296 lines
34 KiB
XML
296 lines
34 KiB
XML
<document id="D9180549EAFE4CBD98003336F8D3F548" ID-CLB-Dataset="26477" ID-DOI="10.11646/zootaxa.4615.3.8" ID-GBIF-Dataset="9238318c-4409-45cc-96cf-1aa17c4006d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3246211" ID-ZooBank="2AA8C6E7-11D5-4443-8E0E-9FB19856B0CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560509668797" checkinUser="plazi" docAuthor="Mondaca, José, Pizarro-Araya, Jaime & Alfaro, Fermín M." docDate="2019" docId="03E6ED6DA675BF6EFF50B44DE9FE9FF9" docLanguage="en" docName="zootaxa.4615.3.8.pdf" docOrigin="Zootaxa 4615 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luispenaia paposo Mondaca, Pizarro-Araya" docType="treatment" docVersion="7" lastPageNumber="557" masterDocId="FFDF9515A670BF66FFC7B247ED0A9D29" masterDocTitle="Revision of the genus Luispenaia Martínez (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), with description of three new species from the Atacama Desert, Chile" masterLastPageNumber="562" masterPageNumber="549" pageNumber="554" updateTime="1698734911197" updateUser="ExternalLinkService">
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<mods:mods id="EE9F7DD84419D60BF66DBF82F65C64E0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1F7E35E2C36F38ECDA2478378A088AA6">
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<mods:title id="2410CC83ABCD923293C0D920BCCB661E">Revision of the genus Luispenaia Martínez (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), with description of three new species from the Atacama Desert, Chile</mods:title>
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</mods:titleInfo>
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<mods:name id="032EAA1EA1CDFD7B3ACB970FADF79D68" type="personal">
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<mods:role id="ABE898B6CFF3FF620CB0C4E07EDA237F">
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<mods:roleTerm id="5AFACB9744390C62BDC9B22C7C869A2B">Author</mods:roleTerm>
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<mods:namePart id="F2BE888D06C6AA9E63FA3A1F09195F18">Mondaca, José</mods:namePart>
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</mods:name>
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<mods:roleTerm id="4DDD2EBE464EC75E031BE782768F7199">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5A6C7DC2A25C5762AE6E1B50D0AC086D">Pizarro-Araya, Jaime</mods:namePart>
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</mods:name>
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<mods:name id="8064546236C911BCB5E1175CB21443C7" type="personal">
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<mods:roleTerm id="66B3A135DA5224C83AB30706DBC535CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="438061E5E1CA24EF911C0EEC78412D28">Alfaro, Fermín M.</mods:namePart>
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<mods:typeOfResource id="3DC927A7C5AF441BD4AE148B50281916">text</mods:typeOfResource>
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<mods:title id="DBFF2E975F009E043F4C98C04F63F4E3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3F53545F0695230CAF333EAF27D422FE">
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<mods:date id="F6D60C21B5C4184563303AE3C284675B">2019</mods:date>
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<mods:number id="5C7434F9D2A1AD23EB1499EFBE09AA58">2019-06-14</mods:number>
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<mods:number id="510079A3FE77E3EBB49C4966704D5CEB">4615</mods:number>
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<mods:number id="89DDAA4B80F89F033BE3C5B6472D32FA">3</mods:number>
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<mods:classification id="74942D94ABF84600D20C9F28E067B77F">journal article</mods:classification>
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<mods:identifier id="BAB2C670F16A13AEF427A8B8652EE76F" type="DOI">10.11646/zootaxa.4615.3.8</mods:identifier>
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<treatment id="03E6ED6DA675BF6EFF50B44DE9FE9FF9" ID-GBIF-Taxon="158223213" LSID="urn:lsid:plazi:treatment:03E6ED6DA675BF6EFF50B44DE9FE9FF9" httpUri="http://treatment.plazi.org/id/03E6ED6DA675BF6EFF50B44DE9FE9FF9" lastPageId="8" lastPageNumber="557" pageId="5" pageNumber="554">
