252 lines
33 KiB
XML
252 lines
33 KiB
XML
<document id="8DD8BCBA1C49BC6252BA6FC150D5E36A" ID-DOI="10.11646/zootaxa.5260.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7794920" ID-ZooBank="7171477F-DB75-4CF6-9507-3329F5D7A9F7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680527871645" checkinUser="plazi" docAuthor="Blake, James A. & Maciolek, Nancy J." docDate="2023" docId="03F1A22EFF91CB27FF7282C3FC24FDC2" docLanguage="en" docName="zootaxa.5260.1.1.pdf" docOrigin="Zootaxa 5260 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heterospio southwardorum Blake & Maciolek 2023, new species" docType="treatment" docUuid="A65BA064-DF88-44FC-A223-A47C4CDABCC9" docUuidSource="ZooBank" docVersion="4" lastPageNumber="32" masterDocId="FFC8DA56FF8CCB38FFE58038FFEAFFA3" masterDocTitle="New species and records of Heterospio (Annelida, Longosomatidae) from continental shelf, slope and abyssal depths of the Atlantic Ocean, Pacific Ocean, Indian Ocean and adjacent seas" masterLastPageNumber="74" masterPageNumber="1" pageNumber="30" updateTime="1699481072705" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="B9BC3C75DC8E5777FDD20D81EEAA050B">New species and records of Heterospio (Annelida, Longosomatidae) from continental shelf, slope and abyssal depths of the Atlantic Ocean, Pacific Ocean, Indian Ocean and adjacent seas</mods:title>
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<mods:roleTerm id="0AD67862D94D8650EE9B4C347986CB67">Author</mods:roleTerm>
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<mods:namePart id="B62DE2690254C5E65C978D650927AADB">Blake, James A.</mods:namePart>
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<mods:nameIdentifier id="A66362D076BDBD5509B12E8CAD08BA9F" type="email">jablake9@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="CA18328DA0A5754EC34254D81675D905">Author</mods:roleTerm>
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<mods:namePart id="EC5D53CCF4F5580F96B6FF0D11E5CA39">Maciolek, Nancy J.</mods:namePart>
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<mods:nameIdentifier id="B407606157AEDAF4A6972178A5F14191" type="ORCID">0000-0003-0195-0713</mods:nameIdentifier>
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<mods:nameIdentifier id="E76C7755C1BD7BF394283E2BDEF711B2" type="email">njmaciolek@gmail.com</mods:nameIdentifier>
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<mods:title id="D6FD9711C5979565CF75A62886108610">Zootaxa</mods:title>
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<mods:part id="40E4F44509FE1DF31045261B154C9C16">
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<mods:date id="4C7BDD85CEF4E3FA293A3D739E742F6D">2023</mods:date>
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<mods:number id="5D97E6A9B364320FD9FBF878FE95C9AC">2023-04-03</mods:number>
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<mods:number id="A5F7E3393509C8BD8C55A68532C4D262">5260</mods:number>
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<mods:detail id="8CA90A10BBA3787EF18F073FD76287E5" type="issue">
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<mods:number id="F2D6F16488651E6FB65D2609C6E238B2">1</mods:number>
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<mods:url id="A825FF2DB0A0C1935A4D64E5FC6923BC">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
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<mods:classification id="3B42164E6AB7D70F6D5F95430358DB9C">journal article</mods:classification>
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<mods:identifier id="A9B1D912A4BA2AA1F3C8D7703CA9942C" type="DOI">10.11646/zootaxa.5260.1.1</mods:identifier>
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<mods:identifier id="EEBDAC3C540CD20A39E26EA66F250797" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F1A22EFF91CB27FF7282C3FC24FDC2" ID-DOI="http://doi.org/10.5281/zenodo.7798809" ID-Zenodo-Dep="7798809" LSID="urn:lsid:zoobank.org:act:A65BA064-DF88-44FC-A223-A47C4CDABCC9" httpUri="http://treatment.plazi.org/id/03F1A22EFF91CB27FF7282C3FC24FDC2" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
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<subSubSection id="C34240B3FF91CB25FF7282C3FD90FCB7" box="[151,634,762,789]" pageId="29" pageNumber="30" type="nomenclature">
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<paragraph id="8BE71338FF91CB25FF7282C3FD90FCB7" blockId="29.[151,996,762,861]" box="[151,634,762,789]" pageId="29" pageNumber="30">
