes highlighted in pale pink. Tips labelled with GBIF codes indicate newly sequenced specimens; PT 002, ES 029, and ES 030 codes designate sequences from the project FREDIE; the GMGMQ 2692 - 18 code is from BOLD; the other codes correspond to sequences obtained from GenBank. The three lineages, Baetis rifensis sensu lato (s. l.), Baetis scambus s. l., and Baetis fuscatus s. l., are labelled above or below their respective branches. Circles on branches indicate Bayesian posterior probabilities&gt; 0.9." figureDoi="10.3897/zookeys.1180.109298.figure10" httpUri="https://binary.pensoft.net/fig/905327" pageId="0" pageNumber="27">10</figureCitation>
). The two species-delimitation methods were largely concordant for the other MOTUs, except for the
<taxonomicName id="3AC7DE39D3838F95199E2EDC82CF8C33" lsidName="B. fuscatus" pageId="0" pageNumber="27" rank="species" species="fuscatus">
<emphasis id="C0AE252F70EED5B2DBA2A9A67DC93A54" italics="true" pageId="0" pageNumber="27">B. fuscatus</emphasis>
</taxonomicName>
<emphasis id="9CEEB5FC22512AAAB6716423AD55A2C2" italics="true" pageId="0" pageNumber="27">sensu lato</emphasis>
(
<emphasis id="88314A3182E045353A188B080276FB35" italics="true" pageId="0" pageNumber="27">s.l.</emphasis>
) clade, which was split into five and eight MOTUs according to the ASAP and mPTP methods, respectively.
</paragraph>
<caption id="D030B8EC0F97CB6DC11F87BEDE607286" doi="10.3897/zookeys.1180.109298.figure10" httpUri="https://binary.pensoft.net/fig/905327" pageId="0" pageNumber="27" start="Figure 10" startId="F10">
<paragraph id="F3491A6C687251672AFABB28CBF6464D" pageId="0" pageNumber="27">
<emphasis id="A23793412ADCBFEE753A071C0C7B7A4D" bold="true" pageId="0" pageNumber="27">Figure 10.</emphasis>
Bayesian (BEAST) maximum clade credibility COI tree of the
<taxonomicName id="8F6168979F48BB30497CA8697F2D87D8" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis fuscatus" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="6664EEF3CF42B8ADB0B2A26B3C825E6F" italics="true" pageId="0" pageNumber="27">Baetis fuscatus</emphasis>
</taxonomicName>
group in the Western Palaearctic. Coloured vertical boxes indicate species-delimitation hypothesis (i.e., MOTUs) according to the mPTP and ASAP methods. For each ASAP-based MOTU, the corresponding species names (where available) and the country/region of origin are provided, with the newly described species specified in bold and the associated GBIF codes highlighted in pale pink. Tips labelled with GBIF codes indicate newly sequenced specimens; PT002, ES029, and ES030 codes designate sequences from the project FREDIE; the GMGMQ2692-18 code is from BOLD; the other codes correspond to sequences obtained from GenBank. The three lineages,
<taxonomicName id="13C75DABEF72A874B8E00950D3B1157D" authorityName="El Yaagoubi, Vuataz &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis rifensis" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="CA7EAC3B96AE882CD40C4978AFF6D449" italics="true" pageId="0" pageNumber="27">Baetis rifensis</emphasis>
</taxonomicName>
<emphasis id="BB49905C1CA8D354064E613680D0D92A" italics="true" pageId="0" pageNumber="27">sensu lato</emphasis>
(
<emphasis id="DF9612336E058758F6C8EC443BE6B399" italics="true" pageId="0" pageNumber="27">s.l.</emphasis>
),
<taxonomicName id="65E1F88EABF8EA8ED42A38B917BE5D83" authorityName="Eaton" authorityYear="1870" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis scambus" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="scambus">
<emphasis id="744DE86AE97CB4E0F0C25512A770D1D8" italics="true" pageId="0" pageNumber="27">Baetis scambus</emphasis>
</taxonomicName>
<emphasis id="9F3B4E3A054782C81314DFF686785814" italics="true" pageId="0" pageNumber="27">s.l.</emphasis>
, and
<taxonomicName id="5E16DBA7EB511EFC504D99B54DC841FB" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis fuscatus" order="Ephemeroptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="31BB9F7A9940EED90A5F2D2362D1A06E" italics="true" pageId="0" pageNumber="27">Baetis fuscatus</emphasis>
</taxonomicName>
<emphasis id="47D790DD59987361843CC5B4FB46C3DB" italics="true" pageId="0" pageNumber="27">s.l.</emphasis>
, are labelled above or below their respective branches. Circles on branches indicate Bayesian posterior probabilities&gt;0.9.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>