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<subSubSection id="C3550FF0A675BF63FF50B44DEEDB9B0D" box="[151,977,1546,1573]" pageId="5" pageNumber="554" type="nomenclature">
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<paragraph id="8BF05C7BA675BF63FF50B44DEEDB9B0D" blockId="5.[151,977,1546,1608]" box="[151,977,1546,1573]" pageId="5" pageNumber="554">
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<heading id="D0B8EB17A675BF63FF50B44DEEDB9B0D" bold="true" box="[151,977,1546,1573]" fontSize="11" level="1" pageId="5" pageNumber="554" reason="1">
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<emphasis id="B93B8069A675BF63FF50B44DEEDB9B0D" bold="true" box="[151,977,1546,1573]" pageId="5" pageNumber="554">
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<taxonomicName id="4C4F27F8A675BF63FF50B44DEFB49B0D" authority="Mondaca, Pizarro-Araya" authorityName="Mondaca, Pizarro-Araya" box="[151,702,1546,1573]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="paposo">
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<emphasis id="B93B8069A675BF63FF50B44DEC719B0C" bold="true" box="[151,379,1546,1573]" italics="true" pageId="5" pageNumber="554">Luispenaia paposo</emphasis>
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Mondaca, Pizarro-Araya
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</taxonomicName>
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, & Alfaro new species
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3550FF0A675BF63FF50B469EC8B9B61" box="[151,385,1581,1608]" pageId="5" pageNumber="554" type="description">
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<paragraph id="8BF05C7BA675BF63FF50B469EC8B9B61" blockId="5.[151,977,1546,1608]" box="[151,385,1581,1608]" pageId="5" pageNumber="554">
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(
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<figureCitation id="137440FEA675BF63FF58B469EDF69B61" box="[159,252,1582,1608]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="5" pageNumber="554">Figs. 11</figureCitation>
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,
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<figureCitation id="137440FEA675BF63FECFB46AEC2C9B61" box="[264,294,1581,1608]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="5" pageNumber="554">15</figureCitation>
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,
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<figureCitation id="137440FEA675BF63FEF5B469EC739B61" box="[306,377,1582,1608]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="5" pageNumber="554">21–23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3550FF0A675BF63FF50B431EF4C9A6C" pageId="5" pageNumber="554" type="materials_examined">
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||
<paragraph id="8BF05C7BA675BF63FF50B431EFE79BB8" blockId="5.[151,1437,1653,2041]" box="[151,749,1653,1681]" pageId="5" pageNumber="554">
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<materialsCitation id="3B275626A675BF63FF50B431EFE39BB8" ID-GBIF-Occurrence="2265605695" box="[151,745,1653,1681]" country="Chile" location="Paposo" pageId="5" pageNumber="554" specimenCount="1" stateProvince="Antofagasta" typeStatus="holotype">
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||
<emphasis id="B93B8069A675BF63FF50B431EC3C9BB9" bold="true" box="[151,310,1653,1680]" pageId="5" pageNumber="554">
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||
<typeStatus id="54F4E2D9A675BF63FF50B431EDD89BB9" box="[151,210,1654,1680]" pageId="5" pageNumber="554">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3581CEBA675BF63FEFBB431EC719BB9" box="[316,379,1654,1680]" name="Chile" pageId="5" pageNumber="554">Chile</collectingCountry>
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,
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<collectingRegion id="498B9299A675BF63FE40B431EF839BB9" box="[391,649,1654,1680]" country="Chile" name="Antofagasta" pageId="5" pageNumber="554">Región de Antofagasta</collectingRegion>
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||
,
