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<document id="69FF1FDBE4CD2E1C586C41C8E7C610B2" ID-DOI="10.11646/zootaxa.5489.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13211257" ID-ZooBank="5AAD9922-550B-4B65-AA9F-C2A10949D372" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="felipe" checkinTime="1722790157296" checkinUser="plazi" docAuthor="Ulitzka, Manfred R." docDate="2024" docId="03D2F065A6290476FF261178FDFBFEF7" docLanguage="en" docName="zootaxa.5489.1.6.pdf" docOrigin="Zootaxa 5489 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5489.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campanithrips kyakhei Ulitzka, 2024, gen. et sp. n." docType="treatment" docVersion="4" lastPageNumber="104" masterDocId="FFEB881DA62D0473FFB11266FFB6FF92" masterDocTitle="Late Cretaceous thrips (Thysanoptera) from Hti Lin amber" masterLastPageNumber="106" masterPageNumber="99" pageNumber="103" updateTime="1726016948364" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="6A2337419C15FA05B396F9F8A2D6488A">Late Cretaceous thrips (Thysanoptera) from Hti Lin amber</mods:title>
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<mods:namePart id="B5B55F80D359C2BBC156377229534D92">Ulitzka, Manfred R.</mods:namePart>
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<mods:date id="8C939DCDC35D402475305DC57E021A11">2024</mods:date>
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<mods:number id="C9DA3F23B72477B720182369F89A4A9C">2024-07-31</mods:number>
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<mods:identifier id="89CED17EA2D20468D68A035696F31506" type="DOI">10.11646/zootaxa.5489.1.6</mods:identifier>
<mods:identifier id="FDFF2C9A319D5C772C65E0A4B51A5E0A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="05B68FA423827F16859DF54BE4540F18" type="Zenodo-Dep">13211257</mods:identifier>
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<treatment id="03D2F065A6290476FF261178FDFBFEF7" ID-DOI="http://doi.org/10.5281/zenodo.13212260" ID-Zenodo-Dep="13212260" LSID="urn:lsid:plazi:treatment:03D2F065A6290476FF261178FDFBFEF7" httpUri="http://treatment.plazi.org/id/03D2F065A6290476FF261178FDFBFEF7" lastPageId="5" lastPageNumber="104" pageId="4" pageNumber="103">
<subSubSection id="C36112F8A6290477FF261178FDE2FCAB" box="[151,596,798,825]" pageId="4" pageNumber="103" type="nomenclature">
<paragraph id="8BC44173A6290477FF261178FDE2FCAB" blockId="4.[151,596,798,861]" box="[151,596,798,825]" pageId="4" pageNumber="103">
<heading id="D08CF61FA6290477FF261178FDE2FCAB" bold="true" box="[151,596,798,825]" fontSize="11" level="1" pageId="4" pageNumber="103" reason="1">
<emphasis id="B90F9D61A6290477FF261178FDE2FCAB" bold="true" box="[151,596,798,825]" pageId="4" pageNumber="103">
<taxonomicName id="4C7B3AF0A6290477FF261178FE04FCAB" authorityName="Ulitzka" authorityYear="2024" box="[151,434,798,825]" class="Insecta" family="Stenurothripidae" genus="Campanithrips" kingdom="Animalia" order="Thysanoptera" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="kyakhei" status="gen. et sp. n.">
<emphasis id="B90F9D61A6290477FF261178FE04FCAB" bold="true" box="[151,434,798,825]" italics="true" pageId="4" pageNumber="103">Campanithrips kyakhei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C201AA6290477FE081179FDE2FCAB" box="[441,596,799,825]" pageId="4" pageNumber="103" rank="species">gen. et sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36112F8A6290477FF261125FD86F807" pageId="4" pageNumber="103" type="description">
<paragraph id="8BC44173A6290477FF261125FEBDFCCF" blockId="4.[151,596,798,861]" box="[151,267,835,861]" pageId="4" pageNumber="103">
(
<figureCitation id="13405DF6A6290477FF2E1125FEB5FCCF" box="[159,259,835,861]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1674,1698]" captionTargetBox="[151,1435,705,1650]" captionTargetId="figure-241@3.[151,1435,705,1650]" captionTargetPageId="3" captionText="FIGURES 1–3. New Terebrantia from Hti Lin amber. (1) Avitrips yellae gen. et sp. n. holotype female (MU-Fos-128/1), lateral view. (2–3) Campanithrips kyakhei gen. et sp. n. holotype female (MU-Fos-129/1); (2) dorsal view, encircled red: pollen grain attached to the antenna of the thrips; (3) left fore wing." figureDoi="http://doi.org/10.5281/zenodo.13211259" httpUri="https://zenodo.org/record/13211259/files/figure.png" pageId="4" pageNumber="103">Figs 2–3</figureCitation>
)
</paragraph>
<paragraph id="8BC44173A6290477FF2611EDFB23FB83" blockId="4.[151,1437,907,2013]" pageId="4" pageNumber="103">
<emphasis id="B90F9D61A6290477FF2611EDFF5DFC37" box="[151,235,907,933]" italics="true" pageId="4" pageNumber="103">Female</emphasis>
. Preserved in dorso-ventral position. Body fully distended; wings slightly spread apart from body, laterally overlapping abdomen on the right side. Hind legs and left mid leg spread, others bent under body or head, respectively. Posterior right side of head and thorax largely hidden by opaque streaks and bubbles. Left antenna bent inwards; most antennal segments permeated by air inclusions. Median parts of abdomen translucent.
</paragraph>
<paragraph id="8BC44173A6290477FF76167DFCEBFBCB" blockId="4.[151,1437,907,2013]" pageId="4" pageNumber="103">Colour. Body, legs and antennae uniformly yellowish brown; major body setae pale, fringes and wing setae dark; wings shaded, veins slightly darker than wing membrane.</paragraph>
<paragraph id="8BC44173A6290477FF761605FC9EF92F" blockId="4.[151,1437,907,2013]" pageId="4" pageNumber="103">
Head (
<figureCitation id="13405DF6A6290477FEA21605FEEEFBEF" box="[275,344,1123,1149]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1674,1698]" captionTargetBox="[151,1435,705,1650]" captionTargetId="figure-241@3.[151,1435,705,1650]" captionTargetPageId="3" captionText="FIGURES 1–3. New Terebrantia from Hti Lin amber. (1) Avitrips yellae gen. et sp. n. holotype female (MU-Fos-128/1), lateral view. (2–3) Campanithrips kyakhei gen. et sp. n. holotype female (MU-Fos-129/1); (2) dorsal view, encircled red: pollen grain attached to the antenna of the thrips; (3) left fore wing." figureDoi="http://doi.org/10.5281/zenodo.13211259" httpUri="https://zenodo.org/record/13211259/files/figure.png" pageId="4" pageNumber="103">Fig. 2</figureCitation>
) about as long as wide, with compound eyes largely protruding anteriorly. Sculpture, ocelli and head setae not assessable. Mouth cone short and rounded. Antennae 9-segmented; segment I slightly conical; II basally with a short but rather strong pedicle, then reverse conically widened and almost parallel in the middle, apically with a narrow, recessed ring; III with a pedicle at base, then inverted conical, distally asymmetrical due to the attachment of a large exterior sense cone. IV–VI (including VII on the left) decreasing in diameter, rounded, each much longer than wide. Sense cone on IV not assessable, only its area of attachment visible; V–VII with long setae; VIII inverse conical; XI style-shaped. Pronotum wider than long; pronotal chaetotaxy as well as pterothoracic structure not assessable. Fore wings (
<figureCitation id="13405DF6A6290477FD8B1739FDCAFAEB" box="[570,636,1375,1401]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1674,1698]" captionTargetBox="[151,1435,705,1650]" captionTargetId="figure-241@3.[151,1435,705,1650]" captionTargetPageId="3" captionText="FIGURES 1–3. New Terebrantia from Hti Lin amber. (1) Avitrips yellae gen. et sp. n. holotype female (MU-Fos-128/1), lateral view. (2–3) Campanithrips kyakhei gen. et sp. n. holotype female (MU-Fos-129/1); (2) dorsal view, encircled red: pollen grain attached to the antenna of the thrips; (3) left fore wing." figureDoi="http://doi.org/10.5281/zenodo.13211259" httpUri="https://zenodo.org/record/13211259/files/figure.png" pageId="4" pageNumber="103">Fig. 3</figureCitation>
) narrow but somewhat widened in basal third, distally from the middle slightly bent forwards, tip softly pointed; with two cross veins (RP1 not visible or not developed; see
<bibRefCitation id="EFEA3C82A6290477FA8117E5FF62FA53" author="Nel, P. &amp; Azar, D. &amp; Prokop, J. &amp; Roques, P. &amp; Hodebert, G. &amp; Nel, A." pageId="4" pageNumber="103" pagination="385 - 399" refId="ref5287" refString="Nel, P., Azar, D., Prokop, J., Roques, P., Hodebert, G. &amp; Nel, A. (2012) From Carboniferous to Recent: wing venation enlightens evolution of thysanopteran lineage. Journal of Systematic Palaeontology, 10 (2), 385 - 399. https: // doi. org / 10.1080 / 14772019.2011.598578" type="journal article" year="2012">
Nel
<emphasis id="B90F9D61A6290477FAD317E5FA2BFA0F" box="[1378,1437,1411,1437]" italics="true" pageId="4" pageNumber="103">et al.</emphasis>
2012
</bibRefCitation>
); first vein with a complete row of 12, second vein with 8 setae, both veins offset by a gap between the last and the penultimate, giving the impression of one missing seta; costa with a row of 18 long setae; wing fringes straight; wing membrane with microtrichia; clavus not assessable. Hind wing (
<figureCitation id="13405DF6A6290477FC751789FBBCF99B" box="[964,1034,1519,1545]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1674,1698]" captionTargetBox="[151,1435,705,1650]" captionTargetId="figure-241@3.[151,1435,705,1650]" captionTargetPageId="3" captionText="FIGURES 1–3. New Terebrantia from Hti Lin amber. (1) Avitrips yellae gen. et sp. n. holotype female (MU-Fos-128/1), lateral view. (2–3) Campanithrips kyakhei gen. et sp. n. holotype female (MU-Fos-129/1); (2) dorsal view, encircled red: pollen grain attached to the antenna of the thrips; (3) left fore wing." figureDoi="http://doi.org/10.5281/zenodo.13211259" httpUri="https://zenodo.org/record/13211259/files/figure.png" pageId="4" pageNumber="103">Fig. 3</figureCitation>
) setae straight; membrane without microtrichia. Fore legs with femora not enlarged; femora and tibiae with curved setae on outer edge; fore tarsi not clearly assessable, however, with hamus. Mid legs slender; femora with 3, tibiae with 4 long ventral setae; tarsi two-segmented. Hind legs slender; tibiae with a row of 3 setae and a pair of distal spines; tarsi two-segmented. Abdomen not assessable in terms of sculpture and chaetotaxy; segments VIII and IX cone-shaped, X distinctly prolonged; distal abdominal setae long and pointed; ovipositor strong.
</paragraph>
<paragraph id="8BC44173A6290477FF7614A1FD86F807" blockId="4.[151,1437,907,2013]" pageId="4" pageNumber="103">
<emphasis id="B90F9D61A6290477FF7614A1FEC8F973" bold="true" box="[199,382,1735,1761]" pageId="4" pageNumber="103">Measurements.</emphasis>
Female (in microns): Body, length 665. Head, length 80; largest width 100. Eyes, length 55; width about 35. Pronotum, length and setae not measurable; width 110. Pterothorax not measurable. Abdomen, length 405; largest width 145 (segment V); segment X length 40. Setae on tergite IX,
<geoCoordinate id="EE4F27B4A6290477FBE11569FB38F8BB" box="[1104,1166,1806,1833]" degrees="1" direction="south" minutes="45" orientation="latitude" pageId="4" pageNumber="103" precision="925" value="-1.75">s1 45</geoCoordinate>
,
<geoCoordinate id="EE4F27B4A6290477FB281569FB61F8BB" box="[1177,1239,1807,1833]" degrees="2" direction="south" minutes="30" orientation="latitude" pageId="4" pageNumber="103" precision="925" value="-2.5">s2 30</geoCoordinate>
,
<geoCoordinate id="EE4F27B4A6290477FB531569FA97F8BB" box="[1250,1313,1806,1833]" degrees="3" direction="south" minutes="55" orientation="latitude" pageId="4" pageNumber="103" precision="925" value="-3.9166667">s3 55</geoCoordinate>
. Antennae, length 180; length (largest width) of segment I 15 (15), II 35 (15), III 25 (10), IV 25 (10), V 20 (10), VI 20 (10), VII 15 (10), VIII 15 (5), IX 15 (5). Sense cones (segment III), length 5; basal width 5. Fore wings, length 455; width in the middle 35; costal setae about 30.
</paragraph>
</subSubSection>
<subSubSection id="C36112F8A6290476FF7615F8FD32FE8F" lastPageId="5" lastPageNumber="104" pageId="4" pageNumber="103" type="materials_examined">
<paragraph id="8BC44173A6290477FF7615F8FBF2F84F" blockId="4.[151,1437,907,2013]" pageId="4" pageNumber="103">
<emphasis id="B90F9D61A6290477FF7615F8FE21F82A" bold="true" box="[199,407,1950,1976]" pageId="4" pageNumber="103">Material studied.</emphasis>
<materialsCitation id="3B134B2EA6290477FE2E15F9FBF2F84F" country="Myanmar" location="Magway Region" pageId="4" pageNumber="103" specimenCode="MU-Fos-129/1" specimenCount="1" stateProvince="Magway Region" typeStatus="holotype">
<typeStatus id="54C0FFD1A6290477FE2E15F9FDB1F82B" box="[415,519,1951,1977]" pageId="4" pageNumber="103" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7D8AFAA6290477FDA015F9FDEBF82B" box="[529,605,1951,1977]" pageId="4" pageNumber="103" type="female">female</specimenCount>
<specimenCode id="DBDDE908A6290477FDD715F9FCA7F82B" box="[614,785,1951,1977]" pageId="4" pageNumber="103">MU-Fos-129/1</specimenCode>
; inclusion in Hti Lin amber,
<collectingRegion id="49BF8F91A6290477FBEF15F9FAABF82B" box="[1118,1309,1951,1977]" country="Myanmar" name="Magway" pageId="4" pageNumber="103">Magway Region</collectingRegion>
,
<collectingCountry id="F36C01E3A6290477FA9B15F9FA2FF82B" box="[1322,1433,1951,1977]" name="Myanmar" pageId="4" pageNumber="103">Myanmar</collectingCountry>
. The holotype was donated by Patrick Müller and is stored in the author’s collection.
</materialsCitation>
</paragraph>
<paragraph id="8BC44173A6280476FF7612F0FD32FE8F" blockId="5.[151,1436,150,357]" pageId="5" pageNumber="104">
<emphasis id="B90F9D61A6280476FF7612F0FEC4FF22" bold="true" box="[199,370,150,176]" pageId="5" pageNumber="104">Syninclusions.</emphasis>
Parts of an insect leg, some small brownish droplets, air bubbles, brownish stains of unknown origin, numerous tiny crystal-like particles and a pollen grain (
<figureCitation id="13405DF6A6280476FCD812DDFC1BFF47" box="[873,941,187,213]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1674,1698]" captionTargetBox="[151,1435,705,1650]" captionTargetId="figure-241@3.[151,1435,705,1650]" captionTargetPageId="3" captionText="FIGURES 1–3. New Terebrantia from Hti Lin amber. (1) Avitrips yellae gen. et sp. n. holotype female (MU-Fos-128/1), lateral view. (2–3) Campanithrips kyakhei gen. et sp. n. holotype female (MU-Fos-129/1); (2) dorsal view, encircled red: pollen grain attached to the antenna of the thrips; (3) left fore wing." figureDoi="http://doi.org/10.5281/zenodo.13211259" httpUri="https://zenodo.org/record/13211259/files/figure.png" pageId="5" pageNumber="104">Fig. 3</figureCitation>
) attached to the right antenna of the thrips inclusion (similar in shape to pollen of extant
<taxonomicName id="4C7B3AF0A6280476FD1512B9FD5EFF6B" box="[676,744,223,249]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="104" phylum="Tracheophyta" rank="genus">
<emphasis id="B90F9D61A6280476FD1512B9FD5EFF6B" box="[676,744,223,249]" italics="true" pageId="5" pageNumber="104">Cycas</emphasis>
</taxonomicName>
or respectively
<taxonomicName id="4C7B3AF0A6280476FC2E12B9FB9EFF6B" box="[927,1064,223,249]" class="Cycadopsida" family="Cycadaceae" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="104" phylum="Tracheophyta" rank="family">Cycadaceae</taxonomicName>
; length 22, width 9 microns (see
<bibRefCitation id="EFEA3C82A6280476FF261365FE26FE8F" author="APSA Members" box="[151,400,259,285]" pageId="5" pageNumber="104" refId="ref4430" refString="APSA Members (2007) The Australasian Pollen and Spore Atlas. Version 1.0. Australian National University, Canberra. Available from: https: // apsa. anu. edu. au / samples / sample. php? id = 301 - 4 - 4 (accessed 29 December 2023)" type="url" year="2007">APSA Members 2007</bibRefCitation>
;
<bibRefCitation id="EFEA3C82A6280476FE2A1365FDC1FE8F" author="Vovides, A. P. &amp; Guevara, R. &amp; Coiro, M. &amp; Galicia, S. &amp; Iglesias, C." box="[411,631,259,285]" pageId="5" pageNumber="104" pagination="182 - 197" refId="ref6791" refString="Vovides, A. P., Guevara, R., Coiro, M., Galicia, S. &amp; Iglesias, C. (2021) Pollen morphology of the Megamexican cycads reveals the potential of morphometrics to identify cycad genera. Botanical Sciences, 99 (1), 182 - 197. https: // doi. org / 10.17129 / botsci. 2696" type="journal article" year="2021">
Vovides
<emphasis id="B90F9D61A6280476FE4D1365FD82FE8F" box="[508,564,259,285]" italics="true" pageId="5" pageNumber="104">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36112F8A6280476FF761341FDFBFEF7" pageId="5" pageNumber="104" type="etymology">
<paragraph id="8BC44173A6280476FF761341FDFBFEF7" blockId="5.[151,1436,150,357]" pageId="5" pageNumber="104">
<emphasis id="B90F9D61A6280476FF761341FEF8FED3" bold="true" box="[199,334,295,321]" pageId="5" pageNumber="104">Etymology.</emphasis>
The species epithet
<taxonomicName id="4C7B3AF0A6280476FD841341FD3DFED3" authorityName="Ulitzka" authorityYear="2024" box="[565,651,295,321]" class="Insecta" family="Stenurothripidae" genus="Campanithrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="kyakhei">
<emphasis id="B90F9D61A6280476FD841341FD3DFED3" box="[565,651,295,321]" italics="true" pageId="5" pageNumber="104">kyakhei</emphasis>
</taxonomicName>
was chosen in honour of the amber deposit ‘Kyakhe’ (meaning ‘tiger bite’) from which this fossil originates.
</paragraph>
</subSubSection>
</treatment>
</document>
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