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<heading id="D0AFA454FF91CB25FF7282C3FD90FCB7" bold="true" box="[151,634,762,789]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="4">
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<emphasis id="B92CCF2AFF91CB25FF7282C3FD90FCB7" bold="true" box="[151,634,762,789]" pageId="29" pageNumber="30">
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<taxonomicName id="4C5868BBFF91CB25FF7282C3FE09FCB6" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[151,483,762,789]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="southwardorum" status="sp. nov.">
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<emphasis id="B92CCF2AFF91CB25FF7282C3FE09FCB6" bold="true" box="[151,483,762,789]" italics="true" pageId="29" pageNumber="30">Heterospio southwardorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21F7251FF91CB25FE0F82C3FD90FCB7" box="[490,634,762,789]" pageId="29" pageNumber="30" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34240B3FF91CB25FF728327FC0EFCFE" pageId="29" pageNumber="30" type="description">
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<paragraph id="8BE71338FF91CB25FF728327FEE9FC9A" blockId="29.[151,996,762,861]" box="[151,259,799,825]" pageId="29" pageNumber="30">
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<figureCitation id="13630FBDFF91CB25FF728327FEE9FC9A" box="[151,259,799,825]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Figure 12</figureCitation>
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</paragraph>
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<paragraph id="8BE71338FF91CB25FF72837AFC0EFCFE" blockId="29.[151,996,762,861]" box="[151,996,834,861]" pageId="29" pageNumber="30">
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<uri id="FFC91F3AFF91CB25FF72837AFC0EFCFE" box="[151,996,834,861]" pageId="29" pageNumber="30">
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urn:lsid:zoobank.org:act:
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<uuid id="FFFE29EDFF91CB25FE43837AFC0EFCFE" box="[422,996,834,861]" pageId="29" pageNumber="30">A65BA064-DF88-44FC-A223-A47C4CDABCC9</uuid>
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</uri>
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</subSubSection>
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<subSubSection id="C34240B3FF91CB25FF7283B2FB05FC6A" pageId="29" pageNumber="30" type="materials_examined">
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<paragraph id="8BE71338FF91CB25FF7283B2FB05FC6A" blockId="29.[151,1437,906,2013]" pageId="29" pageNumber="30">
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<emphasis id="B92CCF2AFF91CB25FF7283B2FC72FC07" bold="true" box="[151,920,906,933]" pageId="29" pageNumber="30">Material examined. NE Atlantic Ocean, Bay of Biscay, off Bilbao,</emphasis>
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<materialsCitation id="3B301965FF91CB25FC7A83B2FC05FC6A" collectingDate="1967-07-19" collectionCode="R, V" collectorName="A. & E. Southward" country="Spain" elevation="641" latitude="43.717" location="Spain" longLatPrecision="68" longitude="-3.797" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
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<emphasis id="B92CCF2AFF91CB25FC7A83B2FC09FC07" bold="true" box="[927,995,906,932]" pageId="29" pageNumber="30">
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<collectingCountry id="F34F53A8FF91CB25FC7A83B2FC09FC07" box="[927,995,906,932]" name="Spain" pageId="29" pageNumber="30">Spain</collectingCountry>
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</emphasis>
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,
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<collectionCode id="ED498BFDFF91CB25FC1483B3FBEEFC06" box="[1009,1028,907,933]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="30">R</collectionCode>
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/
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<collectionCode id="ED498BFDFF91CB25FBE983B3FBCAFC06" box="[1036,1056,907,933]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="29" pageNumber="30" type="Museum">V</collectionCode>
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<taxonomicName id="4C5868BBFF91CB25FBC383B3FB29FC06" authority=", Cruise" authorityName="Cruise" box="[1062,1219,907,933]" class="Hydrozoa" family="Corynidae" genus="Sarsia" kingdom="Animalia" order="Anthoathecata" pageId="29" pageNumber="30" phylum="Cnidaria" rank="genus">
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<emphasis id="B92CCF2AFF91CB25FBC383B3FB84FC06" box="[1062,1134,907,933]" italics="true" pageId="29" pageNumber="30">Sarsia</emphasis>
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, Cruise
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</taxonomicName>
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5,
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<emphasis id="B92CCF2AFF91CB25FB0083B2FAA6FC07" bold="true" box="[1253,1356,906,932]" pageId="29" pageNumber="30">Sta. 5-56</emphasis>
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,
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<date id="FFE635F8FF91CB25FABC83B3FF38FC6A" pageId="29" pageNumber="30" value="1967-07-19">
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<collectingDate id="EFA2CC10FF91CB25FABC83B3FF38FC6A" pageId="29" pageNumber="30" value="1967-07-19">19 Jul 1967</collectingDate>
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</date>
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, coll.
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<collectorName id="26AD76EEFF91CB25FEF18397FE04FC6A" box="[276,494,943,969]" pageId="29" pageNumber="30">A. & E. Southward</collectorName>
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, EBS,
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<geoCoordinate id="EE6C75FFFF91CB25FDDF8397FD40FC6A" box="[570,682,943,969]" degrees="43.717" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="43.717">43.717°N</geoCoordinate>
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,
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<geoCoordinate id="EE6C75FFFF91CB25FD508397FCCDFC6A" box="[693,807,943,969]" degrees="03.797" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="55" value="-3.797">03.797°W</geoCoordinate>
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,
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<quantity id="4CA0BEDDFF91CB25FCD68397FC96FC6A" box="[819,892,943,969]" metricMagnitude="2" metricUnit="m" metricValue="6.409999999999999" pageId="29" pageNumber="30" unit="m" value="641.0">
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<elevation id="0075F40BFF91CB25FCD68397FC96FC6A" box="[819,892,943,969]" metricMagnitude="2" metricUnit="m" metricValue="6.409999999999999" pageId="29" pageNumber="30" unit="m" value="641.0">641 m</elevation>
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</quantity>
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,
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<typeStatus id="54E3AD9AFF91CB25FC6D8397FC05FC6A" box="[904,1007,943,969]" pageId="29" pageNumber="30" type="holotype">
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<emphasis id="B92CCF2AFF91CB25FC6D8397FC05FC6A" bold="true" box="[904,1007,943,969]" pageId="29" pageNumber="30">holotype</emphasis>
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</typeStatus>
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</materialsCitation>
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<materialsCitation id="3B301965FF91CB25FC108396FB00FC6A" box="[1013,1258,942,969]" collectionCode="V" country="Spain" pageId="29" pageNumber="30" specimenCode="LACM-AHF 13283" specimenCount="1" typeStatus="holotype">
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<emphasis id="B92CCF2AFF91CB25FC108396FC14FC6B" bold="true" box="[1013,1022,942,968]" pageId="29" pageNumber="30">(</emphasis>
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<specimenCode id="DBFEBB43FF91CB25FC1A8397FB09FC6A" box="[1023,1251,943,969]" collectionCode="LACM-AHF" pageId="29" pageNumber="30">LACM-AHF 13283</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34240B3FF91CB27FF2283EAFBB0FF12" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="description">
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<paragraph id="8BE71338FF91CB25FF2283EAFEA9FB02" blockId="29.[151,1437,906,2013]" pageId="29" pageNumber="30">
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<emphasis id="B92CCF2AFF91CB25FF2283EAFEBBFC4F" bold="true" box="[199,337,978,1004]" pageId="29" pageNumber="30">Description</emphasis>
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. A moderately sized species,
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<typeStatus id="54E3AD9AFF91CB25FD7C83EBFD11FC4E" box="[665,763,979,1005]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