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||
<location id="8E900AA0A675BF63FD53B430EFE39BB8" LSID="urn:lsid:plazi:treatment:03E6ED6DA675BF6EFF50B44DE9FE9FF9:8E900AA0A675BF63FD53B430EFE39BB8" box="[660,745,1655,1681]" country="Chile" name="Paposo" pageId="5" pageNumber="554" stateProvince="Antofagasta">Paposo</location>
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</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="8BF05C7BA675BF63FF00B4DDEF4C9A6C" blockId="5.[151,1437,1653,2041]" pageId="5" pageNumber="554">
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||
<emphasis id="B93B8069A675BF63FF00B4DDEC7E9B9D" bold="true" box="[199,372,1689,1716]" pageId="5" pageNumber="554">Type material.</emphasis>
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||
<materialsCitation id="3B275626A675BF63FEBCB4DDE9809BD4" ID-GBIF-Occurrence="2412528341" collectingDate="2015-10-30" collectionCode="MNNC" collectorName="J. Pizarro-F. Alfaro" country="Chile" location="Paposo Norte Natural Monument" longitude="70.52698" municipality="Paposo Norte Natural Monument" pageId="5" pageNumber="554" specimenCount="1" specimenCount-male="1" stateProvince="ANTOFAGASTA" typeStatus="holotype">
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||
<typeStatus id="54F4E2D9A675BF63FEBCB4DDECE99B9D" box="[379,483,1690,1716]" pageId="5" pageNumber="554" type="holotype">Holotype</typeStatus>
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||
<specimenCount id="9D4997F2A675BF63FE2EB4DDEF159B9D" box="[489,543,1690,1716]" pageId="5" pageNumber="554" type="male">male</specimenCount>
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at
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<collectionCode id="ED5EC4BEA675BF63FD86B4DDEF9F9B9D" box="[577,661,1690,1716]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="5" pageNumber="554">MNNC</collectionCode>
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||
labeled: a) “
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<collectingCountry id="F3581CEBA675BF63FCE7B4DDEE7F9B9D" box="[800,885,1690,1716]" name="Chile" pageId="5" pageNumber="554">CHILE</collectingCountry>
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<collectingRegion id="498B9299A675BF63FCBCB4DDE94E9B9D" box="[891,1092,1690,1716]" country="Chile" name="Antofagasta" pageId="5" pageNumber="554">ANTOFAGASTA</collectingRegion>
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||
/
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||
<collectingMunicipality id="6B94C601A675BF63FB9FB4DCEDC59BF0" country="Chile" longitude="70.52698" name="Paposo Norte Natural Monument" pageId="5" pageNumber="554" stateProvince="Antofagasta">Paposo Norte Natural Monument</collectingMunicipality>
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(costa), 24°51’ /
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<geoCoordinate id="EE7B3ABCA675BF63FE5BB4FAECFB9BF1" box="[412,497,1725,1752]" pageId="5" pageNumber="554">05.2’’S</geoCoordinate>
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-
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<geoCoordinate id="EE7B3ABCA675BF63FE3EB4FAEF939BF1" box="[505,665,1725,1752]" degrees="70" direction="east" minutes="31" orientation="longitude" pageId="5" pageNumber="554" precision="1" seconds="37.1" value="70.52698">70°31’37.1’’O</geoCoordinate>
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/
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||
<quantity id="4CB7F19EA675BF63FD72B4F9EFF89BF0" box="[693,754,1726,1753]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="554" unit="m" value="43.0">43 m</quantity>
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,
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<collectingDate id="EFB58353A675BF63FCC6B4FAEEAA9BF1" box="[769,928,1725,1752]" pageId="5" pageNumber="554">
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||
27/
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||
<date id="FFF17ABBA675BF63FCE2B4FAEEAA9BF1" box="[805,928,1725,1752]" pageId="5" pageNumber="554" value="2015-10-30">30-X-2015</date>
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||
</collectingDate>
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||
/ col.
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<collectorName id="26BA39ADA675BF63FC29B4F8E9C79BF1" box="[1006,1229,1726,1753]" pageId="5" pageNumber="554">J. Pizarro-F. Alfaro</collectorName>
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” b) “
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<taxonomicName id="4C4F27F8A675BF63FAC9B4F8E8809BF1" authorityName="Martinez" authorityYear="1972" box="[1294,1418,1727,1752]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B93B8069A675BF63FAC9B4F8E8809BF1" box="[1294,1418,1727,1752]" italics="true" pageId="5" pageNumber="554">Luispenaia</emphasis>
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||
</taxonomicName>
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||
/
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||
<taxonomicName id="4C4F27F8A675BF63FF50B4A4EDE29BD5" authority="Mondaca & Pizarro-Araya & Alfaro, 2019" authorityName="Mondaca & Pizarro-Araya & Alfaro" authorityYear="2019" box="[151,232,1763,1788]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="paposo" status="sp. nov.">
|
||
<emphasis id="B93B8069A675BF63FF50B4A4EDE29BD5" box="[151,232,1763,1788]" italics="true" pageId="5" pageNumber="554">paposo</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A2083D12A675BF63FF36B4A5EC479BD5" box="[241,333,1762,1788]" pageId="5" pageNumber="554" rank="species">
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<emphasis id="B93B8069A675BF63FF36B4A5EC479BD5" bold="true" box="[241,333,1762,1788]" pageId="5" pageNumber="554">sp. nov.</emphasis>
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||
</taxonomicNameLabel>
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/ Mondaca, Pizarro-Araya & Alfaro / HOLOTYPE” (red label, typeset)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B275626A675BF63FB51B4A4EF4C9A6C" ID-GBIF-Occurrence="2412528342" collectionCode="JMEC" country="Chile" pageId="5" pageNumber="554" stateProvince="Antofagasta" typeStatus="paratype">
|
||
<collectorName id="26BA39ADA675BF63FB51B4A4E9CD9BD4" box="[1174,1223,1763,1789]" pageId="5" pageNumber="554">Two</collectorName>
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||
male
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||
<typeStatus id="54F4E2D9A675BF63FAD7B4A4E8719BD4" box="[1296,1403,1763,1789]" pageId="5" pageNumber="554" type="paratype">paratypes</typeStatus>
|
||
(1
|
||
<collectionCode id="ED5EC4BEA675BF63FF50B541EDEB9A09" box="[151,225,1798,1824]" pageId="5" pageNumber="554">JMEC</collectionCode>
|
||
, 1
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||
<collectionCode id="ED5EC4BEA675BF63FEC6B541EC509A09" box="[257,346,1798,1824]" pageId="5" pageNumber="554">LEULS</collectionCode>
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||
) labeled as
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||
<typeStatus id="54F4E2D9A675BF63FE1AB541EF359A09" box="[477,575,1798,1824]" pageId="5" pageNumber="554" type="holotype">holotype</typeStatus>
|
||
except: b) “
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||
<taxonomicName id="4C4F27F8A675BF63FD00B540EE499A09" authorityName="Martinez" authorityYear="1972" box="[711,835,1799,1824]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B93B8069A675BF63FD00B540EE499A09" box="[711,835,1799,1824]" italics="true" pageId="5" pageNumber="554">Luispenaia</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C4F27F8A675BF63FC9FB540EEA39A09" authority="Mondaca & Pizarro-Araya & Alfaro, 2019" authorityName="Mondaca & Pizarro-Araya & Alfaro" authorityYear="2019" box="[856,937,1799,1824]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="paposo" status="sp. nov.">
|
||
<emphasis id="B93B8069A675BF63FC9FB540EEA39A09" box="[856,937,1799,1824]" italics="true" pageId="5" pageNumber="554">paposo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2083D12A675BF63FC77B541E9039A09" box="[944,1033,1798,1824]" pageId="5" pageNumber="554" rank="species">
|
||
<emphasis id="B93B8069A675BF63FC77B541E9039A09" bold="true" box="[944,1033,1798,1824]" pageId="5" pageNumber="554">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
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||
/ Mondaca, Pizarro-Araya & Alfaro /
|
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<typeStatus id="54F4E2D9A675BF63FF61B56DEC3F9A6D" box="[166,309,1834,1860]" pageId="5" pageNumber="554" type="paratype">PARATYPE</typeStatus>
|
||
” (yellow label, typeset).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3550FF0A675BF63FF00B50AEE3A9AD1" pageId="5" pageNumber="554" type="diagnosis">
|
||
<paragraph id="8BF05C7BA675BF63FF00B50AEE3A9AD1" blockId="5.[151,1437,1653,2041]" pageId="5" pageNumber="554">
|
||
<emphasis id="B93B8069A675BF63FF00B50AEC489A41" bold="true" box="[199,322,1869,1896]" pageId="5" pageNumber="554">Diagnosis.</emphasis>
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This species is distinguished from all other species in the genus
|
||
<taxonomicName id="4C4F27F8A675BF63FBDAB508E9939A41" authorityName="Martinez" authorityYear="1972" box="[1053,1177,1871,1896]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93B8069A675BF63FBDAB508E9939A41" box="[1053,1177,1871,1896]" italics="true" pageId="5" pageNumber="554">Luispenaia</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: body dorsally black shiny, covered with white and yellowish-white setae (
|
||
<figureCitation id="137440FEA675BF63FB1FB535E8269AA5" box="[1240,1324,1906,1932]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="5" pageNumber="554">Fig. 21</figureCitation>
|
||
). Clypeus distal and laterally rounded, reflexed upturned; lateral margin between clypeal base and ocular canthus rounded, slightly projected externally (
|
||
<figureCitation id="137440FEA675BF63FE25B5FDEF3E9AFD" box="[482,564,1978,2004]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="5" pageNumber="554">Fig. 11</figureCitation>
|
||
). Antennae with 8 antennomeres; club with 4 antennomeres (
|
||
<figureCitation id="137440FEA675BF63FB20B5FDE8319AFD" box="[1255,1339,1977,2004]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="5" pageNumber="554">Fig. 15</figureCitation>
|
||
). Pronotum suboval, broadest in middle, laterally arcuate (
|
||
<figureCitation id="137440FEA675BF63FD17B599EE289AD1" box="[720,802,2014,2040]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="5" pageNumber="554">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF300CF3A676BF60FF50B58BE9099ACC" box="[151,1027,1996,2021]" httpUri="https://zenodo.org/record/3246215/files/figure.png" pageId="6" pageNumber="555" startId="6.[151,250,1996,2021]" targetBox="[192,1388,211,1949]" targetPageId="6">
|
||
<paragraph id="8BF05C7BA676BF60FF50B58BE9099ACC" blockId="6.[151,1027,1996,2021]" box="[151,1027,1996,2021]" pageId="6" pageNumber="555">
|
||
<emphasis id="B93B8069A676BF60FF50B58BEC199ACC" bold="true" box="[151,275,1996,2021]" pageId="6" pageNumber="555">FIGURE 8.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C4F27F8A676BF60FE71B58AEF2D9ACD" authorityName="Martinez" authorityYear="1972" box="[438,551,1997,2020]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="555" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93B8069A676BF60FE71B58AEF2D9ACD" box="[438,551,1997,2020]" italics="true" pageId="6" pageNumber="555">Luispenaia</emphasis>
|
||
</taxonomicName>
|
||
species in the Atacama Desert, northern Chile.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF300CF3A677BF61FF50B7A3EE9198D5" box="[151,923,1508,1533]" httpUri="https://zenodo.org/record/3246218/files/figure.png" pageId="7" pageNumber="556" startId="7.[151,250,1508,1533]" targetBox="[153,1434,212,1496]" targetPageId="7">
|
||
<paragraph id="8BF05C7BA677BF61FF50B7A3EE9198D5" blockId="7.[151,923,1508,1533]" box="[151,923,1508,1533]" pageId="7" pageNumber="556">
|
||
<emphasis id="B93B8069A677BF61FF50B7A3EC1998D4" bold="true" box="[151,275,1508,1533]" pageId="7" pageNumber="556">FIGURE 9.</emphasis>
|
||
Male holotype of
|
||
<taxonomicName id="4C4F27F8A677BF61FE09B7A2EE5A98D4" authority="Martinez. Scale" authorityName="Martinez. Scale" box="[462,848,1508,1533]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="556" phylum="Arthropoda" rank="species" species="paradoxa">
|
||
<emphasis id="B93B8069A677BF61FE09B7A2EFAC98D4" box="[462,678,1509,1533]" italics="true" pageId="7" pageNumber="556">Luispenaia paradoxa</emphasis>
|
||
Martínez. Scale
|
||
</taxonomicName>
|
||
: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3550FF0A677BF6EFF00B461EF549C86" lastPageId="8" lastPageNumber="557" pageId="7" pageNumber="556" type="description">
|
||
<paragraph id="8BF05C7BA677BF6EFF00B461EF3F9C6A" blockId="7.[151,1437,1574,2034]" lastBlockId="8.[151,1437,153,720]" lastPageId="8" lastPageNumber="557" pageId="7" pageNumber="556">
|
||
<emphasis id="B93B8069A677BF61FF00B461EF2F9B68" bold="true" box="[199,549,1574,1601]" pageId="7" pageNumber="556">
|
||
Description of male
|
||
<typeStatus id="54F4E2D9A677BF61FE79B460EF2F9B68" box="[446,549,1575,1601]" pageId="7" pageNumber="556" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Length 6.0 mm, wide 2.3.
|
||
<emphasis id="B93B8069A677BF61FCAEB460EEA99B68" box="[873,931,1575,1601]" italics="true" pageId="7" pageNumber="556">Body</emphasis>
|
||
: Slightly convex, dorsally black shiny, with white and yellowish-white setae; setae long, moderately dense on head, pronotum, and elytra (
|
||
<figureCitation id="137440FEA677BF61FB6BB40CE9F49B4C" box="[1196,1278,1611,1637]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="7" pageNumber="556">Fig. 21</figureCitation>
|
||
).
|
||
<emphasis id="B93B8069A677BF61FAD5B40CE8459B4C" box="[1298,1359,1611,1637]" italics="true" pageId="7" pageNumber="556">Head</emphasis>
|
||
: Clypeus distal and laterally rounded, reflexed upturned; dorsal surface moderately punctate and setose; punctures large; frontoclypeal suture well defined, frontal width equal to 7 times eye widths, with surface densely punctate, punc- tures large; eye canthus short and wide, rounded externally (
|
||
<figureCitation id="137440FEA677BF61FCE8B4F0EE8A9BF8" box="[815,896,1719,1745]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="7" pageNumber="556">Fig. 11</figureCitation>
|
||
). Antennae with 8 antennomeres; antennomere 1 robust, claviform; antennomere 2 submonoliform; antennomere 3 subcylindrical; antennomere 4 flattened; antennal club with 4 antennomeres glabrous, subequal in length, arched in dorsal view (
|
||
<figureCitation id="137440FEA677BF61FBC6B4B8E95E9A30" box="[1025,1108,1790,1817]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="7" pageNumber="556">Fig. 15</figureCitation>
|
||
).
|
||
<emphasis id="B93B8069A677BF61FBAFB547E9DD9A30" box="[1128,1239,1792,1817]" italics="true" pageId="7" pageNumber="556">Pronotum</emphasis>
|
||
: Slightly convex, approximately suboval, broadest in middle, laterally arcuate; surface densely punctate and setose; setae recumbent, directed posteriorly; longitudinal midline impunctate, lateral margin with long, dense, laterally directed setae (
|
||
<figureCitation id="137440FEA677BF61FAA8B500EDBE9AAC" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="7" pageNumber="556">Fig. 11</figureCitation>
|
||
).
|
||
<emphasis id="B93B8069A677BF61FF0BB52CEC309AAC" box="[204,314,1899,1925]" italics="true" pageId="7" pageNumber="556">Scutellum</emphasis>
|
||
: Triangular, longer than wide; sides broadly arcuate.
|
||
<emphasis id="B93B8069A677BF61FC6FB52CEEE49AAC" box="[936,1006,1899,1925]" italics="true" pageId="7" pageNumber="556">Elytra</emphasis>
|
||
: Slightly convex, surface moderately setose; setae recumbent, directed posteriorly, with striae poorly defined; lateral margin with long, moderately dense, laterally directed setae.
|
||
<emphasis id="B93B8069A677BF61FE59B5F3ECD99AE4" box="[414,467,1972,1997]" italics="true" pageId="7" pageNumber="556">Legs</emphasis>
|
||
: Moderately setose and shiny, with white and yellowish-white, long setae. Protibiae tridentate, with teeth acute distally; protarsomere 5 longer than protarsomeres 3–4 combined; mesotibiae and metati- biae widened medial and apically, with one oblique spinose carina externally, apical edge of the mesotibiae with row of long equal macrosetae, metatibiae with row of very small equal macrosetae barely perceptible; mesotarsomere 5 subequal in length to mesotarsomeres 3–4 combined.
|
||
<emphasis id="B93B8069A678BF6EFD34B2A6EE579DD2" box="[755,861,225,251]" italics="true" pageId="8" pageNumber="557">Genitalia</emphasis>
|
||
: Parameres moderately long, wider at base (frontal view), laterally cuneiform, slightly curved ventrally, distally attenuate and rounded at apex; phallobase 1.04 times longer than parameres (
|
||
<figureCitation id="137440FEA678BF6EFE58B36EEF209C6A" box="[415,554,297,323]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="8" pageNumber="557">Figs. 22–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF05C7BA678BF6EFF00B30AECA89C4E" blockId="8.[151,1437,153,720]" box="[199,418,333,359]" pageId="8" pageNumber="557">
|
||
<emphasis id="B93B8069A678BF6EFF00B30AEC2F9C4E" bold="true" box="[199,293,333,359]" pageId="8" pageNumber="557">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BF05C7BA678BF6EFF00B337EF549C86" blockId="8.[151,1437,153,720]" pageId="8" pageNumber="557">
|
||
<emphasis id="B93B8069A678BF6EFF00B337EC359CA2" bold="true" box="[199,319,368,395]" pageId="8" pageNumber="557">Variation.</emphasis>
|
||
Length 5.9–6.0 mm. Dorsal color light brown, dark brown or black shiny. Not differing significantly from the
|
||
<typeStatus id="54F4E2D9A678BF6EFF39B3D2EC6A9C86" box="[254,352,405,431]" pageId="8" pageNumber="557" type="holotype">holotype</typeStatus>
|
||
in external characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3550FF0A678BF6EFF00B3FEE9E19CFA" box="[199,1259,441,467]" pageId="8" pageNumber="557" type="etymology">
|
||
<paragraph id="8BF05C7BA678BF6EFF00B3FEE9E19CFA" blockId="8.[151,1437,153,720]" box="[199,1259,441,467]" pageId="8" pageNumber="557">
|
||
<emphasis id="B93B8069A678BF6EFF00B3FEEC449CFA" bold="true" box="[199,334,441,467]" pageId="8" pageNumber="557">Etymology.</emphasis>
|
||
Derived from Paposo, origin locality of the new species located in northern
|
||
<collectingCountry id="F3581CEBA678BF6EFB60B3FEE9ED9CFA" box="[1191,1255,441,467]" name="Chile" pageId="8" pageNumber="557">Chile</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3550FF0A678BF6EFF00B39BE9FE9FF9" pageId="8" pageNumber="557" type="distribution">
|
||
<paragraph id="8BF05C7BA678BF6EFF00B39BE97E9CDE" blockId="8.[151,1437,153,720]" box="[199,1140,476,503]" pageId="8" pageNumber="557">
|
||
<emphasis id="B93B8069A678BF6EFF00B39BEC6B9CDE" bold="true" box="[199,353,476,503]" pageId="8" pageNumber="557">Distribution.</emphasis>
|
||
Paposo Norte Natural Monument,
|
||
<collectingRegion id="498B9299A678BF6EFD2FB39AEEC19CDE" box="[744,971,477,503]" country="Chile" name="Antofagasta" pageId="8" pageNumber="557">Antofagasta Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3581CEBA678BF6EFC11B39AE9199CDE" box="[982,1043,477,503]" name="Chile" pageId="8" pageNumber="557">Chile</collectingCountry>
|
||
(
|
||
<figureCitation id="137440FEA678BF6EFBE5B39AE96C9CDE" box="[1058,1126,477,503]" captionStart="FIGURE 8" captionStartId="6.[151,250,1996,2021]" captionTargetBox="[192,1388,211,1949]" captionTargetId="figure@6.[151,1435,170,1984]" captionTargetPageId="6" captionText="FIGURE 8. Distribution of Luispenaia species in the Atacama Desert, northern Chile." httpUri="https://zenodo.org/record/3246215/files/figure.png" pageId="8" pageNumber="557">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF05C7BA678BF6EFF00B047E9FE9FF9" blockId="8.[151,1437,153,720]" pageId="8" pageNumber="557">
|
||
<emphasis id="B93B8069A678BF6EFF00B047ECB59F32" bold="true" box="[199,447,512,539]" pageId="8" pageNumber="557">Habitat and ecology.</emphasis>
|
||
<taxonomicName id="4C4F27F8A678BF6EFE00B045EF979F32" authorityName="Mondaca & Pizarro-Araya & Alfaro" authorityYear="2019" box="[455,669,514,539]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="557" phylum="Arthropoda" rank="species" species="paposo">
|
||
<emphasis id="B93B8069A678BF6EFE00B045EF979F32" box="[455,669,514,539]" italics="true" pageId="8" pageNumber="557">Luispenaia paposo</emphasis>
|
||
</taxonomicName>
|
||
has been collected during spring by using pitfall trap in the coastal dunes in Paposo Natural Monument (northern
|
||
<collectingCountry id="F3581CEBA678BF6EFD69B062EFE59F69" box="[686,751,549,576]" name="Chile" pageId="8" pageNumber="557">Chile</collectingCountry>
|
||
), in the few places of the area that harbor some vegetation, which is reduced to some scattered plants adapted to psammophilous environments (
|
||
<figureCitation id="137440FEA678BF6EFB85B00EE9A29F4A" box="[1090,1192,585,611]" captionStart="FIGURES 30" captionStartId="11.[151,264,1675,1700]" captionTargetBox="[151,1435,801,1663]" captionTargetId="figure@11.[151,1435,801,1663]" captionTargetPageId="11" captionText="FIGURES 30. Type locality of Luispenaia species. A, Llanos de Challe National Park (Atacama Region, Chile), habitat of L. paradoxa; B, Boqueron Chañar (Atacama Region, Chile), habitat of L. atacamensis; C, Paposo Norte Natural Monument (Antofagasta Region, Chile), habitat of L. paposo; D, Quebrada Los Yales (Antofagasta Region, Chile), habitat of L. paulseni." httpUri="https://zenodo.org/record/3246226/files/figure.png" pageId="8" pageNumber="557">Fig. 30C</figureCitation>
|
||
). In this area there are two relatively abundant species that correspond to the dominant association;
|
||
<taxonomicName id="4C4F27F8A678BF6EFC2BB02AE8029FAE" authority="Meyen" authorityName="Meyen" authorityYear="1834" box="[1004,1288,621,647]" class="Magnoliopsida" family="Apocynaceae" genus="Skytanthus" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="557" phylum="Tracheophyta" rank="species" species="acutus">
|
||
<emphasis id="B93B8069A678BF6EFC2BB02AE9BE9FAE" box="[1004,1204,621,647]" italics="true" pageId="8" pageNumber="557">Skytanthus acutus</emphasis>
|
||
Meyen
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4F27F8A678BF6EFAFAB029ECDE9F82" authority="Barneoud. The" authorityName="Barneoud. The" authorityYear="1847" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Animalia" order="Caryophyllales" pageId="8" pageNumber="557" phylum="Tracheophyta" rank="species" species="maritima">
|
||
<emphasis id="B93B8069A678BF6EFAFAB029EC229F82" italics="true" pageId="8" pageNumber="557">Tetragonia maritima</emphasis>
|
||
Barnéoud. The
|
||
</taxonomicName>
|
||
vegetational floor that takes place in this area is the coastal Mediterranean desert scrub of
|
||
<taxonomicName id="4C4F27F8A678BF6EFF71B0F2EF129FE6" authority="Phil." authorityName="Phil." box="[182,536,693,719]" class="Magnoliopsida" family="Asteraceae" genus="Gypothamnium" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="557" phylum="Tracheophyta" rank="species" species="pinifolium">
|
||
<emphasis id="B93B8069A678BF6EFF71B0F2ECD69FE6" box="[182,476,693,719]" italics="true" pageId="8" pageNumber="557">Gypothamnium pinifolium</emphasis>
|
||
Phil.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4F27F8A678BF6EFD89B0F2E9FA9FF9" authority="Phil. (Luebert & Pliscoff 2006)" authorityName="Phil. (Luebert & Pliscoff" authorityYear="2006" box="[590,1264,693,720]" class="Magnoliopsida" family="Boraginaceae" genus="Heliotropium" kingdom="Plantae" order="Boraginales" pageId="8" pageNumber="557" phylum="Tracheophyta" rank="species" species="pycnophyllum">
|
||
<emphasis id="B93B8069A678BF6EFD89B0F2EE829FE6" box="[590,904,693,719]" italics="true" pageId="8" pageNumber="557">Heliotropium pycnophyllum</emphasis>
|
||
Phil. (
|
||
<bibRefCitation id="EFDE218AA678BF6EFC16B0F2E9E39FF9" author="Luebert, F. & Pliscoff, P." box="[977,1257,693,720]" pageId="8" pageNumber="557" refId="ref7924" refString="Luebert, F. & Pliscoff, P. (2006) Sinopsis Bioclimatica y Vegetacional de Chile. Editorial Universitaria, Santiago, 316 pp." type="book" year="2006">Luebert & Pliscoff 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |