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<document id="DA00C33BD37AC34043BEB2294EB717AE" ID-DOI="10.11646/zootaxa.5443.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10997023" ID-ZooBank="85B08D1D-489A-43A9-9E66-86755024D9FB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1713521164659" checkinUser="plazi" docAuthor="Yanega, Douglas, Goemans, Geert, Dam, Matthew Van, Gómez-Marco, Francesc &amp; Hoddle, Mark" docDate="2024" docId="03C3A664FFBCFFF158C57DCCFACAFD57" docLanguage="en" docName="zootaxa.5443.1.1.pdf" docOrigin="Zootaxa 5443 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scaralina veracruzensis Yanega &amp; Van Dam 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="44" masterDocId="FFFADE1CFF96FFDA58527E46FF9FFFD2" masterDocTitle="Description of a new genus of North and Central American planthoppers (Hemiptera: Fulgoridae) with fourteen new species" masterLastPageNumber="53" masterPageNumber="1" pageNumber="43" updateTime="1713793342614" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="2627C16C1B927A960B9E651F3D10F5FF">Description of a new genus of North and Central American planthoppers (Hemiptera: Fulgoridae) with fourteen new species</mods:title>
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<mods:namePart id="600EE2562A3C16E5E9B8406D55A8B110">Goemans, Geert</mods:namePart>
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<mods:namePart id="306D23243A297C6E45DC0000899D3865">Dam, Matthew Van</mods:namePart>
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<mods:date id="5F3DF68EF74D2E3859E62ECE2A295446">2024</mods:date>
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<paragraph id="8BD51772FFBCFFF058C57DCCFCBDFC77" blockId="42.[151,802,906,969]" box="[151,802,906,933]" pageId="42" pageNumber="43">
<heading id="D09DA01EFFBCFFF058C57DCCFCBDFC77" bold="true" box="[151,802,906,933]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="B91ECB60FFBCFFF058C57DCCFCBDFC77" bold="true" box="[151,802,906,933]" pageId="42" pageNumber="43">
<taxonomicName id="4C6A6CF1FFBCFFF058C57DCCFD28FC77" authority="Yanega &amp; Van Dam, 2024" authorityName="Yanega &amp; Van Dam" authorityYear="2024" box="[151,695,906,933]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="veracruzensis">
<emphasis id="B91ECB60FFBCFFF058C57DCCFE26FC77" bold="true" box="[151,441,906,933]" italics="true" pageId="42" pageNumber="43">Scaralina veracruzensis</emphasis>
Yanega &amp;
<collectingRegion id="49AED990FFBCFFF05A137DCDFDEDFC77" box="[577,626,907,933]" country="Turkey" name="Van" pageId="42" pageNumber="43">Van</collectingRegion>
Dam
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,
<taxonomicNameLabel id="A22D761BFFBCFFF05A967DCDFCBDFC77" box="[708,802,907,933]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C37044F9FFBCFFF058C57DE9FECEFC1B" box="[151,337,942,969]" pageId="42" pageNumber="43" type="description">
<paragraph id="8BD51772FFBCFFF058C57DE9FECEFC1B" blockId="42.[151,802,906,969]" box="[151,337,942,969]" pageId="42" pageNumber="43">
(
<figureCitation id="13510BF7FFBCFFF058CD7DE9FF69FC1B" box="[159,246,943,969]" captionStart="FIGURES 3641" captionStartId="22.[152,270,1305,1329]" captionTargetBox="[151,1435,181,1281]" captionTargetId="figure-24@22.[151,1435,181,1281]" captionTargetPageId="22" captionText="FIGURES 3641. Faces of Scaralina species. (36) S. obfusca (male); (37) S. obrienae; (38) S. orientalis; (39) S. rileyi; (40) S. sullivani; (41) S. veracruzensis." figureDoi="http://doi.org/10.5281/zenodo.10997061" httpUri="https://zenodo.org/record/10997061/files/figure.png" pageId="42" pageNumber="43">Figs 41</figureCitation>
,
<figureCitation id="13510BF7FFBCFFF059507DE8FEBFFC1B" box="[258,288,942,969]" captionStart="FIGURES 5156" captionStartId="25.[152,270,1304,1328]" captionTargetBox="[151,1435,181,1280]" captionTargetId="figure-17@25.[151,1435,181,1280]" captionTargetPageId="25" captionText="FIGURES 5156. Heads and nota of Scaralina species. (51) S. obfusca (female); (52) S. obrienae; (53) S. orientalis; (54) S. rileyi; (55) S. sullivani; (56) S. veracruzensis." figureDoi="http://doi.org/10.5281/zenodo.10997065" httpUri="https://zenodo.org/record/10997065/files/figure.png" pageId="42" pageNumber="43">56</figureCitation>
,
<figureCitation id="13510BF7FFBCFFF0597E7DE9FED6FC1B" box="[300,329,943,969]" captionStart="FIGURES 6972" captionStartId="28.[152,270,1154,1178]" captionTargetBox="[151,1435,181,1131]" captionTargetId="figure-24@28.[151,1435,181,1131]" captionTargetPageId="28" captionText="FIGURES 6972. Forewings of Scaralina species. (69) S. rileyi; (70) S. sullivani; (71) S. veracruzensis; (72) Scaralis (Alphinoides) inbio." figureDoi="http://doi.org/10.5281/zenodo.10997073" httpUri="https://zenodo.org/record/10997073/files/figure.png" pageId="42" pageNumber="43">71</figureCitation>
)
</paragraph>
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<subSubSection id="C37044F9FFBCFFF058C57DB1FAFBFBC3" box="[151,1380,1015,1041]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BD51772FFBCFFF058C57DB1FAFBFBC3" blockId="42.[151,1437,1015,2013]" box="[151,1380,1015,1041]" pageId="42" pageNumber="43">
<emphasis id="B91ECB60FFBCFFF058C57DB1FE87FBC3" bold="true" box="[151,280,1015,1041]" pageId="42" pageNumber="43">Etymology</emphasis>
. This species is named for the Mexican state from which the only known specimen was collected.
</paragraph>
</subSubSection>
<subSubSection id="C37044F9FFBCFFF058957A5CFB7BFA13" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BD51772FFBCFFF058957A5CFB7BFA13" blockId="42.[151,1437,1015,2013]" pageId="42" pageNumber="43">
<emphasis id="B91ECB60FFBCFFF058957A5CFEA5FBE6" bold="true" box="[199,314,1050,1076]" pageId="42" pageNumber="43">Diagnosis</emphasis>
. While this taxon somewhat resembles
<taxonomicName id="4C6A6CF1FFBCFFF05AAC7A5DFCE4FBE7" box="[766,891,1051,1077]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sullivani">
<emphasis id="B91ECB60FFBCFFF05AAC7A5DFCE4FBE7" box="[766,891,1051,1077]" italics="true" pageId="42" pageNumber="43">S. sullivani</emphasis>
</taxonomicName>
, which is also from
<collectingRegion id="49AED990FFBCFFF05C377A5DFB53FBE7" box="[1125,1228,1051,1077]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="42" pageNumber="43">Veracruz</collectingRegion>
, it differs in many features. The pattern of coloration is different in many respects (e.g.,
<taxonomicName id="4C6A6CF1FFBCFFF05BDC7A79FC71FB8B" box="[910,1006,1087,1113]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sullivani">
<emphasis id="B91ECB60FFBCFFF05BDC7A79FC71FB8B" box="[910,1006,1087,1113]" italics="true" pageId="42" pageNumber="43">sullivani</emphasis>
</taxonomicName>
has a paler face, many light abdominal markings, and lacks a strong black cell at the claval junction); the costal crossveins are completely evanescent basally, much as in
<taxonomicName id="4C6A6CF1FFBCFFF059267AC1FE54FB73" box="[372,459,1159,1185]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="obfusca">
<emphasis id="B91ECB60FFBCFFF059267AC1FE54FB73" box="[372,459,1159,1185]" italics="true" pageId="42" pageNumber="43">obfusca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A6CF1FFBCFFF05A517AC1FDF8FB73" box="[515,615,1159,1185]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="obrienae">
<emphasis id="B91ECB60FFBCFFF05A517AC1FDF8FB73" box="[515,615,1159,1185]" italics="true" pageId="42" pageNumber="43">obrienae</emphasis>
</taxonomicName>
; the frons in
<taxonomicName id="4C6A6CF1FFBCFFF05AAF7AC1FC08FB72" box="[765,919,1159,1184]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="veracruzensis">
<emphasis id="B91ECB60FFBCFFF05AAF7AC1FC08FB72" box="[765,919,1159,1184]" italics="true" pageId="42" pageNumber="43">veracruzensis</emphasis>
</taxonomicName>
has rather little expansion of the lower lateral lobes, so the lateral margins of the frons are nearly straight, rather than concave; the latero-basal carina of the clypeus is evanescent distally, but complete in
<taxonomicName id="4C6A6CF1FFBCFFF05AFC7A89FC91FB3B" box="[686,782,1231,1257]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sullivani">
<emphasis id="B91ECB60FFBCFFF05AFC7A89FC91FB3B" box="[686,782,1231,1257]" italics="true" pageId="42" pageNumber="43">sullivani</emphasis>
</taxonomicName>