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<emphasis id="B92CCF2AFF91CB25FCE483EAFCE0FC4F" bold="true" box="[769,778,978,1004]" pageId="29" pageNumber="30">(</emphasis>
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LACM-AHF 13283) incomplete with 13 setigers,
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<quantity id="4CA0BEDDFF91CB25FADC83EBFA77FC4E" box="[1337,1437,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="29" pageNumber="30" unit="mm" value="21.3">21.3 mm</quantity>
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long,
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<quantity id="4CA0BEDDFF91CB25FF3283CFFED1FBB2" box="[215,315,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="29" pageNumber="30" unit="mm" value="0.43">0.43 mm</quantity>
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wide across thoracic segments,
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<quantity id="4CA0BEDDFF91CB25FD4683CFFCEDFBB2" box="[675,775,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="29" pageNumber="30" unit="mm" value="0.37">0.37 mm</quantity>
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wide across middle abdominal segment. Thorax with eight crowded segments followed by elongate setiger 9. Setigers 1–7 twice as wide as long; setiger 8 slightly longer, about as wide as long; setiger 9 greatly elongated, as long as setigers 1–8 combined. Abdominal setigers 10–13 long, narrow, encompassing about
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<quantity id="4CA0BEDDFF91CB25FE39845BFDC1FBDE" box="[476,555,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="29" pageNumber="30" unit="mm" value="16.0">16 mm</quantity>
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of total length of
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<typeStatus id="54E3AD9AFF91CB25FD17845BFCB2FBDE" box="[754,856,1123,1149]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
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. Color in alcohol opaque white to light tan; pigment entirely absent.
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</paragraph>
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<paragraph id="8BE71338FF91CB25FF228493FBDEFA3E" blockId="29.[151,1437,906,2013]" pageId="29" pageNumber="30">
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Pre-setiger region short, as long as first two thoracic setigers (
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<figureCitation id="13630FBDFF91CB25FC578493FBA8FB66" box="[946,1090,1195,1221]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12A–B</figureCitation>
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). Prostomium flattened, disklike, broadly rounded anteriorly (
|
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<figureCitation id="13630FBDFF91CB25FDF384F7FD49FB4A" box="[534,675,1231,1257]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12A–B</figureCitation>
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); eyespots absent; nuchal organs elongate slits anterior to anterior peristomial ring (
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<figureCitation id="13630FBDFF91CB25FE8584CBFE26FAAE" box="[352,460,1267,1293]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12A</figureCitation>
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). Peristomium with two rings; first ring narrow, dorsally visible posterior to nuchal organs, not continuing ventrally; second ring large, similar in size and shape to following setigers, interrupted by broad dorsal crest that merges with setiger 1 (
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<figureCitation id="13630FBDFF91CB25FD2C8503FCD9FAF6" box="[713,819,1339,1365]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12A</figureCitation>
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), entire on ventral surface, interrupted by mouth (
|
||
<figureCitation id="13630FBDFF91CB25FA8A8503FF23FADA" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12B</figureCitation>
|
||
). Dorsal tentacles not present, but scars evident in notch between peristomial rings. Mouth a triangular-shaped opening surrounded laterally by thickened lobes (
|
||
<figureCitation id="13630FBDFF91CB25FD2485BBFCC2FA3E" box="[705,808,1411,1437]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12B</figureCitation>
|
||
); proboscis not everted.
|
||
</paragraph>
|
||
<paragraph id="8BE71338FF91CB25FF22859FFCE3F942" blockId="29.[151,1437,906,2013]" pageId="29" pageNumber="30">
|
||
Branchiae or their stubs present on setigers 2–7 (
|
||
<figureCitation id="13630FBDFF91CB25FD0A859FFCB2FA62" box="[751,856,1447,1473]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12A</figureCitation>
|
||
); intact branchiae short, oval-shaped. All parapodia biramous with setal fascicles arising from near anterior edge of segment. Thoracic region with eight short setigers (
|
||
<figureCitation id="13630FBDFF91CB25FF7A85D7FEE0F9AA" box="[159,266,1519,1545]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12A</figureCitation>
|
||
), each slightly wider than long; setiger 8 slightly longer than those preceding. Setiger 9 elongated, as long as setigers 1–8 combined. All thoracic setigers slightly flattened dorsally with notopodia of setigers 1–7 weakly inflated and elevated over dorsum; notopodia of setigers 8–9 with thickened lobes from which setae arise; neuropodia of setigers 1–9 with short, postsetal lobes (
|
||
<figureCitation id="13630FBDFF91CB25FD148663FCBCF9D6" box="[753,854,1627,1653]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12B</figureCitation>
|
||
), most evident on setigers 1–7. Abdominal segments from setiger 10 rounded in cross section with parapodia on anterior borders as transverse ridge encircling body, producing cinctures or rows of setae similar to those of some cirratulids; these ridges interrupted dorsally, ventrally, and laterally denoting location of noto- and neuropodia.
|
||
</paragraph>
|
||
<paragraph id="8BE71338FF91CB25FF2286D3FC3EF81A" blockId="29.[151,1437,906,2013]" pageId="29" pageNumber="30">
|
||
All thoracic notopodia of setigers1–9 with 12–16 long capillaries in spreading fascicles; capillaries of neuropodia more numerous, with 25–30 arranged in tight, dense fascicle. Setae of noto- and neuropodia of following setigers in two transverse rows; setigers 10–11 all capillaries, rows with distinct dorsal and lateral gaps; setiger 12 with all capillaries in notopodia and a few acicular spines in anterior row of neuropodia; setiger 13 with cinctures of 10–12 spines in anterior row and 25–30 capillaries (
|
||
<figureCitation id="13630FBDFF91CB25FD388743FCA8F836" box="[733,834,1915,1941]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12C</figureCitation>
|
||
) in posterior row; capillaries with thin fine tip; spines flattened, tapering to narrow pointed tip, none with aristate tip (
|
||
<figureCitation id="13630FBDFF91CB25FCBA87A7FC2DF81A" box="[863,967,1951,1977]" captionStart="FIGURE 12" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[212,1370,197,1904]" captionTargetId="figure-18@30.[192,1395,181,1920]" captionTargetPageId="30" captionText="FIGURE 12. Heterospio southwardorum n. sp. Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13." figureDoi="http://doi.org/10.5281/zenodo.7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="29" pageNumber="30">Fig. 12D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE71338FF91CB25FF2287FBFD87F87E" blockId="29.[151,1437,906,2013]" box="[199,621,1987,2013]" pageId="29" pageNumber="30">Posterior bulbous section not present.</paragraph>
|
||
<caption id="DF2743B0FF92CB26FF7287AFFB3EF877" ID-DOI="http://doi.org/10.5281/zenodo.7794944" ID-Zenodo-Dep="7794944" httpUri="https://zenodo.org/record/7794944/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1943,1967]" targetBox="[212,1370,197,1904]" targetPageId="30">
|
||
<paragraph id="8BE71338FF92CB26FF7287AFFB3EF877" blockId="30.[151,1436,1943,2004]" pageId="30" pageNumber="31">
|
||
<emphasis id="B92CCF2AFF92CB26FF7287AFFEF2F80C" bold="true" box="[151,280,1943,1967]" pageId="30" pageNumber="31">FIGURE 12</emphasis>
|
||
.
