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<document id="AC4F86A88D29EB83F29E959353FF8CBD" ID-DOI="10.5252/cryptogamie-bryologie2024v45a4" ID-GBIF-Dataset="be13208d-9646-4a62-b0aa-de2668950ba3" ID-ISSN="1776-0992" ID-Zenodo-Dep="13378493" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1724747919433" checkinUser="plazi" docAuthor="Hugonnot, Vincent, Kučera, Jan, Osman, Imen Ben, Daoud-Bouattour, Amina &amp; Muller, Serge D." docDate="2024" docId="03F1F227FFB07E52EA456845FB382231" docLanguage="en" docName="bryologie.2024.45.4.pdf" docOrigin="Cryptogamie, Bryologie 45 (4)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/bryologie2024v45a4.pdf" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Pterygoneurum sampaianum Guim." docType="treatment" docVersion="4" lastPageNumber="46" masterDocId="FFC88A5FFFB87E59E93F6F39FF82251D" masterDocTitle="Pterygoneurum sampaianum (Guim.) Guim.: range extension to Africa, first mentions in France, confirmation of specific status and improved morphological circumscription" masterLastPageNumber="48" masterPageNumber="37" pageNumber="43" updateTime="1725378201199" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="06932A7C7DF2E2D747840570AD2E1873">Pterygoneurum sampaianum (Guim.) Guim.: range extension to Africa, first mentions in France, confirmation of specific status and improved morphological circumscription</mods:title>
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<mods:namePart id="4ECD2502324A827E3CBB529D82C7C4B8">Hugonnot, Vincent</mods:namePart>
<mods:affiliation id="47E9EBD4F9F7786F1566DA3227E4B974">Le Bourg, 43380 Blassac (France) vincent. hugonnot @ wanadoo. fr</mods:affiliation>
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<mods:namePart id="2256A9302403451D625CEB65B68F2DA8">Kučera, Jan</mods:namePart>
<mods:affiliation id="C699D2B5A896997A236D02BAA19961C7">Department of Botany, Faculty of Science, University of South Bohemia, Branišovská 1760, CZ- 370 05 České Budějovice (Czech Republic) kucera @ prf. jcu. cz</mods:affiliation>
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<mods:namePart id="82D1E219661BF214088B10A643B74052">Osman, Imen Ben</mods:namePart>
<mods:affiliation id="A6781A94B53924615A325BC323B6CD70">Département de Biologie, Faculté des Sciences de Tunis, Université Tunis El-Manar, 2092 Tunis (Tunisia) &amp; LR 18 ES 13 BiCADE, Faculté des Lettres, des Arts et des Humanités de Manouba, Université de la Manouba, 2010 Manouba (Tunisia)</mods:affiliation>
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<mods:namePart id="9FFE11AC1360BB4076DFE06B153594AA">Daoud-Bouattour, Amina</mods:namePart>
<mods:affiliation id="786F2BB01CAEC91EB180AD7E82B8C9A8">Département de Biologie, Faculté des Sciences de Tunis, Université Tunis El-Manar, 2092 Tunis (Tunisia) and LR 18 ES 13 BiCADE, Faculté des Lettres, des Arts et des Humanités de Manouba, Université de la Manouba, 2010 Manouba (Tunisia) imanbenosmen @ gmail. com daoudamina 200 @ yahoo. fr</mods:affiliation>
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<mods:namePart id="F333EC903ACF35B6862527D33410E12C">Muller, Serge D.</mods:namePart>
<mods:affiliation id="04EC9DB39E13D8EA75FDEE715CFEDDA7">Institut des Sciences de l’Évolution (ISEM), Université de Montpellier, CNRS, IRD, EPHE, 34095 Montpellier cedex 05 (France) serge. muller @ umontpellier. fr</mods:affiliation>
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(Guim.) Guim.
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• Tunisian Dorsal, 
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• 
<materialsCitation id="3B30496CFFB17E50E9BC6E4FFD0424B4" ID-GBIF-Occurrence="4928362312" collectingDate="2021-03-04" collectorName="Hugonnot." country="France" county="Barmettaz" elevation="1300" latitude="45.220066" location="Savoie" longLatPrecision="1" longitude="6.7471724" municipality="Maurienne valley" pageId="9" pageNumber="44" specimenCount="1">
<location id="8E8715EAFFB17E50E9BC6E4FFF452493" LSID="urn:lsid:plazi:treatment:03F1F227FFB07E52EA456845FB382231:8E8715EAFFB17E50E9BC6E4FFF452493" box="[131,199,374,398]" country="France" county="Barmettaz" latitude="45.220066" longLatPrecision="1" longitude="6.7471724" municipality="Maurienne valley" name="Savoie" pageId="9" pageNumber="44">Savoie</location>
, 
<location id="8E8715EAFFB17E50E9ED6E4FFEA22493" LSID="urn:lsid:plazi:treatment:03F1F227FFB07E52EA456845FB382231:8E8715EAFFB17E50E9ED6E4FFEA22493" box="[210,288,374,398]" country="France" county="Barmettaz" latitude="45.220066" longLatPrecision="1" longitude="6.7471724" municipality="Maurienne valley" name="Aussois" pageId="9" pageNumber="44">Aussois</location>
, 
<collectingMunicipality id="6B83D94BFFB17E50E8136E4FFE5D2493" box="[300,479,374,398]" pageId="9" pageNumber="44">Maurienne valley</collectingMunicipality>
, 
<collectingCounty id="62863BBDFFB17E50E8D46E4FFDD72490" box="[491,597,374,397]" pageId="9" pageNumber="44">Barmettaz</collectingCounty>
; 
<geoCoordinate id="EE6C25F6FFB17E50EB5A6E4CFC862490" box="[613,772,373,397]" degrees="45" direction="north" minutes="13" orientation="latitude" pageId="9" pageNumber="44" precision="1" seconds="12.23" value="45.220066">45°13’12.23”N</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50E9BB6EA9FE9524B5" box="[132,279,400,424]" degrees="06" direction="east" minutes="44" orientation="longitude" pageId="9" pageNumber="44" precision="1" seconds="49.82" value="6.7471724">06°44’49.82”E</geoCoordinate>
; 
<quantity id="4CA0EED4FFB17E50E81C6EA9FEF024B5" box="[291,370,400,424]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="44" unit="m" value="1300.0">
<elevation id="0075A402FFB17E50E81C6EA9FEF024B5" box="[291,370,400,424]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="44" unit="m" value="1300.0">1300 m</elevation>
</quantity>
; 
<date id="FFE665F1FFB17E50E8426EA9FE6624B5" box="[381,484,400,424]" pageId="9" pageNumber="44" value="2021-03-04">
<collectingDate id="EFA29C19FFB17E50E8426EA9FE6624B5" box="[381,484,400,424]" pageId="9" pageNumber="44" value="2021-03-04">4.III.2021</collectingDate>
</date>
; leg. 
<collectorName id="26AD26E7FFB17E50EB286EA8FD0424B4" box="[535,646,401,425]" pageId="9" pageNumber="44">Hugonnot.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BE74331FFB17E50E9BB6E95FE6A2731" blockId="9.[131,776,215,557]" pageId="9" pageNumber="44">
<materialsCitation id="3B30496CFFB17E50E9BB6E95FF4924EA" ID-GBIF-Occurrence="4928362302" collectingDate="2020-02-12" country="Spain" latitude="42.07903" location="Torroella de Montgri" longLatPrecision="1" longitude="3.19476" pageId="9" pageNumber="44" specimenCount="1">
<emphasis id="B92C9F23FFB17E50E9BB6E95FF3824D9" bold="true" box="[132,186,428,452]" pageId="9" pageNumber="44">
<collectingCountry id="F34F03A1FFB17E50E9BB6E95FF3824D9" box="[132,186,428,452]" name="Spain" pageId="9" pageNumber="44">Spain</collectingCountry>
</emphasis>
• 
<location id="8E8715EAFFB17E50E9F56E93FE0E24DE" LSID="urn:lsid:plazi:treatment:03F1F227FFB07E52EA456845FB382231:8E8715EAFFB17E50E9F56E93FE0E24DE" box="[202,396,426,451]" country="Spain" latitude="42.07903" longLatPrecision="1" longitude="3.19476" name="Torroella de Montgri" pageId="9" pageNumber="44">Torroella de Montgrí</location>
, 
<location id="8E8715EAFFB17E50E8AB6E93FE6824DF" LSID="urn:lsid:plazi:treatment:03F1F227FFB07E52EA456845FB382231:8E8715EAFFB17E50E8AB6E93FE6824DF" box="[404,490,426,450]" country="Spain" latitude="42.07903" longLatPrecision="1" longitude="3.19476" name="L'Estartit" pageId="9" pageNumber="44">L’Estartit</location>
, 
<location id="8E8715EAFFB17E50E8CE6E92FDDF24DF" LSID="urn:lsid:plazi:treatment:03F1F227FFB07E52EA456845FB382231:8E8715EAFFB17E50E8CE6E92FDDF24DF" box="[497,605,426,451]" country="Spain" latitude="42.07903" longLatPrecision="1" longitude="3.19476" name="Cap Castell" pageId="9" pageNumber="44">Cap Castell</location>
; 
<geoCoordinate id="EE6C25F6FFB17E50EB5C6E93FC8324DF" box="[611,769,426,450]" degrees="42" direction="north" minutes="4" orientation="latitude" pageId="9" pageNumber="44" precision="1" seconds="44.5080" value="42.07903">42°4’44.5080”N</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50E9BB6EFCFEB224C0" box="[132,304,453,477]" degrees="03" direction="east" minutes="11" orientation="longitude" pageId="9" pageNumber="44" precision="1" seconds="41.1360" value="3.19476">03°11’41.1360”E</geoCoordinate>
; 
<date id="FFE665F1FFB17E50E8016EFCFE2924C0" box="[318,427,453,477]" pageId="9" pageNumber="44" value="2020-02-12">
<collectingDate id="EFA29C19FFB17E50E8016EFCFE2924C0" box="[318,427,453,477]" pageId="9" pageNumber="44" value="2020-02-12">12.II.2020</collectingDate>
</date>
; Vincent Hugonnot’s personal herbarium
</materialsCitation>
• 
<materialsCitation id="3B30496CFFB17E50E9DC6EE6FE672731" ID-GBIF-Occurrence="4928362321" collectingDate="2015-04-28" collectorName="J. Kucera" country="Spain" elevation="800" latitude="36.980305" location="Barranco de Cacin" longLatPrecision="1" longitude="-2.812972" municipality="Sierra de Gador" pageId="9" pageNumber="44" specimenCode="CBFS 17331, 17332" specimenCount="1" stateProvince="Andalucia">
<collectingRegion id="499C8DD3FFB17E50E9DC6EE6FECA24EA" box="[227,328,479,503]" country="Spain" name="Andalucia" pageId="9" pageNumber="44">Andalucía</collectingRegion>
, Almería, Fondón, 
<collectingMunicipality id="6B83D94BFFB17E50EB376EE6FD2624EA" box="[520,676,479,503]" pageId="9" pageNumber="44">Sierra de Gádor</collectingMunicipality>
, 
<location id="8E8715EAFFB17E50EB906ED9FF63270F" LSID="urn:lsid:plazi:treatment:03F1F227FFB07E52EA456845FB382231:8E8715EAFFB17E50EB906ED9FF63270F" country="Spain" latitude="36.980305" longLatPrecision="1" longitude="-2.812972" municipality="Sierra de Gador" name="Barranco de Cacin" pageId="9" pageNumber="44" stateProvince="Andalucia">Barranco de Cacín</location>
; 
<geoCoordinate id="EE6C25F6FFB17E50E9D06EC3FEFC270F" box="[239,382,506,530]" degrees="36" direction="north" minutes="58" orientation="latitude" pageId="9" pageNumber="44" precision="1" seconds="49.1" value="36.980305">36°58’49.1”N</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50E8B56EC3FD9A270F" box="[394,536,506,530]" degrees="02" direction="west" minutes="48" orientation="longitude" pageId="9" pageNumber="44" precision="1" seconds="46.7" value="-2.812972">02°48’46.7”W</geoCoordinate>
; 
<emphasis id="B92C9F23FFB17E50EB186EC2FDB2270F" box="[551,560,507,530]" italics="true" pageId="9" pageNumber="44">c</emphasis>
. 
<quantity id="4CA0EED4FFB17E50EB016EC3FD01270C" box="[574,643,506,530]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="44" unit="m" value="800.0">
<elevation id="0075A402FFB17E50EB016EC3FD01270C" box="[574,643,506,530]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="44" unit="m" value="800.0">800 m</elevation>
</quantity>
; 
<date id="FFE665F1FFB17E50EBB06EC3FC80270F" box="[655,770,506,530]" pageId="9" pageNumber="44" value="2015-04-28">
<collectingDate id="EFA29C19FFB17E50EBB06EC3FC80270F" box="[655,770,506,530]" pageId="9" pageNumber="44" value="2015-04-28">28.IV.2015</collectingDate>
</date>
; leg. 
<collectorName id="26AD26E7FFB17E50E9916D2CFE892730" box="[174,267,533,557]" pageId="9" pageNumber="44">J. Kučera</collectorName>
; 
<specimenCode id="DBFEEB4AFFB17E50E82A6D2DFE1A2731" box="[277,408,532,556]" collectionCode="CBFS" country="Czech Republic" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15554" lsid="urn:lsid:biocol.org:col:15554" name="University of South Bohemia" pageId="9" pageNumber="44" type="Herbarium">CBFS 17331</specimenCode>
, 
<specimenCode id="DBFEEB4AFFB17E50E89D6D2DFE672731" box="[418,485,532,556]" collectionCode="CBFS" country="Czech Republic" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15554" lsid="urn:lsid:biocol.org:col:15554" name="University of South Bohemia" pageId="9" pageNumber="44" type="Herbarium">17332</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34210BAFFB17E50E9BB6D6FFACC2193" pageId="9" pageNumber="44" type="description">
<paragraph id="8BE74331FFB17E50E9BB6D6FFE072773" blockId="9.[132,389,598,624]" box="[132,389,598,624]" pageId="9" pageNumber="44">
<heading id="D0AFF45DFFB17E50E9BB6D6FFE072773" allCaps="true" box="[132,389,598,624]" fontSize="9" level="2" pageId="9" pageNumber="44" reason="2">
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<smallCapsWord id="8D01D5EDFFB17E50E9C66D62FE072773" baselines="617" box="[249,389,603,622]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="description" pageId="9" pageNumber="44">DESCRIPTION</smallCapsWord>
</heading>
</paragraph>
<paragraph id="8BE74331FFB17E50E9BB6D4FFE37260D" blockId="9.[130,777,630,2028]" pageId="9" pageNumber="44">
Previous morphological descriptions did not fully account for the morphological variability observed in the studied specimens whose identification was confirmed using molecular data. We therefore provide an emended morphological description of 
<taxonomicName id="4C5838B2FFB17E50E9BB6DCFFE2C2612" box="[132,430,758,783]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="9" pageNumber="44" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB17E50E9BB6DCFFE2C2612" box="[132,430,758,783]" italics="true" pageId="9" pageNumber="44">Pterygoneurum sampaianum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE74331FFB17E50E9A46C2CFC39276D" blockId="9.[130,777,630,2028]" lastBlockId="9.[810,1457,215,1167]" pageId="9" pageNumber="44">
Plants gregarious, in loose colonies characteristically consisting of very small sterile individuals and much larger fertile individuals, shortly bulbiform, whitish green to pale green, often with discolored tips, 1.25-1.8 (3.5) mm high. Axillary hairs 
<quantity id="4CA0EED4FFB17E50E9816CACFE8326B2" box="[190,257,917,943]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="9" pageNumber="44" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3-4 in</quantity>
axils of upper leaves, 9-12 ×100-130 µm, made of 1-2 basal cells short and 3-5 feebly colored and longer cells; terminal cell 9-10 ×30-35 µm. Subterranean part of stem thick and brown, 0.19-0.24× 
<quantity id="4CA0EED4FFB17E50E8C16CCDFD1D2112" box="[510,671,1012,1039]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="9" pageNumber="44" unit="mm" value="0.5" valueMax="0.55" valueMin="0.45">0.45-0.55 mm</quantity>
, of thick-walled cells, with or without 1-2 non-chlorophyllous scales, proximally attached to substrate by 5-10 large brown rhizoids, 36-50 µm wide near base, often more than 
<quantity id="4CA0EED4FFB17E50EB6F6B6DFD122172" box="[592,656,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="44" unit="mm" value="6.0">6 mm</quantity>
long, ramified, strongly decreasing in size towards the apex (to 8 µm wide) and becoming hyaline, smooth but coated with fine soil material. Leafy stem aerial, unbranched or occasionally branched (often the smaller branch sterile, reduced to few leaves), with cortex undifferentiated (externally with smaller cells with barely thickened walls), central strand present but weak, of several cells. Proximal above-ground leaves reduced and hardly chlorophyllose, equally wide or wider than long, trapezoidal, suborbicular to rectangular, deeply concave, with none or very short acumen, awnless; basal marginal cells strongly differentiated into 2-4 cell-wide hyaline border, of long rectangular cells (5-7 ×40-50 µm), ascending along the margin; median and apical cells often very thick-walled. Distal leaves erect, deeply concave to convolute, awnless or with short straight hyaline smooth awn (70-100 µm long), individual leaves on some shoots occasionally bear a longer, up to 400 µm long, at times even flexuous hair, or many leaves piliferous on the same shoot; leaves wider than long, trapezoidal, 0.7-0.9× 
<quantity id="4CA0EED4FFB17E50E84E698BFE7B23D0" box="[369,505,1714,1741]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="9" pageNumber="44" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7-0.8 mm</quantity>
, rarely longer than wide 0.4-0.5× 
<quantity id="4CA0EED4FFB17E50E9D669EBFEF223F1" box="[233,368,1746,1772]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="9" pageNumber="44" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8-0.9 mm</quantity>
, widely rounded to truncate at apex, with plane margins; basal marginal cells forming a more or less distinct hyaline border, 3-6 cells wide, ascending to apical 1/3 of rectangular cells 10-14 × 40-70 µm, border totally missing at times; basal cells short-rectangular to quadrate, 15-22× 18-45 µm; median cells quadrate to rectangular, 8-15×8-20 µm; apical cells rounded quadrate to rounded rectangular, 7-13×8-18 µm, with thick walls, often hyaline; margin dentate from mid-leaf to apex, teeth formed by the protruding cells, often geminate. Costa 45-66 µm wide at base, adaxially expanded into 2-3 large lamellae to 15 cells high, running from base to apex, crenulated at margin by protruding conical cells (apex mamilla like, rarely with two mamilla-like ornamentations); terminal cells of lamellae 12-14×15-18 µm; surface of lamellae occasionally with short outgrowths of conical cells or small laminal outgrowths; cells of lamellae 14-20 ×18-30 µm; surface cells of costa thick-walled both abaxially and adaxially; abaxial walls of lamina cells more thickened than adaxial ones in transverse section, costa with 2-3 poorly differentiated guide cells, dorsal stereid layer not developed. Autoicous, male buds at stem base on ground level.
</paragraph>
<paragraph id="8BE74331FFB17E50EA7C6D4FFACC2193" blockId="9.[810,1457,215,1167]" pageId="9" pageNumber="44">
Sporophyte with red-brown urn. Seta 0.076 ×0.09× 1.4- 2.5 (3) mm, dextrorse; epidermal cells 8-13× 18-57 µm, with slightly thickened walls. Seta in transverse section with one layer of epidermal cell with outer tangentially thickened walls, central strand present. Urn shortly cylindrical, rarely cupulate, with faint longitudinal plicae when dry, wide-mouthed, 320-600 × 740-890 µm; exothecial cells irregularly rectangular, 18-32× 40-87 µm, with rather thin walls. Epidermal cell transverse section with tangentially not or very weakly thickened outer walls. Stomata 1-2, phaneroporous, at base of urn. Annulus poorly differentiated, of 1-2 rows of rectangular, barely vesiculose cells. Mouth with no trace of peristome (either at rim or in the operculum). Operculum with short and blunt beak, 
<quantity id="4CA0EED4FFB17E50EA546B2DFB8A2132" box="[875,1032,1044,1071]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="9" pageNumber="44" unit="mm" value="0.525" valueMax="0.65" valueMin="0.4">0.40-0.65 mm</quantity>
long. Spores spherical, 35-58 µm across diameter, often ellipsoid or potato-shaped. Calyptra narrowly cylindrical at first, at maturity cucullate, rostrate, subobtuse at apex, covering urn at length (
<quantity id="4CA0EED4FFB17E50EDBB6B4DFA852192" box="[1156,1287,1140,1167]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="9" pageNumber="44" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4-0.7 mm</quantity>
long).
</paragraph>
</subSubSection>
<subSubSection id="C34210BAFFB17E52EA126B8DFAA12692" lastPageId="11" lastPageNumber="46" pageId="9" pageNumber="44" type="biology_ecology">
<paragraph id="8BE74331FFB17E50EA126B8DFC0F21D1" blockId="9.[810,1457,1204,2028]" box="[813,909,1204,1230]" pageId="9" pageNumber="44">
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</paragraph>
<paragraph id="8BE74331FFB17E52EA126BEDFD07278D" blockId="9.[810,1457,1204,2028]" lastBlockId="11.[130,777,215,2028]" lastPageId="11" lastPageNumber="46" pageId="9" pageNumber="44">
Our observations allow us to identify the ecological requirements of 
<taxonomicName id="4C5838B2FFB17E50EAB26BCDFB372010" box="[909,1205,1268,1293]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="9" pageNumber="44" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB17E50EAB26BCDFB372010" box="[909,1205,1268,1293]" italics="true" pageId="9" pageNumber="44">Pterygoneurum sampaianum</emphasis>
</taxonomicName>
. The species is typically found in arid or semiarid climates. In the Maurienne valley of the French Alps, the locality is recognized as being among the driest in 
<collectingCountry id="F34F03A1FFB17E50EAB66A6DFC562073" box="[905,980,1364,1390]" name="France" pageId="9" pageNumber="44">France</collectingCountry>
, with an average annual rainfall of 
<quantity id="4CA0EED4FFB17E50EC776A6AFA272070" box="[1352,1445,1363,1390]" metricMagnitude="-1" metricUnit="m" metricValue="5.18" pageId="9" pageNumber="44" unit="mm" value="518.0">518 mm</quantity>
/ year, with the vegetation referred to as 
<emphasis id="B92C9F23FFB17E50EDF26A4AFA2D2090" box="[1229,1455,1395,1421]" italics="true" pageId="9" pageNumber="44">Festucetalia valesiacae</emphasis>
Br.-Bl.&amp;Tüxen ex Br.-Bl. 1949. The dominants include 
<taxonomicName id="4C5838B2FFB17E50EC596AADFBEB20D0" authority="Schleich. ex Gaudin" authorityName="Gaudin" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="species" species="valesiaca">
<emphasis id="B92C9F23FFB17E50EC596AADFC0520D0" italics="true" pageId="9" pageNumber="44">Festuca valesiaca</emphasis>
Schleich. ex Gaudin
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB17E50ED4B6A8AFA8B20D0" box="[1140,1289,1459,1485]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="species" species="capillata">
<emphasis id="B92C9F23FFB17E50ED4B6A8AFA8B20D0" box="[1140,1289,1459,1485]" italics="true" pageId="9" pageNumber="44">Stipa capillata</emphasis>
</taxonomicName>
L., 
<taxonomicName id="4C5838B2FFB17E50EC0B6A8AFCF520F0" authority="Borbas" authorityName="Borbas" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="species" species="eriocaulis">
<emphasis id="B92C9F23FFB17E50EC0B6A8AFA2D20D0" box="[1332,1455,1459,1485]" italics="true" pageId="9" pageNumber="44">S. eriocaulis</emphasis>
Borbás
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB17E50EA406AEAFA2E20F0" authority="(Rouy) P. Fourn." authorityName="P. Fourn." baseAuthorityName="Rouy" box="[895,1452,1491,1517]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="subSpecies" species="seguieriana" subSpecies="loiseleurii">
<emphasis id="B92C9F23FFB17E50EA406AEAFBD820F0" box="[895,1114,1491,1517]" italics="true" pageId="9" pageNumber="44">Euphorbia seguieriana</emphasis>
subsp. 
<emphasis id="B92C9F23FFB17E50ED9E6AEAFB7C20F0" box="[1185,1278,1491,1517]" italics="true" pageId="9" pageNumber="44">loiseleurii</emphasis>
(Rouy) P.Fourn.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB17E50EA156ACAFB1D2310" authority="(Honck.) Gaudin" authorityName="Gaudin" baseAuthorityName="Honck." box="[810,1183,1523,1549]" class="Liliopsida" family="Poaceae" genus="Koeleria" kingdom="Plantae" order="Poales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="species" species="vallesiana">
<emphasis id="B92C9F23FFB17E50EA156ACAFC612310" box="[810,995,1523,1549]" italics="true" pageId="9" pageNumber="44">Koeleria vallesiana</emphasis>
(Honck.) Gaudin
</taxonomicName>
and 
<taxonomicName id="4C5838B2FFB17E50EDF26ACAFA322310" authority="L. In" authorityName="L. In" box="[1229,1456,1523,1549]" class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="species" species="pusillum">
<emphasis id="B92C9F23FFB17E50EDF26ACAFAF52310" box="[1229,1399,1523,1549]" italics="true" pageId="9" pageNumber="44">Galium pusillum</emphasis>
L. In
</taxonomicName>
the Isle-Crémieu, North Isère, 
<collectingCountry id="F34F03A1FFB17E50ED56692AFB322330" box="[1129,1200,1555,1581]" name="France" pageId="9" pageNumber="44">France</collectingCountry>
(Vénérieu, Simiaud and La Chanas; derelict marl quarry; 
<geoCoordinate id="EE6C25F6FFB17E50EDAF690BFA952350" box="[1168,1303,1586,1613]" degrees="45" direction="north" minutes="39" orientation="latitude" pageId="9" pageNumber="44" precision="15" seconds="43" value="45.661945">45°39’43”N</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50EC1D690BFA2A2350" box="[1314,1448,1586,1613]" degrees="05" direction="west" minutes="17" orientation="longitude" pageId="9" pageNumber="44" precision="15" seconds="17" value="-5.2880554">05°17’17”W</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50EA12696BFC352370" box="[813,951,1618,1645]" degrees="45" direction="north" minutes="39" orientation="latitude" pageId="9" pageNumber="44" precision="15" seconds="36" value="45.66">45°39’36”N</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50EAFA696BFBCD2370" box="[965,1103,1618,1645]" degrees="05" direction="west" minutes="17" orientation="longitude" pageId="9" pageNumber="44" precision="15" seconds="17" value="-5.2880554">05°17’17”W</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50ED60696BFB6B2370" box="[1119,1257,1618,1645]" degrees="45" direction="north" minutes="39" orientation="latitude" pageId="9" pageNumber="44" precision="15" seconds="18" value="45.655003">45°39’18”N</geoCoordinate>
, 
<geoCoordinate id="EE6C25F6FFB17E50EDC8696BFAF62370" box="[1271,1396,1618,1645]" degrees="5" direction="west" minutes="16" orientation="longitude" pageId="9" pageNumber="44" precision="15" seconds="33" value="-5.2758336">5°16’33”W</geoCoordinate>
; 238 and 
<quantity id="4CA0EED4FFB17E50EA62694BFC262390" box="[861,932,1650,1677]" metricMagnitude="2" metricUnit="m" metricValue="2.18" pageId="9" pageNumber="44" unit="m" value="218.0">218 m</quantity>
a.s.l.; 
<date id="FFE665F1FFB17E50EAD6694BFB042391" box="[1001,1158,1650,1676]" pageId="9" pageNumber="44" value="2023-03-03">3 March 2023</date>
; obs. S. Geoffroy, A. Labroche, Y. Longeot and M. Maglio) a calcareous plateau with an average annual rainfall much higher, of 
<quantity id="4CA0EED4FFB17E50EC2C698BFA2A23D1" box="[1299,1448,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="9.6" metricValueMax="9.7" metricValueMin="9.5" pageId="9" pageNumber="44" unit="mm" value="960.0" valueMax="970.0" valueMin="950.0">950-970 mm</quantity>
/ year, the vegetation where 
<taxonomicName id="4C5838B2FFB17E50ED7569EAFB6A23F1" box="[1098,1256,1747,1772]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="9" pageNumber="44" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB17E50ED7569EAFB6A23F1" box="[1098,1256,1747,1772]" italics="true" pageId="9" pageNumber="44">P. sampaianum</emphasis>
</taxonomicName>
have been spotted is a mosaic of mesoxerophilous grasslands of 
<emphasis id="B92C9F23FFB17E50EC1E69CBFBD62236" italics="true" pageId="9" pageNumber="44">Tetragonolobo maritimi -Bromenion erecti</emphasis>
<collectionCode id="ED49DBF4FFB17E50ED61682BFBEB2231" box="[1118,1129,1810,1836]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="9" pageNumber="44" type="Herbarium">J</collectionCode>
.-M.Royer 2006 and 
<emphasis id="B92C9F23FFB17E50EC5E682BFBBE2256" italics="true" pageId="9" pageNumber="44">Teucrio montani -Bromenion erecti</emphasis>
<collectionCode id="ED49DBF4FFB17E50ED7E680BFBCE2251" box="[1089,1100,1842,1868]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="9" pageNumber="44" type="Herbarium">J</collectionCode>
.-M.Royer 2006 and thermophilous semi-wet meadows with 
<taxonomicName id="4C5838B2FFB17E50ED546868FA9E2276" box="[1131,1308,1873,1899]" class="Liliopsida" family="Poaceae" genus="Molinia" kingdom="Plantae" order="Poales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="species" species="caerulea">
<emphasis id="B92C9F23FFB17E50ED546868FA9E2276" box="[1131,1308,1873,1899]" italics="true" pageId="9" pageNumber="44">Molinia caerulea</emphasis>
</taxonomicName>
(L.) Moench (
<emphasis id="B92C9F23FFB17E50EA096848FAAC2296" box="[822,1326,1905,1932]" italics="true" pageId="9" pageNumber="44">Deschampsio mediae -Molinienion arundinaceae</emphasis>
B.Foucault 2008) developed on outcrops of marl-limestone soil with temporary hydromorphy. In the eastern Pyrenees, the average annual rainfall is approximately 
<quantity id="4CA0EED4FFB17E50ED4568E8FB5122F6" box="[1146,1235,2001,2028]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="44" unit="mm" value="550.0">550 mm</quantity>
/year. The vegetation comprises degraded grasslands and garrigues with 
<taxonomicName id="4C5838B2FFB37E52EB9F6FEEFE47240F" authority="(Pers.) P. Beauv." authorityName="P. Beauv." baseAuthorityName="Pers." class="Liliopsida" family="Poaceae" genus="Brachypodium" kingdom="Plantae" order="Poales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="retusum">
<emphasis id="B92C9F23FFB37E52EB9F6FEEFE97240C" italics="true" pageId="11" pageNumber="46">Brachypodium retusum</emphasis>
(Pers.) P.Beauv.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E8EE6FC1FD30240C" box="[465,690,247,273]" class="Magnoliopsida" family="Lamiaceae" genus="Rosmarinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="B92C9F23FFB37E52E8EE6FC1FD30240C" box="[465,690,247,273]" italics="true" pageId="11" pageNumber="46">Rosmarinus officinalis</emphasis>
</taxonomicName>
L., and is referred to the alliances 
<emphasis id="B92C9F23FFB37E52E8A26E2EFF3E244C" italics="true" pageId="11" pageNumber="46">Phlomido lychnitidis -Brachypodion retusi</emphasis>
(Rivas-Martínez 1978) Mateo 1983 and 
<taxonomicName id="4C5838B2FFB37E52EBBB6E01FE8F246C" authority="Br." authorityName="Br." class="Bryopsida" family="Pottiaceae" genus="Rosmarinion" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="officinalis">
<emphasis id="B92C9F23FFB37E52EBBB6E01FF64246C" italics="true" pageId="11" pageNumber="46">Rosmarinion officinalis</emphasis>
Br.
</taxonomicName>
-Bl. ex Molinier. In 
<collectingCountry id="F34F03A1FFB37E52E8E66E6EFDA9246C" box="[473,555,343,369]" name="Tunisia" pageId="11" pageNumber="46">Tunisia</collectingCountry>
, 
<taxonomicName id="4C5838B2FFB37E52EB0D6E61FF60248D" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52EB0D6E61FF60248D" italics="true" pageId="11" pageNumber="46">Pterygoneurum sampaianum</emphasis>
</taxonomicName>
was collected from the Tunisian Dorsal, which has an average annual rainfall of 
<emphasis id="B92C9F23FFB37E52E8906EAEFE3A24AD" box="[431,440,407,432]" italics="true" pageId="11" pageNumber="46">c</emphasis>
. 
<quantity id="4CA0EED4FFB37E52E8F86EAEFDA324AC" box="[455,545,407,433]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="46" unit="mm" value="400.0">400 mm</quantity>
/year. The plants were growing in low matorral vegetation composed primarily of solitary 
<taxonomicName id="4C5838B2FFB37E52E9E56EEEFE3E24ED" authority="Mill." authorityName="Mill." box="[218,444,470,496]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="halepensis">
<emphasis id="B92C9F23FFB37E52E9E56EEEFE0224ED" box="[218,384,470,496]" italics="true" pageId="11" pageNumber="46">Pinus halepensis</emphasis>
Mill.
</taxonomicName>
and 
<taxonomicName id="4C5838B2FFB37E52E8CE6EEEFDED24ED" box="[497,623,470,497]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B92C9F23FFB37E52E8CE6EEEFDED24ED" box="[497,623,470,497]" italics="true" pageId="11" pageNumber="46">Quercus ilex</emphasis>
</taxonomicName>
L. 
<taxonomicName id="4C5838B2FFB37E52EBAB6EEFFED2270D" authority="(Desf.) Samp." authorityName="Samp." baseAuthorityName="Desf." pageId="11" pageNumber="46" rank="subSpecies" subSpecies="ballota">
subsp. 
<emphasis id="B92C9F23FFB37E52EBE26EEFFF29270D" italics="true" pageId="11" pageNumber="46">ballota</emphasis>
(Desf.) Samp.
</taxonomicName>
, with 
<taxonomicName id="4C5838B2FFB37E52E8A76ECEFDE7270D" authority="Dunal" authorityName="Dunal" box="[408,613,502,529]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="clusii">
<emphasis id="B92C9F23FFB37E52E8A76ECEFD94270D" box="[408,534,502,529]" italics="true" pageId="11" pageNumber="46">Cistus clusii</emphasis>
Dunal
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EB4B6ECEFD62270D" box="[628,736,503,529]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="creticus">
<emphasis id="B92C9F23FFB37E52EB4B6ECEFD62270D" box="[628,736,503,529]" italics="true" pageId="11" pageNumber="46">C. creticus</emphasis>
</taxonomicName>
L., 
<taxonomicName id="4C5838B2FFB37E52E9B96D2FFEC0272D" box="[134,322,534,560]" class="Magnoliopsida" family="Rhamnaceae" genus="Rhamnus" kingdom="Plantae" order="Rosales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="lycioides">
<emphasis id="B92C9F23FFB37E52E9B96D2FFEC0272D" box="[134,322,534,560]" italics="true" pageId="11" pageNumber="46">Rhamnus lycioides</emphasis>
</taxonomicName>
L. and 
<taxonomicName id="4C5838B2FFB37E52E8AE6D2EFDF3272D" box="[401,625,534,560]" class="Magnoliopsida" family="Lamiaceae" genus="Rosmarinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="46" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="B92C9F23FFB37E52E8AE6D2EFDF3272D" box="[401,625,534,560]" italics="true" pageId="11" pageNumber="46">Rosmarinus officinalis</emphasis>
</taxonomicName>
, belonging to the order 
<emphasis id="B92C9F23FFB37E52E9D66D0EFD9D274D" box="[233,543,566,593]" italics="true" pageId="11" pageNumber="46">
<taxonomicName id="4C5838B2FFB37E52E9D66D0EFEBA274D" box="[233,312,567,592]" order="Pistacio" pageId="11" pageNumber="46" rank="order">Pistacio</taxonomicName>
-Rhamnetalia alaterni
</emphasis>
Rivas-Martinez 1975 and the recently described alliance 
<emphasis id="B92C9F23FFB37E52EB3E6D6EFD2C276D" box="[513,686,598,624]" italics="true" pageId="11" pageNumber="46">Loto dorycnium -</emphasis>
<taxonomicName id="4C5838B2FFB37E52EB916D6EFE04278D" authority="Djebaili 2017" authorityName="Djebaili" authorityYear="2017" class="Bryopsida" family="Pottiaceae" genus="Pinetum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="halepensis">
<emphasis id="B92C9F23FFB37E52EB916D6EFF6B278D" italics="true" pageId="11" pageNumber="46">Pinetum halepensis</emphasis>
Djebaïli 2017
</taxonomicName>
(
<bibRefCitation id="EFC93EC0FFB37E52E8A96D4FFDF4278D" author="MEDDOUR R. &amp; MEDDOUR-SAHAR O. &amp; ZERAIA L. &amp; MUCINA L." box="[406,630,630,656]" pageId="11" pageNumber="46" pagination="127 - 163" refId="ref9125" refString="MEDDOUR R., MEDDOUR-SAHAR O., ZERAIA L. &amp; MUCINA L. 2017. - Syntaxonomic synopsis of the forest and tall scrub vegetation of Northern Algeria. Lazaroa 38 (2): 127 - 163. http: // dx. doi. org / 10.5209 / LAZA. 53272" type="journal article" year="2017">
Meddour 
<emphasis id="B92C9F23FFB37E52EB3C6D4EFDB4278D" box="[515,566,630,656]" italics="true" pageId="11" pageNumber="46">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<caption id="DF2713B9FFB27E53E82168EEFA9522F4" ID-DOI="http://doi.org/10.5281/zenodo.13378503" ID-Zenodo-Dep="13378503" box="[286,1303,2007,2025]" httpUri="https://zenodo.org/record/13378503/files/figure.png" pageId="10" pageNumber="45" startId="10.[286,297,2007,2025]" targetBox="[173,1416,215,1878]" targetPageId="10" targetType="figure">
<paragraph id="8BE74331FFB27E53E82168EEFA9522F4" blockId="10.[286,1303,2007,2025]" box="[286,1303,2007,2025]" pageId="10" pageNumber="45">
<smallCapsWord id="8D01D5EDFFB27E53E82168EEFEB522F5" baselines="2020,2021" box="[286,311,2007,2025]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="45">FIG</smallCapsWord>
. 4. — Habitat of 
<taxonomicName id="4C5838B2FFB27E53E8FA68EEFCAC22F4" authority="(Guim.) Guim." authorityName="Guim." baseAuthorityName="Guim." box="[453,814,2007,2025]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="10" pageNumber="45" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB27E53E8FA68EEFD3622F4" box="[453,692,2007,2025]" italics="true" pageId="10" pageNumber="45">Pterygoneurum sampaianum</emphasis>
(Guim.) Guim.
</taxonomicName>
(11 March 2020, Tunisia, Siliana Governorate, Sidi Ameur).
</paragraph>
</caption>
<paragraph id="8BE74331FFB37E52E9A46DAFFDCC20F0" blockId="11.[130,777,215,2028]" pageId="11" pageNumber="46">
In the newly-discovered French populations, the species grows with rich assemblages of pioneer, mainly ruderal bryophytes, often belonging to the family 
<taxonomicName id="4C5838B2FFB37E52E8FA6DECFDB727F2" box="[453,565,725,751]" class="Bryopsida" family="Pottiaceae" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="family">Pottiaceae</taxonomicName>
. In Savoy, the most frequently associated species are 
<taxonomicName id="4C5838B2FFB37E52E8F16DCCFEAB2632" authority="(W. D. J. Koch ex Schultz) Kindb." authorityName="Kindb." baseAuthorityName="Schultz" class="Bryopsida" family="Pottiaceae" genus="Aloina" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="aloides">
<emphasis id="B92C9F23FFB37E52E8F16DCCFDDB2612" box="[462,601,757,783]" italics="true" pageId="11" pageNumber="46">Aloina aloides</emphasis>
(W.D.J.Koch ex Schultz) Kindb.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E80E6C2FFDC72632" authority="Hedw." authorityName="Hedw." box="[305,581,789,815]" class="Bryopsida" family="Bryaceae" genus="Bryum" kingdom="Plantae" order="Bryales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="dichotomum">
<emphasis id="B92C9F23FFB37E52E80E6C2FFE752632" box="[305,503,789,815]" italics="true" pageId="11" pageNumber="46">Bryum dichotomum</emphasis>
Hedw.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EB746C2CFEA52652" authority="(Brid.) K. Saito" authorityName="K. Saito" baseAuthorityName="Brid." class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="acutus">
<emphasis id="B92C9F23FFB37E52EB746C2CFC872632" box="[587,773,789,815]" italics="true" pageId="11" pageNumber="46">Didymodon acutus</emphasis>
(Brid.) K.Saito
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E80C6C0CFC85264D" authority="(Hedw.) P. Beauv" authorityName="P. Beauv" baseAuthorityName="Hedw." box="[307,775,821,848]" class="Bryopsida" family="Pottiaceae" genus="Streblotrichum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="convolutum">
<emphasis id="B92C9F23FFB37E52E80C6C0CFDCF2652" box="[307,589,821,847]" italics="true" pageId="11" pageNumber="46">Streblotrichum convolutum</emphasis>
(Hedw.) P.Beauv
</taxonomicName>
and 
<taxonomicName id="4C5838B2FFB37E52E98C6C6CFDD62672" authority="(Sm.) Lindb. In Isere" authorityName="Lindb. In Isere" baseAuthorityName="Sm." box="[179,596,853,880]" class="Bryopsida" family="Pottiaceae" genus="Tortula" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="atrovirens">
<emphasis id="B92C9F23FFB37E52E98C6C6CFEE92672" box="[179,363,853,879]" italics="true" pageId="11" pageNumber="46">Tortula atrovirens</emphasis>
(Sm.) Lindb. In Isère
</taxonomicName>
, main associated species are 
<taxonomicName id="4C5838B2FFB37E52E9CA6C4CFEDB2692" box="[245,345,885,911]" class="Bryopsida" family="Pottiaceae" genus="Aloina" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="aloides">
<emphasis id="B92C9F23FFB37E52E9CA6C4CFEDB2692" box="[245,345,885,911]" italics="true" pageId="11" pageNumber="46">A. aloides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E85A6C4CFC812692" authority="(Bruch &amp; Schimp.) Limpr." authorityName="Limpr." baseAuthorityName="Bruch &amp; Schimp." box="[357,771,885,911]" class="Bryopsida" family="Pottiaceae" genus="Aloina" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="ambigua">
<emphasis id="B92C9F23FFB37E52E85A6C4CFE5F2692" box="[357,477,885,911]" italics="true" pageId="11" pageNumber="46">A. ambigua</emphasis>
(Bruch &amp; Schimp.) Limpr.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E9BC6CACFDDF26B2" authority="(Hedw.) R. H. Zander" authorityName="R. H. Zander" baseAuthorityName="Hedw." box="[131,605,917,943]" class="Bryopsida" family="Pottiaceae" genus="Microbryum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="curvicollum">
<emphasis id="B92C9F23FFB37E52E9BC6CACFEF926B2" box="[131,379,917,943]" italics="true" pageId="11" pageNumber="46">Microbryum curvicollum</emphasis>
(Hedw.) R.H.Zander
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EB5C6CAFFE5F26D2" authority="(Brid.) Blockeel &amp; A. J. E. Sm." authorityName="Blockeel &amp; A. J. E. Sm." baseAuthorityName="Brid." class="Bryopsida" family="Pottiaceae" genus="Pottiopsis" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="caespitosa">
<emphasis id="B92C9F23FFB37E52EB5C6CAFFF2826D3" italics="true" pageId="11" pageNumber="46">Pottiopsis caespitosa</emphasis>
(Brid.) Blockeel &amp; A.J.E.Sm.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E8DB6C8CFE7D26F3" authority="(Mull. Hal.) Holyoak &amp; N. Pedersen" authorityName="Holyoak &amp; N. Pedersen" baseAuthorityName="Mull. Hal." class="Oligohymenophorea" family="Hysterocinetidae" genus="Ptychostomum" kingdom="Chromista" order="Scuticociliatida" pageId="11" pageNumber="46" phylum="Ciliophora" rank="species" species="imbricatulum">
<emphasis id="B92C9F23FFB37E52E8DB6C8CFC8726D2" box="[484,773,949,975]" italics="true" pageId="11" pageNumber="46">Ptychostomum imbricatulum</emphasis>
(Müll.Hal.) Holyoak &amp; N.Pedersen
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EB396CEDFC8126F3" authority="Broth." authorityName="Broth." box="[518,771,980,1006]" class="Bryopsida" family="Pottiaceae" genus="Tortula" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="caucasica">
<emphasis id="B92C9F23FFB37E52EB396CEDFD3626F3" box="[518,692,980,1006]" italics="true" pageId="11" pageNumber="46">Tortula caucasica</emphasis>
Broth.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E9BD6CCCFEDE2113" authority="Broth." authorityName="Broth." box="[130,348,1012,1038]" class="Bryopsida" family="Pottiaceae" genus="Tortula" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="lindbergii">
<emphasis id="B92C9F23FFB37E52E9BD6CCCFE892113" box="[130,267,1012,1038]" italics="true" pageId="11" pageNumber="46">T. lindbergii</emphasis>
Broth.
</taxonomicName>
and 
<taxonomicName id="4C5838B2FFB37E52E8A56CCCFD582112" authority="(Hedw.) Mitt. In" authorityName="Mitt. In" baseAuthorityName="Hedw." box="[410,730,1012,1039]" class="Bryopsida" family="Pottiaceae" genus="Pottia" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="truncata">
<emphasis id="B92C9F23FFB37E52E8A56CCCFD932113" box="[410,529,1013,1038]" italics="true" pageId="11" pageNumber="46">T. truncata</emphasis>
(Hedw.) Mitt. In
</taxonomicName>
the Mediterranean localities they are 
<taxonomicName id="4C5838B2FFB37E52E8E26B2DFF732153" authority="(Spruce) Mull. Hal." authorityName="Mull. Hal." baseAuthorityName="Spruce" class="Bryopsida" family="Pottiaceae" genus="Acaulon" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="triquetrum">
<emphasis id="B92C9F23FFB37E52E8E26B2DFD242133" box="[477,678,1044,1070]" italics="true" pageId="11" pageNumber="46">Acaulon triquetrum</emphasis>
(Spruce) Müll.Hal.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E83F6B0DFE112153" box="[256,403,1076,1102]" class="Bryopsida" family="Pottiaceae" genus="Aloina" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="aloides">
<emphasis id="B92C9F23FFB37E52E83F6B0DFE112153" box="[256,403,1076,1102]" italics="true" pageId="11" pageNumber="46">Aloina aloides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E89D6B0CFDF62153" box="[418,628,1076,1102]" class="Bryopsida" family="Bryaceae" genus="Bryum" kingdom="Plantae" order="Bryales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="dichotomum">
<emphasis id="B92C9F23FFB37E52E89D6B0CFDF62153" box="[418,628,1076,1102]" italics="true" pageId="11" pageNumber="46">Bryum dichotomum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EBBB6B0CFE282172" authority="R. Wilczek &amp; Demaret" authorityName="R. Wilczek &amp; Demaret" authorityYear="1976" class="Bryopsida" family="Bryaceae" genus="Bryum" kingdom="Plantae" order="Bryales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="gemmilucens">
<emphasis id="B92C9F23FFB37E52EBBB6B0CFF2F2173" italics="true" pageId="11" pageNumber="46">B. gemmilucens</emphasis>
R.Wilczek &amp; Demaret
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E88A6B6DFDF52173" box="[437,631,1108,1134]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="acutus">
<emphasis id="B92C9F23FFB37E52E88A6B6DFDF52173" box="[437,631,1108,1134]" italics="true" pageId="11" pageNumber="46">Didymodon acutus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EBBA6B6DFF7C2193" class="Bryopsida" family="Pottiaceae" genus="Microbryum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="curvicollum">
<emphasis id="B92C9F23FFB37E52EBBA6B6DFF7C2193" italics="true" pageId="11" pageNumber="46">Microbryum curvicollum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5838B2FFB37E52E8086B4CFC842192" authority="(Sm.) R. H. Zander. Poorer" authorityName="R. H. Zander. Poorer" baseAuthorityName="Sm." box="[311,774,1140,1167]" class="Bryopsida" family="Pottiaceae" genus="Microbryum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="davallianum">
<emphasis id="B92C9F23FFB37E52E8086B4CFE602193" box="[311,482,1140,1166]" italics="true" pageId="11" pageNumber="46">M. davallianum</emphasis>
(Sm.) R.H.Zander. Poorer
</taxonomicName>
communities are generally dominated by scarce individuals of only two species, 
<taxonomicName id="4C5838B2FFB37E52E8596B8CFDB421D3" box="[358,566,1204,1230]" class="Bryopsida" family="Bryaceae" genus="Bryum" kingdom="Plantae" order="Bryales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="dichotomum">
<emphasis id="B92C9F23FFB37E52E8596B8CFDB421D3" box="[358,566,1204,1230]" italics="true" pageId="11" pageNumber="46">Bryum dichotomum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5838B2FFB37E52EB4F6B8DFE8221F3" class="Bryopsida" family="Pottiaceae" genus="Streblotrichum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="convolutum">
<emphasis id="B92C9F23FFB37E52EB4F6B8DFE8221F3" italics="true" pageId="11" pageNumber="46">Streblotrichum convolutum</emphasis>
</taxonomicName>
. In 
<collectingCountry id="F34F03A1FFB37E52E80E6BEDFE0721F3" box="[305,389,1236,1262]" name="Tunisia" pageId="11" pageNumber="46">Tunisia</collectingCountry>
, the species grows mostly without direct associates. In Andalucía, 
<taxonomicName id="4C5838B2FFB37E52E8F76BCDFDE02010" box="[456,610,1268,1293]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52E8F76BCDFDE02010" box="[456,610,1268,1293]" italics="true" pageId="11" pageNumber="46">P. sampaianum</emphasis>
</taxonomicName>
grew in fissures of exposed conglomerate rock outcrops, and notable associated species included 
<taxonomicName id="4C5838B2FFB37E52E8526A0AFE7C2050" box="[365,510,1331,1357]" class="Bryopsida" family="Pottiaceae" genus="Aloina" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="aloides">
<emphasis id="B92C9F23FFB37E52E8526A0AFE7C2050" box="[365,510,1331,1357]" italics="true" pageId="11" pageNumber="46">Aloina aloides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EB336A0AFEB72070" authority="(Corb.) E. Maier" authorityName="E. Maier" authorityYear="1998" baseAuthorityName="Corb." class="Bryopsida" family="Ptychomitriaceae" genus="Campylostelium" kingdom="Plantae" order="Grimmiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="pitardii">
<emphasis id="B92C9F23FFB37E52EB336A0AFC852050" box="[524,775,1331,1357]" italics="true" pageId="11" pageNumber="46">Campylostelium pitardii</emphasis>
(Corb.) E.Maier
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E8006A6AFC802073" authority="Holz. &amp; E. B. Bartram" authorityName="Holz. &amp; E. B. Bartram" box="[319,770,1363,1390]" class="Bryopsida" family="Pottiaceae" genus="Crossidium" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="aberrans">
<emphasis id="B92C9F23FFB37E52E8006A6AFD922070" box="[319,528,1363,1389]" italics="true" pageId="11" pageNumber="46">Crossidium aberrans</emphasis>
Holz. &amp; E.B.Bartram
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E9BB6A4AFE732090" authority="(Brid.) K. Saito" authorityName="K. Saito" baseAuthorityName="Brid." box="[132,497,1395,1421]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="acutus">
<emphasis id="B92C9F23FFB37E52E9BB6A4AFEC52090" box="[132,327,1395,1421]" italics="true" pageId="11" pageNumber="46">Didymodon acutus</emphasis>
(Brid.) K.Saito
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E8C36A4DFE1A20B0" authority="(J. Froehl.) J. A. Jimenez &amp; M. J. Cano" authorityName="J. A. Jimenez &amp; M. J. Cano" baseAuthorityName="J. Froehl." class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="desertorum">
<emphasis id="B92C9F23FFB37E52E8C36A4DFD122090" box="[508,656,1395,1421]" italics="true" pageId="11" pageNumber="46">D. desertorum</emphasis>
(J.Froehl.) J.A.Jiménez &amp; M.J.Cano
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52E89B6AAAFD3120B0" authority="Hedw." authorityName="Hedw." box="[420,691,1427,1453]" class="Bryopsida" family="Encalyptaceae" genus="Encalypta" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="vulgaris">
<emphasis id="B92C9F23FFB37E52E89B6AAAFDE020B0" box="[420,610,1427,1453]" italics="true" pageId="11" pageNumber="46">Encalypta vulgaris</emphasis>
Hedw.
</taxonomicName>
, 
<taxonomicName id="4C5838B2FFB37E52EB816AADFE3220D0" authority="(Turner)" baseAuthorityName="Turner" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="muhlenbergii">
<emphasis id="B92C9F23FFB37E52EB816AADFECD20D0" italics="true" pageId="11" pageNumber="46">Entosthodon muhlenbergii</emphasis>
(Turner)
</taxonomicName>
Fife, 
<taxonomicName id="4C5838B2FFB37E52E8D86A8AFEDB20F0" authority="(Brid.) R. H. Zander" authorityName="R. H. Zander" baseAuthorityName="Brid." class="Bryopsida" family="Pottiaceae" genus="Pseudocrossidium" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="revolutum">
<emphasis id="B92C9F23FFB37E52E8D86A8AFC8720D0" box="[487,773,1459,1485]" italics="true" pageId="11" pageNumber="46">Pseudocrossidium revolutum</emphasis>
(Brid.) R.H.Zander
</taxonomicName>
and 
<taxonomicName id="4C5838B2FFB37E52E8B06AEAFDCA20F0" box="[399,584,1491,1517]" class="Bryopsida" family="Pottiaceae" genus="Tortula" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="atrovirens">
<emphasis id="B92C9F23FFB37E52E8B06AEAFDCA20F0" box="[399,584,1491,1517]" italics="true" pageId="11" pageNumber="46">Tortula atrovirens</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE74331FFB37E52E9A46ACAFEFF2296" blockId="11.[130,777,215,2028]" pageId="11" pageNumber="46">
Most significantly, 
<taxonomicName id="4C5838B2FFB37E52E84F6ACAFD1C2311" box="[368,670,1523,1548]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52E84F6ACAFD1C2311" box="[368,670,1523,1548]" italics="true" pageId="11" pageNumber="46">Pterygoneurum sampaianum</emphasis>
</taxonomicName>
generally occurs in very open micro-communities (
<figureCitation id="13635FB4FFB37E52EB71692BFD0F2330" box="[590,653,1554,1581]" captionStart="FIG" captionStartId="10.[286,297,2007,2025]" captionTargetBox="[173,1416,215,1878]" captionTargetId="figure-45@10.[746,1322,768,1324]" captionTargetPageId="10" captionText="FIG. 4. — Habitat of Pterygoneurum sampaianum (Guim.) Guim. (11 March 2020, Tunisia, Siliana Governorate, Sidi Ameur)." figureDoi="http://doi.org/10.5281/zenodo.13378503" httpUri="https://zenodo.org/record/13378503/files/figure.png" pageId="11" pageNumber="46">Fig. 4</figureCitation>
), with low cover of vegetation, and very sparse bryophytes, which strongly suggests a high sensitivity to competition. Although this is a generally well-known trait of 
<taxonomicName id="4C5838B2FFB37E52E8E7694BFDC62391" box="[472,580,1650,1676]" class="Bryopsida" family="Pottiaceae" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="family">Pottiaceae</taxonomicName>
communities, the species’ apparent inability to invade more closed populations of larger species in the immediate vicinity appears to be significant. In the Pyrénées-Orientales and in Aude (
<collectingCountry id="F34F03A1FFB37E52EBB769EBFD5723F1" box="[648,725,1746,1772]" name="France" pageId="11" pageNumber="46">France</collectingCountry>
), the species is strongly dependant on repeated trampling of paths in limestone plateaus where it grows in linear populations on a restricted fringe at the transition between small grassland communities and totally bare substrate (corresponding to car tracks or hiking paths).
</paragraph>
<paragraph id="8BE74331FFB37E52E9A468A8FC4F272D" blockId="11.[130,777,215,2028]" lastBlockId="11.[811,1457,215,911]" pageId="11" pageNumber="46">
The general association of 
<taxonomicName id="4C5838B2FFB37E52E89468ABFD4D22B6" box="[427,719,1938,1963]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52E89468ABFD4D22B6" box="[427,719,1938,1963]" italics="true" pageId="11" pageNumber="46">Pterygoneurum sampaianum</emphasis>
</taxonomicName>
with fine clay and constraining substrates is also important. Initially it was believed to grow on the chalk clay of the Algarve (
<bibRefCitation id="EFC93EC0FFB37E52EA096FEEFC6E25EC" author="GUIMARAES A. L. M." box="[822,1004,215,241]" pageId="11" pageNumber="46" pagination="52 - 52" refId="ref8511" refString="GUIMARAES A. L. M. 1917. - Uma Pottia nova do Algarve. Annaes Scientificos da Academia Polytechnica do Porto 12: 52 - 52." type="journal article" year="1917">Guimarães 1917</bibRefCitation>
) and on gypsiferous or saline soils in SE and NE Spain (
<bibRefCitation id="EFC93EC0FFB37E52EAF16FCEFB1B240F" author="GUERRA J. &amp; ROS R. M. &amp; CANO M. J. &amp; CASARES M." box="[974,1177,247,274]" pageId="11" pageNumber="46" pagination="125 - 135" refId="ref8414" refString="GUERRA J., ROS R. M., CANO M. J. &amp; CASARES M. 1995 b. - Gypsiferous outcrops in SE Spain, refuges of rare, vulnerable and endangered bryophytes and lichens. Cryptogamie, Bryologie Lichenologie 16: 125 - 135." type="journal article" year="1995">
Guerra 
<emphasis id="B92C9F23FFB37E52ED216FC1FBD2240C" box="[1054,1104,247,273]" italics="true" pageId="11" pageNumber="46">et al.</emphasis>
1995b
</bibRefCitation>
). 
<bibRefCitation id="EFC93EC0FFB37E52ED916FCEFA2D240C" author="CASAS C. &amp; BRUGUES M." box="[1198,1455,247,273]" pageId="11" pageNumber="46" pagination="103 - 114" refId="ref8332" refString="CASAS C. &amp; BRUGUES M. 1978. - Nova aportacio al coneixement de la brioflora dels Monegros. Anales del Instituto Botanico A. J. Cavanilles 35: 103 - 114." type="journal article" year="1978">Casas &amp; Brugués (1978)</bibRefCitation>
describe the microhabitat as a fine layer of clay deposited in the depressions. In 
<collectingCountry id="F34F03A1FFB37E52EAC16E0EFBD2244C" box="[1022,1104,311,337]" name="Tunisia" pageId="11" pageNumber="46">Tunisia</collectingCountry>
, the species is strongly linked to degraded habitats, mainly the flanks of small mountainous systems where fluvial flows during violent storms can carve rills and shape badlands characteristic of severely eroded soils. Important quantities of very fine-grained detritic “glacis” accumulate at the foot of the slopes, periodically regenerating new bare habitat suitable for the most diminutive 
<taxonomicName id="4C5838B2FFB37E52ECBD6ECEFCF7272D" class="Bryopsida" family="Pottiaceae" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="family">Pottiaceae</taxonomicName>
species.
</paragraph>
<paragraph id="8BE74331FFB37E52EA7C6D0FFAA12692" blockId="11.[811,1457,215,911]" pageId="11" pageNumber="46">
An examination of the subterranean system of 
<taxonomicName id="4C5838B2FFB37E52EC716D0EFC7F276D" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52EC716D0EFC7F276D" italics="true" pageId="11" pageNumber="46">Pterygoneurum sampaianum</emphasis>
</taxonomicName>
, made of large and thick-walled rhizoids (compared to the significantly thinner rhizoids of 
<taxonomicName id="4C5838B2FFB37E52EC7F6D4EFA27278D" box="[1344,1445,631,656]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="ovatum">
<emphasis id="B92C9F23FFB37E52EC7F6D4EFA27278D" box="[1344,1445,631,656]" italics="true" pageId="11" pageNumber="46">P. ovatum</emphasis>
</taxonomicName>
) anchored to semi-buried succulent stem, suggests that it could be linked to perennation. In the allegedly short-lived 
<taxonomicName id="4C5838B2FFB37E52EC666D8CFC2327F2" class="Bryopsida" family="Pottiaceae" genus="Acaulon" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="triquetrum">
<emphasis id="B92C9F23FFB37E52EC666D8CFC2327F2" italics="true" pageId="11" pageNumber="46">Acaulon triquetrum</emphasis>
</taxonomicName>
, long-term monitoring allowed 
<bibRefCitation id="EFC93EC0FFB37E52EC306DECFA3327ED" author="AHRENS M." box="[1295,1457,725,752]" pageId="11" pageNumber="46" pagination="239 - 274" refId="ref8186" refString="AHRENS M. 2003. - Untersuchungen zum Lebenszyklus von Acaulon triquetrum (Bryopsida, Pottiaceae). Herzogia 16: 239 - 274." type="journal article" year="2003">Ahrens (2003)</bibRefCitation>
to uncover the species’ remarkable persistence which turns out to be pauciennial instead. This calls for further studies of similar small pottiaceous mosses since our knowledge of their biology is based on generalizations that do not seem to hold up with a careful individual examination.
</paragraph>
</subSubSection>
<subSubSection id="C34210BAFFB37E52EA126C8CFB382231" pageId="11" pageNumber="46" type="conservation">
<paragraph id="8BE74331FFB37E52EA126C8CFBA926D0" blockId="11.[813,1067,949,975]" box="[813,1067,949,975]" pageId="11" pageNumber="46">
<heading id="D0AFF45DFFB37E52EA126C8CFBA926D0" allCaps="true" box="[813,1067,949,975]" fontSize="9" level="2" pageId="11" pageNumber="46" reason="2">
<smallCapsWord id="8D01D5EDFFB37E52EA126C8CFC5526D0" baselines="968,968" box="[813,983,949,975]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Conservation" pageId="11" pageNumber="46">CONSERVATION</smallCapsWord>
<smallCapsWord id="8D01D5EDFFB37E52EAE16C83FBA926D0" baselines="968" box="[990,1067,954,973]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="status" pageId="11" pageNumber="46">STATUS</smallCapsWord>
</heading>
</paragraph>
<paragraph id="8BE74331FFB37E52EA126CEDFB382231" blockId="11.[811,1457,980,1836]" pageId="11" pageNumber="46">
The new data on 
<taxonomicName id="4C5838B2FFB37E52EAE76CECFB7726F3" box="[984,1269,981,1006]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52EAE76CECFB7726F3" box="[984,1269,981,1006]" italics="true" pageId="11" pageNumber="46">Pterygoneurum sampaianum</emphasis>
</taxonomicName>
require to address the possible adjustments to its so far estimated conservation status. Based on the rare occurrences recorded so far in Europe, the species was evaluated as Near Threatened in the last version of European Red List of Mosses (
<bibRefCitation id="EFC93EC0FFB37E52EDB26B6DFAEE2173" author="SERGIO C. &amp; CANO M. J." box="[1165,1388,1108,1134]" pageId="11" pageNumber="46" refId="ref9372" refString="SERGIO C. &amp; CANO M. J. 2019. - Pterygoneurum sampaianum (Europe assessment). The IUCN Red List of Threatened Species 2019: e. T 87562666 A 87729799. Accessed on 03 October 2023." type="journal volume" year="2019">Sérgio &amp; Cano 2019</bibRefCitation>
). This assessment has probably mainly been based on the situation in 
<collectingCountry id="F34F03A1FFB37E52EA746BADFC0A21B3" box="[843,904,1172,1198]" name="Spain" pageId="11" pageNumber="46">Spain</collectingCountry>
where it is assessed as Least Concern (
<bibRefCitation id="EFC93EC0FFB37E52EC076BADFBC721D3" author="BRUGUES M. &amp; GONZALEZ-MANCEBO J. M." pageId="11" pageNumber="46" pagination="26 - 42" refId="ref8277" refString="BRUGUES M. &amp; GONZALEZ-MANCEBO J. M. 2012. - Lista Roja de los Briofitos Amenazados de Espana, in GARILLETI R. &amp; ALBERTOS B. (eds), Atlas y Libro Rojo de los Briofitos amenazados de Espana. Organismo Autonomo Parques Nacionales, Madrid: 26 - 42." type="book chapter" year="2012">Brugués &amp; González-Mancebo 2012</bibRefCitation>
), as elsewhere in its distribution area, the species was evaluated as Regionally Extinct (RE) in 
<collectingCountry id="F34F03A1FFB37E52EA736BCAFC252010" box="[844,935,1267,1293]" name="Portugal" pageId="11" pageNumber="46">Portugal</collectingCountry>
and Data Deficient in 
<collectingCountry id="F34F03A1FFB37E52ED9D6BCAFA9D2010" box="[1186,1311,1267,1293]" name="Switzerland" pageId="11" pageNumber="46">Switzerland</collectingCountry>
(
<bibRefCitation id="EFC93EC0FFB37E52EC126BCAFC4E2033" author="HODGETTS N. &amp; LOCKHART N." pageId="11" pageNumber="46" pagination="1 - 214" refId="ref8577" refString="HODGETTS N. &amp; LOCKHART N. 2020. - Checklist and country status of European bryophytes. Update 2020. Irish Wildlife Manuals 123: 1 - 214. http: // hdl. handle. net / 2262 / 93026" type="journal article" year="2020">Hodgetts &amp; Lockhart 2020</bibRefCitation>
). With the now more accurate morphological circumscription and better knowledge about its habitat in the Mediterranean region, additional records can be expected in the near future. However, vast areas of apparently favourable habitats that do not seem to be occupied by 
<taxonomicName id="4C5838B2FFB37E52EC2E6AADFA3220B0" box="[1297,1456,1428,1453]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52EC2E6AADFA3220B0" box="[1297,1456,1428,1453]" italics="true" pageId="11" pageNumber="46">P. sampaianum</emphasis>
</taxonomicName>
exist in the Mediterranean. Considering the species’ apparent weakness in competition, ecological specialization, and apparent rarity, its threat status cannot currently be lowered but rather, it can be discussed to be evaluated as Vulnerable with respect to the population size which nearly certainly is smaller than 10 000 mature individuals and small individual populations hardly exceeding one hundred individuals in conjunction with the possibly decreasing overall size based on the suitable habitat degradation in the future risk assessment. Therefore, conservation efforts should focus on preserving the habitats where 
<taxonomicName id="4C5838B2FFB37E52EAEC69CAFBF02211" box="[979,1138,1779,1804]" class="Bryopsida" family="Pottiaceae" genus="Pterygoneurum" kingdom="Plantae" order="Pottiales" pageId="11" pageNumber="46" phylum="Bryophyta" rank="species" species="sampaianum">
<emphasis id="B92C9F23FFB37E52EAEC69CAFBF02211" box="[979,1138,1779,1804]" italics="true" pageId="11" pageNumber="46">P. sampaianum</emphasis>
</taxonomicName>
occurs, ensuring the survival of this species for future generations.
</paragraph>
</subSubSection>
</treatment>
</document>