. The strongly contrasting pattern on the mesonotum, and greatly reduced basal forewing markings, coupled with the disproportionately enlarged wing cells also apparently make this taxon distinctive; however, we do acknowledge that these are all features which can vary somewhat within a species (see note regarding
<taxonomicName id="4C6A6CF1FFBCFFF05A687B7DFD01FA87" box="[570,670,1339,1365]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="obrienae">
<emphasis id="B91ECB60FFBCFFF05A687B7DFD01FA87" box="[570,670,1339,1365]" italics="true" pageId="42" pageNumber="43">obrienae</emphasis>
</taxonomicName>
), so it is possible that future collections of additional material will result in some change in the species diagnosis. The loss of the abdomen and accompanying dissection early during the study was very unfortunate, as we never had an opportunity to directly compare the male genital characters to other taxa such as
<taxonomicName id="4C6A6CF1FFBCFFF059347BE1FE59FA13" box="[358,454,1447,1473]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sullivani">
<emphasis id="B91ECB60FFBCFFF059347BE1FE59FA13" box="[358,454,1447,1473]" italics="true" pageId="42" pageNumber="43">sullivani</emphasis>
</taxonomicName>
or
<taxonomicName id="4C6A6CF1FFBCFFF059BE7BE1FDDCFA13" box="[492,579,1447,1473]" class="Insecta" family="Fulgoridae" genus="Scaralina" kingdom="Animalia" order="Hemiptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="obfusca">
<emphasis id="B91ECB60FFBCFFF059BE7BE1FDDCFA13" box="[492,579,1447,1473]" italics="true" pageId="42" pageNumber="43">obfusca</emphasis>
</taxonomicName>
, but were instead limited to early notes that had been taken.
</paragraph>
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<subSubSection id="C37044F9FFBCFFF158957B8CFF22FDEF" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="8BD51772FFBCFFF058957B8CFEC4F933" blockId="42.[151,1437,1015,2013]" pageId="42" pageNumber="43">
<emphasis id="B91ECB60FFBCFFF058957B8CFECEFA36" bold="true" box="[199,337,1482,1508]" pageId="42" pageNumber="43">Description</emphasis>
(male only; female unknown).
<emphasis id="B91ECB60FFBCFFF05AE97B8CFD66FA37" bold="true" box="[699,761,1482,1509]" italics="true" pageId="42" pageNumber="43">Head</emphasis>
(
<figureCitation id="13510BF7FFBCFFF05B587B8DFCFCFA37" box="[778,867,1483,1509]" captionStart="FIGURES 3641" captionStartId="22.[152,270,1305,1329]" captionTargetBox="[151,1435,181,1281]" captionTargetId="figure-24@22.[151,1435,181,1281]" captionTargetPageId="22" captionText="FIGURES 3641. Faces of Scaralina species. (36) S. obfusca (male); (37) S. obrienae; (38) S. orientalis; (39) S. rileyi; (40) S. sullivani; (41) S. veracruzensis." figureDoi="http://doi.org/10.5281/zenodo.10997061" httpUri="https://zenodo.org/record/10997061/files/figure.png" pageId="42" pageNumber="43">Figs 41</figureCitation>
,
<figureCitation id="13510BF7FFBCFFF05B227B8CFC0FFA37" box="[880,912,1482,1509]" captionStart="FIGURES 5156" captionStartId="25.[152,270,1304,1328]" captionTargetBox="[151,1435,181,1280]" captionTargetId="figure-17@25.[151,1435,181,1280]" captionTargetPageId="25" captionText="FIGURES 5156. Heads and nota of Scaralina species. (51) S. obfusca (female); (52) S. obrienae; (53) S. orientalis; (54) S. rileyi; (55) S. sullivani; (56) S. veracruzensis." figureDoi="http://doi.org/10.5281/zenodo.10997065" httpUri="https://zenodo.org/record/10997065/files/figure.png" pageId="42" pageNumber="43">56</figureCitation>
). Rim of vertex mostly pale; three very broad pale marks anteriorly, otherwise black; supra-ocular lobes pale apically; posterior rim mostly pale, except darkened adjacent to supra-ocular lobes; dark spots of vertex posterior in position; black markings very well-defined and restricted to lateral bands and posterior spots. Frons with a narrow pale upper margin, very dark just below this (pale markings at corner of frontal crease near ocellus evanescent), and the ground color below a little less dark, except for scattered fine black spots surrounding setal bases. Lateral margin of frons nearly straight, lower lobes not expanded. Clypeus extensively pale, with some dark markings, latero-basal carina evanescent distally. Rostrum typical for genus.
</paragraph>
<paragraph id="8BD51772FFBCFFF0589578ACFD92F86B" blockId="42.[151,1437,1015,2013]" pageId="42" pageNumber="43">
<emphasis id="B91ECB60FFBCFFF0589578ACFE82F8D7" bold="true" box="[199,285,1770,1797]" italics="true" pageId="42" pageNumber="43">Thorax</emphasis>
(
<figureCitation id="13510BF7FFBCFFF0597B78ADFEE5F8D7" box="[297,378,1770,1797]" captionStart="FIGURES 5156" captionStartId="25.[152,270,1304,1328]" captionTargetBox="[151,1435,181,1280]" captionTargetId="figure-17@25.[151,1435,181,1280]" captionTargetPageId="25" captionText="FIGURES 5156. Heads and nota of Scaralina species. (51) S. obfusca (female); (52) S. obrienae; (53) S. orientalis; (54) S. rileyi; (55) S. sullivani; (56) S. veracruzensis." figureDoi="http://doi.org/10.5281/zenodo.10997065" httpUri="https://zenodo.org/record/10997065/files/figure.png" pageId="42" pageNumber="43">Fig. 56</figureCitation>
). Pronotum ground color pale, orange-tinted tan, dark markings generally very highly contrasting; post-ocular pronotal spots well-defined, large; midline of dorsal pronotal face pale, with black submedian bands curving laterally towards posterior, and anterolateral corners also black. Mesonotum center mostly pale, black markings relatively extensive otherwise; medial carina low; lateral carinae pale, almost straight, very slightly sinuate anteriorly or posteriorly; posterior mesonotal lobe rounded. Dorsal setae long. Legs as in generic diagnosis, markings also highly contrasting.
</paragraph>
<paragraph id="8BD51772FFBCFFF158957985FAF8FE7F" blockId="42.[151,1437,1015,2013]" lastBlockId="43.[151,1437,151,645]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B91ECB60FFBCFFF058957985FE90F80F" bold="true" box="[199,271,1987,2013]" italics="true" pageId="42" pageNumber="43">Wings</emphasis>
(
<figureCitation id="13510BF7FFBCFFF0594C7985FEEFF80F" box="[286,368,1987,2013]" captionStart="FIGURES 6972" captionStartId="28.[152,270,1154,1178]" captionTargetBox="[151,1435,181,1131]" captionTargetId="figure-24@28.[151,1435,181,1131]" captionTargetPageId="28" captionText="FIGURES 6972. Forewings of Scaralina species. (69) S. rileyi; (70) S. sullivani; (71) S. veracruzensis; (72) Scaralis (Alphinoides) inbio." figureDoi="http://doi.org/10.5281/zenodo.10997073" httpUri="https://zenodo.org/record/10997073/files/figure.png" pageId="42" pageNumber="43">Fig. 71</figureCitation>
). Forewings with dark markings on basal half of 1st claval vein somewhat interrupted; a narrow, continuous dark transverse patch from costa to clavus, at the level of the first branch of M (which is abbreviated relative to other species), and semi-translucent orange basal markings; postcostal cell black with well-defined, small, pale orange spots throughout; remainder of wing quite clear, with very limited, faint spotting. Costal crossveins relatively few, mostly straight or slightly arcuate but oblique, very obscure or absent basally; M with 23 main branches, crossveins between them mostly straight, widely spaced, making exceptionally large, elongate-rectangular cells; CuA with 34 branches, rarely more than 1 of these coming directly off of CuA, and the first branch typically precedes the claval vein juncture; greatest distance between Pcu and A
<subScript id="17EE1537FFBDFFF15BFE7FC7FC2BFE5D" attach="left" box="[940,948,385,399]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
veins exceeds greatest distance between A
<subScript id="17EE1537FFBDFFF15DC67FC7FA03FE5D" attach="left" box="[1428,1436,385,399]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
and wing margin; fused vein posterior to juncture very short. Hind wing hyaline except base, which is orange.
</paragraph>
<paragraph id="8BD51772FFBDFFF158957FF1FE2CFE27" blockId="43.[151,1437,151,645]" pageId="43" pageNumber="44">
<emphasis id="B91ECB60FFBDFFF158957FF1FF60FE03" bold="true" box="[199,255,439,465]" pageId="43" pageNumber="44">Note</emphasis>
. Genitalia and abdomen lost after dissection so only preliminary notes were taken]
<emphasis id="B91ECB60FFBDFFF15CCA7FF0FA91FE03" bold="true" box="[1176,1294,438,465]" pageId="43" pageNumber="44">
<emphasis id="B91ECB60FFBDFFF15CCA7FF0FA99FE03" bold="true" box="[1176,1286,438,465]" italics="true" pageId="43" pageNumber="44">Abdomen</emphasis>
.
</emphasis>
Dorsum and venter nearly solid black.
</paragraph>
<paragraph id="8BD51772FFBDFFF158957FB8FF22FDEF" blockId="43.[151,1437,151,645]" pageId="43" pageNumber="44">
<emphasis id="B91ECB60FFBDFFF158957FB8FE1CFDCB" bold="true" box="[199,387,510,537]" italics="true" pageId="43" pageNumber="44">Male terminalia</emphasis>
. Gonostyli black apically. Setose bulge has two rows of spinose setae extending posteriorly (?).
</paragraph>
</subSubSection>
<subSubSection id="C37044F9FFBDFFF158957C01FAF5FDB3" box="[199,1386,582,609]" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BD51772FFBDFFF158957C01FAF5FDB3" blockId="43.[151,1437,151,645]" box="[199,1386,582,609]" pageId="43" pageNumber="44">
<emphasis id="B91ECB60FFBDFFF158957C01FEEAFDB2" bold="true" box="[199,373,582,609]" pageId="43" pageNumber="44">Type material.</emphasis>
<materialsCitation id="3B021D2FFFBDFFF1592E7C00FAF5FDB3" box="[380,1386,582,609]" collectionCode="ASULOB" country="Mexico" location="Veracruz" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<emphasis id="B91ECB60FFBDFFF1592E7C00FE74FDB3" bold="true" box="[380,491,582,609]" pageId="43" pageNumber="44">
<emphasis id="B91ECB60FFBDFFF1592E7C00FE7BFDB3" bold="true" box="[380,484,582,609]" italics="true" pageId="43" pageNumber="44">
<typeStatus id="54D1A9D0FFBDFFF1592E7C00FE7BFDB3" box="[380,484,582,609]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
</emphasis>
,
</emphasis>
male:
<emphasis id="B91ECB60FFBDFFF15A657C00FCB5FDB3" bold="true" box="[567,810,582,609]" pageId="43" pageNumber="44">
<collectingCountry id="F37D57E2FFBDFFF15A657C00FD2DFDB2" box="[567,690,582,608]" name="Mexico" pageId="43" pageNumber="44">MEXICO</collectingCountry>
:
<collectingRegion id="49AED990FFBDFFF15AEF7C01FCB5FDB3" box="[701,810,583,609]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="43" pageNumber="44">Veracruz</collectingRegion>
</emphasis>
: “Serra Pescados nr. Perote” [no date] (
<collectionCode id="ED7B8FB7FFBDFFF15CB57C01FAC1FDB3" box="[1255,1374,583,609]" pageId="43" pageNumber="44">ASULOB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37044F9FFBDFFF158957C2CFACAFD57" box="[199,1365,618,645]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BD51772FFBDFFF158957C2CFACAFD57" blockId="43.[151,1437,151,645]" box="[199,1365,618,645]" pageId="43" pageNumber="44">
<emphasis id="B91ECB60FFBDFFF158957C2CFEC6FD56" bold="true" box="[199,345,618,644]" pageId="43" pageNumber="44">Distribution</emphasis>
. So far known only from the
<typeStatus id="54D1A9D0FFBDFFF15AF77C2DFD4AFD57" box="[677,725,619,645]" pageId="43" pageNumber="44">type</typeStatus>
locality in central
<collectingRegion id="49AED990FFBDFFF15BF47C2DFB92FD57" box="[934,1037,619,645]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="43" pageNumber="44">Veracruz</collectingRegion>
, near the border with
<collectingRegion id="49AED990FFBDFFF15D537C2DFACEFD57" box="[1281,1361,619,645]" country="Mexico" name="Puebla" pageId="43" pageNumber="44">Puebla</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>