|
||
<taxonomicName id="4C5868BBFF92CB26FEC087A0FDD0F813" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[293,570,1944,1968]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="southwardorum" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFF92CB26FEC087A0FDD0F813" box="[293,570,1944,1968]" italics="true" pageId="30" pageNumber="31">Heterospio southwardorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FF92CB26FDDA87A0FD9FF80C" box="[575,629,1944,1967]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype (LACM-AHF Poly 13283): A, anterior end (setiger 1–9), dorsal view; B, anterior end (setiger 1–5), ventral view; C, capillary neuroseta, setiger 13; D, acicular spines, setiger 13.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE71338FF93CB27FF2280AFFBB0FF12" blockId="31.[151,1437,150,609]" box="[199,1114,150,177]" pageId="31" pageNumber="32">
|
||
<emphasis id="B92CCF2AFF93CB27FF2280AFFE3EFF13" bold="true" box="[199,468,150,177]" pageId="31" pageNumber="32">Methyl Green staining</emphasis>
|
||
. No pattern, stain not concentrated anywhere along body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34240B3FF93CB27FF228083FAADFE72" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="8BE71338FF93CB27FF228083FAADFE72" blockId="31.[151,1437,150,609]" pageId="31" pageNumber="32">
|
||
<emphasis id="B92CCF2AFF93CB27FF228083FEDEFF76" bold="true" box="[199,308,187,213]" pageId="31" pageNumber="32">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C5868BBFF93CB27FEA48083FD9BFF76" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[321,625,187,213]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="southwardorum" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFF93CB27FEA48083FD9BFF76" box="[321,625,187,213]" italics="true" pageId="31" pageNumber="32">Heterospio southwardorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FF93CB27FD928083FD5FFF76" box="[631,693,187,213]" pageId="31" pageNumber="32" rank="species">
|
||
<emphasis id="B92CCF2AFF93CB27FD928083FD47FF76" bold="true" box="[631,685,187,213]" pageId="31" pageNumber="32">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is most similar to
|
||
<taxonomicName id="4C5868BBFF93CB27FC988083FBDEFF76" box="[893,1076,187,213]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="mediterranea">
|
||
<emphasis id="B92CCF2AFF93CB27FC988083FBDEFF76" box="[893,1076,187,213]" italics="true" pageId="31" pageNumber="32">H. mediterranea</emphasis>
|
||
</taxonomicName>
|
||
. Both species have a broad disklike prostomium, two peristomial rings, dorsal peristomial crest, and in both species setiger 9 is the first elongate segment, being as long as setigers 1–8 combined. The two species differ in that in
|
||
<taxonomicName id="4C5868BBFF93CB27FBB7813BFACCFEBE" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[1106,1318,259,285]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="southwardorum" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFF93CB27FBB7813BFACCFEBE" box="[1106,1318,259,285]" italics="true" pageId="31" pageNumber="32">H. southwardorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FF93CB27FAD5813BFA9BFEBE" box="[1328,1393,259,285]" pageId="31" pageNumber="32" rank="species">
|
||
<emphasis id="B92CCF2AFF93CB27FAD5813BFA83FEBE" bold="true" box="[1328,1385,259,285]" pageId="31" pageNumber="32">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
the first peristomial ring is narrow and separated from the prostomium by the nuchal groove, branchiae are present on setigers 2–7, capillary setae are present on setigers 10–11, with acicular spines first present on setigers 12–13, and subuluncini are absent. In contrast,
|
||
<taxonomicName id="4C5868BBFF93CB27FDC38157FD35FE2A" box="[550,735,367,393]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="mediterranea">
|
||
<emphasis id="B92CCF2AFF93CB27FDC38157FD35FE2A" box="[550,735,367,393]" italics="true" pageId="31" pageNumber="32">H. mediterranea</emphasis>
|
||
</taxonomicName>
|
||
has both peristomial rings of an equivalent size, branchiae are present on setigers 2–8, and capillary setae are present on setigers 10–12, with acicular spines and subuluncini first present from setiger 13. See
|
||
<tableCitation id="C6DA2683FF93CB27FE32818FFDC0FE72" box="[471,554,439,465]" captionStart="TABLE 1" captionStartId="61.[151,243,462,488]" captionTargetPageId="61" captionText="TABLE 1. Morphological Characteristics of the Known Species of Heterospio." httpUri="http://table.plazi.org/id/DF2743B0FFB1CB05FF7281F6FC00FE4B" pageId="31" pageNumber="32" tableUuid="DF2743B0FFB1CB05FF7281F6FC00FE4B">Table 1</tableCitation>
|
||
for further comparison of
|
||
<taxonomicName id="4C5868BBFF93CB27FCB1818FFBCCFE72" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[852,1062,439,465]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="southwardorum" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFF93CB27FCB1818FFBCCFE72" box="[852,1062,439,465]" italics="true" pageId="31" pageNumber="32">H. southwardorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FF93CB27FBC8818FFB86FE72" box="[1069,1132,439,465]" pageId="31" pageNumber="32" rank="species">
|
||
<emphasis id="B92CCF2AFF93CB27FBC8818FFB8FFE72" bold="true" box="[1069,1125,439,465]" pageId="31" pageNumber="32">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
with its congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34240B3FF93CB27FF2281E3FC04FD9E" pageId="31" pageNumber="32" type="etymology">
|
||
<paragraph id="8BE71338FF93CB27FF2281E3FC04FD9E" blockId="31.[151,1437,150,609]" pageId="31" pageNumber="32">
|
||
<emphasis id="B92CCF2AFF93CB27FF2281E3FEA2FE56" bold="true" box="[199,328,475,501]" pageId="31" pageNumber="32">Etymology</emphasis>
|
||
. This species is named for the late Drs. Alan and Eve Southward, prominent marine zoologists who directed the R/V
|
||
<taxonomicName id="4C5868BBFF93CB27FEB181C7FE76FDBA" box="[340,412,511,537]" class="Hydrozoa" family="Corynidae" genus="Sarsia" kingdom="Animalia" order="Anthoathecata" pageId="31" pageNumber="32" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B92CCF2AFF93CB27FEB181C7FE76FDBA" box="[340,412,511,537]" italics="true" pageId="31" pageNumber="32">Sarsia</emphasis>
|
||
</taxonomicName>
|
||
cruise in the Bay of Biscay during which this species was collected. Dr. Eve Southward was a deep-sea biologist and an authority on annelids of the family
|
||
<taxonomicName id="4C5868BBFF93CB27FCBC821BFC00FD9E" box="[857,1002,547,573]" class="Polychaeta" family="Siboglinidae" kingdom="Animalia" order="Sabellida" pageId="31" pageNumber="32" phylum="Annelida" rank="family">Siboglinidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34240B3FF93CB27FF22827EFC24FDC2" box="[199,974,582,609]" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="8BE71338FF93CB27FF22827EFC24FDC2" blockId="31.[151,1437,150,609]" box="[199,974,582,609]" pageId="31" pageNumber="32">
|
||
<emphasis id="B92CCF2AFF93CB27FF22827EFEB3FDC3" bold="true" box="[199,345,582,608]" pageId="31" pageNumber="32">Distribution</emphasis>
|
||
. North Atlantic Ocean, Bay of Biscay off
|
||
<collectingCountry id="F34F53A8FF93CB27FCD7827FFC9FFDC2" box="[818,885,583,609]" name="Spain" pageId="31" pageNumber="32">Spain</collectingCountry>
|
||
,
|
||
<specimenCount id="9D5ED8B1FF93CB27FC9A827FFC24FDC2" box="[895,974,583,609]" pageId="31" pageNumber="32" type="generic">
|
||
<quantity id="4CA0BEDDFF93CB27FC9A827FFC23FDC2" box="[895,969,583,609]" metricMagnitude="2" metricUnit="m" metricValue="6.409999999999999" pageId="31" pageNumber="32" unit="m" value="641.0">641 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |