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<mods:title id="136E43A95C6125E331A088F704A4EEC5">Revision of the punctatus species-group of Dissomphalus Ashmead (Hymenoptera, Bethylidae)</mods:title>
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<mods:namePart id="9A598E0AA518B4BC924F255DA6321A82">De Brito, Chirlei D.</mods:namePart>
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<mods:affiliation id="7776CC90B421B5A27BF256FB2F9E339D">Universidade Federal do Espírito Santo, Avenida Fernando Ferrari, 514, Goiabeiras, 29075 - 910 Vitória, ES, Brazil.</mods:affiliation>
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<mods:namePart id="0615704D81A4220B2EDBFFE44B51737C">Azevedo, Celso O.</mods:namePart>
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<mods:affiliation id="250CE869C203B1587617CD6E53A7407C">Universidade Federal do Espírito Santo, Avenida Fernando Ferrari, 514, Goiabeiras, 29075 - 910 Vitória, ES, Brazil.</mods:affiliation>
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<emphasis id="B951A033FFB1BA7B3F8B70FE7FC1FD9A" bold="true" box="[151,943,510,537]" pageId="2" pageNumber="391">
Key to males of 
<taxonomicName id="4C2507A2FFB1BA7B3E7C70FF7E65FD9B" authorityName="Ashmead" authorityYear="1893" box="[352,523,510,537]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B951A033FFB1BA7B3E7C70FF7E65FD9B" bold="true" box="[352,523,510,537]" italics="true" pageId="2" pageNumber="391">Dissomphalus</emphasis>
</taxonomicName>
from the 
<taxonomicName id="4C2507A2FFB1BA7B3D9970FE7E92FD9B" baseAuthorityName="Kieffer" baseAuthorityYear="1910" box="[645,764,511,537]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B951A033FFB1BA7B3D9970FE7E92FD9B" bold="true" box="[645,764,511,537]" italics="true" pageId="2" pageNumber="391">punctatus</emphasis>
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species-group
</emphasis>
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<paragraph id="8B9A7C21FFB1BA7B3F8B734279F2FDDA" blockId="2.[151,1437,579,2039]" box="[151,1436,579,602]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B73427CCDFDDA" bold="true" box="[151,163,579,600]" pageId="2" pageNumber="391">1</emphasis>
. Tergal process with tubercle (
<figureCitation id="131E60A4FFB1BA7B3EE173457E09FDDB" box="[509,615,580,602]" captionStart="FIGURE 1" captionStartId="12.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-18@12.[151,1435,181,1615]" captionTargetPageId="12" captionText="FIGURE 1. Dissomphalus sp. males. Patterns of metasomal tergal process in dorsal view. A. D. maryae sp. nov., paratype. B. D. punctatus. C. D. carolinae sp. nov., paratype. D. D. johannae sp. nov., paratype. E. D. deaneae sp. nov., paratype. F. D. margarethae sp. nov., paratype. Scale bars: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.14239041" httpUri="https://zenodo.org/record/14239041/files/figure.png" pageId="2" pageNumber="391">Figs 1D, E</figureCitation>
).................................................................. 
<emphasis id="B951A033FFB1BA7B3A8C734279F2FDDA" bold="true" box="[1424,1436,579,600]" pageId="2" pageNumber="391">2</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B736079F2FDF4" blockId="2.[151,1437,579,2039]" box="[151,1436,609,631]" pageId="2" pageNumber="391">
– Tergal process with tuft of setae (
<figureCitation id="131E60A4FFB1BA7B3D3D73607ECBFDF4" box="[545,677,609,631]" captionStart="FIGURE 1" captionStartId="12.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-18@12.[151,1435,181,1615]" captionTargetPageId="12" captionText="FIGURE 1. Dissomphalus sp. males. Patterns of metasomal tergal process in dorsal view. A. D. maryae sp. nov., paratype. B. D. punctatus. C. D. carolinae sp. nov., paratype. D. D. johannae sp. nov., paratype. E. D. deaneae sp. nov., paratype. F. D. margarethae sp. nov., paratype. Scale bars: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.14239041" httpUri="https://zenodo.org/record/14239041/files/figure.png" pageId="2" pageNumber="391">Figs 1A, C, F</figureCitation>
)............................................................. 
<emphasis id="B951A033FFB1BA7B3A8C736079F2FDF4" bold="true" box="[1424,1436,609,630]" pageId="2" pageNumber="391">3</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B737F79F2FD11" blockId="2.[151,1437,579,2039]" box="[151,1436,638,660]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B737F7CCDFD11" bold="true" box="[151,163,638,659]" pageId="2" pageNumber="391">2</emphasis>
(1). Abaxial margin of harpe projected anterad; basivolsella with spines (
<figureCitation id="131E60A4FFB1BA7B3C6A737F7FACFD11" box="[886,962,638,660]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="2" pageNumber="391">Fig. 4H</figureCitation>
)..................... 
<taxonomicName id="4C2507A2FFB1BA7B3BCF737E7927FD11" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1235,1353,638,660]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="johannae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BCF737E7927FD11" box="[1235,1353,638,660]" italics="true" pageId="2" pageNumber="391">D. johannae</emphasis>
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<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C737F79F2FD11" box="[1360,1436,638,659]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B9A7C21FFB1BA7B3F8B739D79F2FD33" blockId="2.[151,1437,579,2039]" box="[151,1436,668,690]" pageId="2" pageNumber="391">
– Abaxial margin of harpe projected anterad; basivolsella without spines (
<figureCitation id="131E60A4FFB1BA7B3C89739D7F8EFD33" box="[917,992,668,690]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Fig. 3G</figureCitation>
).................... 
<taxonomicName id="4C2507A2FFB1BA7B3BC0739D7924FD33" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1244,1354,668,689]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="deaneae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BC0739D7924FD33" box="[1244,1354,668,689]" italics="true" pageId="2" pageNumber="391">D. deaneae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C739D79F2FD33" bold="true" box="[1360,1436,668,689]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C739D79F2FD33" box="[1360,1436,668,689]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B9A7C21FFB1BA7B3F8B73B879F2FD4C" blockId="2.[151,1437,579,2039]" box="[151,1436,697,719]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B73B87CCDFD4C" bold="true" box="[151,163,697,718]" pageId="2" pageNumber="391">3</emphasis>
(1). Aedeagal dorsal lobe short (
<figureCitation id="131E60A4FFB1BA7B3EF373B87E51FD4C" box="[495,575,697,719]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="2" pageNumber="391">Figs 2A</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3D5573B87E06FD4C" box="[585,616,697,718]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">3A</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3D6E73B87EE1FD4D" box="[626,655,697,719]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">5C</figureCitation>
).............................................................. 
<emphasis id="B951A033FFB1BA7B3A8C73B879F2FD4C" bold="true" box="[1424,1436,697,718]" pageId="2" pageNumber="391">4</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B73D679F2FD69" blockId="2.[151,1437,579,2039]" box="[151,1436,726,748]" pageId="2" pageNumber="391">
– Aedeagal dorsal lobe long or very long (
<figureCitation id="131E60A4FFB1BA7B3D7973D77EDDFD69" box="[613,691,726,748]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Figs 3C</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3DA273D77EB2FD69" box="[702,732,726,747]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="2" pageNumber="391">4C</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3DFA73D77F6FFD6E" box="[742,769,726,748]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">5E</figureCitation>
).................................................... 
<emphasis id="B951A033FFB1BA7B3A9873D779F2FD69" bold="true" box="[1412,1436,726,747]" pageId="2" pageNumber="391">13</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B73F279F2FC88" blockId="2.[151,1437,579,2039]" box="[151,1436,755,778]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B73F27CCDFC8A" bold="true" box="[151,163,755,776]" pageId="2" pageNumber="391">4</emphasis>
(3). Ventral margin of harpe bifurcated (see 
<bibRefCitation id="EFB401D0FFB1BA7B3D7C73F57F0DFC8B" author="Colombo, W. D. &amp; Alencar, I. D. C. C. &amp; Limeira-De-Oliveira, F. &amp; Azevedo, C. O." box="[608,867,756,778]" pageId="2" pageNumber="391" pagination="1 - 40" refId="ref43809" refString="Colombo, W. D., Alencar, I. D. C. C., Limeira-De-Oliveira, F. &amp; Azevedo, C. O. (2018) New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic forest from northeastern Brazil. Zootaxa, 4462 (1), 1 - 40. https: // doi. org / 10.11646 / zootaxa. 4462.1.1" type="journal article" year="2018">Colombo &amp; Azevedo 2018</bibRefCitation>
).................. 
<taxonomicName id="4C2507A2FFB1BA7B3B5173F579F2FC88" authority="Colombo &amp; Azevedo" authorityName="Colombo &amp; Azevedo" authorityYear="2018" box="[1101,1436,756,778]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="acutipupu">
<emphasis id="B951A033FFB1BA7B3B5173F578A4FC8B" box="[1101,1226,756,777]" italics="true" pageId="2" pageNumber="391">D. acutipupu</emphasis>
Colombo &amp; Azevedo
</taxonomicName>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B721079F2FCA4" blockId="2.[151,1437,579,2039]" box="[151,1436,785,807]" pageId="2" pageNumber="391">
– Ventral margin of harpe not bifurcated (
<figureCitation id="131E60A4FFB1BA7B3D4372107EA7FCA4" box="[607,713,785,807]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">Figs 5A, E</figureCitation>
).......................................................... 
<emphasis id="B951A033FFB1BA7B3A8C721079F2FCA4" bold="true" box="[1424,1436,785,806]" pageId="2" pageNumber="391">5</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B722F79F2FCC1" blockId="2.[151,1437,579,2039]" box="[151,1436,814,836]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B722F7CCDFCC1" bold="true" box="[151,163,814,835]" pageId="2" pageNumber="391">5</emphasis>
(4). Basivolsella with spines (
<figureCitation id="131E60A4FFB1BA7B3ECB722F7E49FCC1" box="[471,551,814,836]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="2" pageNumber="391">Figs 4H</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3D2D722F7E21FCC6" box="[561,591,814,836]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">5G</figureCitation>
).................................................................... 
<emphasis id="B951A033FFB1BA7B3A8C722F79F2FCC1" bold="true" box="[1424,1436,814,835]" pageId="2" pageNumber="391">6</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B724D79F2FCE3" blockId="2.[151,1437,579,2039]" box="[151,1436,844,866]" pageId="2" pageNumber="391">
– Basivolsella without spines............................................................................. 
<emphasis id="B951A033FFB1BA7B3A8C724D79F2FCE3" bold="true" box="[1424,1436,844,865]" pageId="2" pageNumber="391">7</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B726879F2FC19" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B72687CCDFCFC" bold="true" box="[151,163,873,894]" pageId="2" pageNumber="391">6</emphasis>
(5). Harpe entirely wide; inner median filament almost same size of outer lobe directed dorsad; basal cover plate in diamond shape (
<figureCitation id="131E60A4FFB1BA7B3FF572877D5AFC1E" box="[233,308,902,924]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">Fig. 5D</figureCitation>
)......................................................................... 
<taxonomicName id="4C2507A2FFB1BA7B3BAE72867927FC19" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1202,1353,902,924]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="margarethae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BAE72867927FC19" box="[1202,1353,902,924]" italics="true" pageId="2" pageNumber="391">D. margarethae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C728779F2FC19" bold="true" box="[1360,1436,902,923]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C728779F2FC19" box="[1360,1436,902,923]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B72A579F2FC54" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
– Harpe wide basally; inner median filament anterior to apex of aedeagal outer lobe posterad; basal cover plate in inverted triangle shape (
<figureCitation id="131E60A4FFB1BA7B3E6972C07DD2FC54" box="[373,444,961,983]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Fig. 3F</figureCitation>
)................................................................ 
<taxonomicName id="4C2507A2FFB1BA7B3BCC72C37924FC54" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1232,1354,961,983]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BCC72C37924FC54" box="[1232,1354,961,983]" italics="true" pageId="2" pageNumber="391">D. dandarae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C72C079F2FC54" bold="true" box="[1360,1436,961,982]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C72C079F2FC54" box="[1360,1436,961,982]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B72DF79F2FC71" blockId="2.[151,1437,579,2039]" box="[151,1436,990,1012]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B72DF7CCDFC71" bold="true" box="[151,163,990,1011]" pageId="2" pageNumber="391">7</emphasis>
(5). Aedeagal dorsal outer lobe directed mesad (
<figureCitation id="131E60A4FFB1BA7B3D9472DF7EBDFC71" box="[648,723,990,1012]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="2" pageNumber="391">Fig. 4A</figureCitation>
)......................................................... 
<emphasis id="B951A033FFB1BA7B3A8C72DF79F2FC71" bold="true" box="[1424,1436,990,1011]" pageId="2" pageNumber="391">8</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B72FD79F3FB92" blockId="2.[151,1437,579,2039]" box="[151,1437,1019,1042]" pageId="2" pageNumber="391">
– Aedeagal dorsal outer lobe not directed mesad............................................................. 
<emphasis id="B951A033FFB1BA7B3A9A72FA79F3FB92" bold="true" box="[1414,1437,1019,1040]" pageId="2" pageNumber="391">11</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B751879F2FBC9" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B75187CCDFBAC" bold="true" box="[151,163,1049,1070]" pageId="2" pageNumber="391">8</emphasis>
(7). Inner median filament aligned with apex of aedeagal dorsal lobes; basal cover plate very wide; aedeagal ventral valve higher than aedeagal dorsal valve (
<figureCitation id="131E60A4FFB1BA7B3EF175377E56FBC9" box="[493,568,1078,1100]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="2" pageNumber="391">Fig. 4A</figureCitation>
)......................................................... 
<taxonomicName id="4C2507A2FFB1BA7B3BE775367927FBC9" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1275,1353,1078,1100]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="elzae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BE775367927FBC9" box="[1275,1353,1078,1100]" italics="true" pageId="2" pageNumber="391">D. elzae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C753779F2FBC9" bold="true" box="[1360,1436,1078,1099]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C753779F2FBC9" box="[1360,1436,1078,1099]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B755579F2FB04" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
– Inner median filament anterior to apex of aedeagal dorsal lobes (
<figureCitation id="131E60A4FFB1BA7B3C7C75557FC4FBEB" box="[864,938,1108,1130]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Fig. 3F</figureCitation>
); basal cover plate narrow (
<figureCitation id="131E60A4FFB1BA7B3BA175557967FBEB" box="[1213,1289,1108,1130]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="2" pageNumber="391">Figs 2F</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3A097555795BFBEB" box="[1301,1333,1108,1129]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">3A</figureCitation>
); aedeagal ventral valve not higher than aedeagal dorsal valve (
<figureCitation id="131E60A4FFB1BA7B3DCB75707F49FB04" box="[727,807,1137,1159]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Figs 3D</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3C2D75707F05FB04" box="[817,875,1137,1159]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">5A, B</figureCitation>
)............................................ 
<emphasis id="B951A033FFB1BA7B3A8C757079F2FB04" bold="true" box="[1424,1436,1137,1158]" pageId="2" pageNumber="391">9</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B758F79F2FB21" blockId="2.[151,1437,579,2039]" box="[151,1436,1166,1188]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B758F7CCDFB21" bold="true" box="[151,163,1166,1187]" pageId="2" pageNumber="391">9</emphasis>
(8). Harpe entirely wide with dorsal margin straight (
<figureCitation id="131E60A4FFB1BA7B3DAA758F7F6BFB26" box="[694,773,1166,1188]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">Figs 5D</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3C0C758F7F43FB26" box="[784,813,1166,1188]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="2" pageNumber="391">6F</figureCitation>
); apex of aedeagal ventral valve parallel (
<figureCitation id="131E60A4FFB1BA7B3BA1758F7968FB26" box="[1213,1286,1166,1188]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">Fig. 5C</figureCitation>
)......... 
<emphasis id="B951A033FFB1BA7B3A98758F79F2FB21" bold="true" box="[1412,1436,1166,1187]" pageId="2" pageNumber="391">10</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B75AD79F2FB5C" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
– Harpe wide basally with posterior margin very incurved; apex of aedeagal ventral valve slightly laterad (
<figureCitation id="131E60A4FFB1BA7B3BD275AD7977FB43" box="[1230,1305,1196,1218]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Fig. 3A</figureCitation>
)........................................................................................... 
<taxonomicName id="4C2507A2FFB1BA7B3BD475CB7924FB5C" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1224,1354,1225,1247]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="ashauerae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BD475CB7924FB5C" box="[1224,1354,1225,1247]" italics="true" pageId="2" pageNumber="391">D. ashauerae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C75C879F2FB5C" bold="true" box="[1360,1436,1225,1246]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C75C879F2FB5C" box="[1360,1436,1225,1246]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B75E779F3FB7E" blockId="2.[151,1437,579,2039]" box="[151,1437,1254,1276]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B75E77CC1FB79" bold="true" box="[151,175,1254,1275]" pageId="2" pageNumber="391">10</emphasis>
(9). Digitus very long; posterior margin of basal cover plate broadly incurved; apex of harpe wide (
<figureCitation id="131E60A4FFB1BA7B3B6D75E778D4FB79" box="[1137,1210,1254,1276]" captionStart="FIGURE 1" captionStartId="12.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-18@12.[151,1435,181,1615]" captionTargetPageId="12" captionText="FIGURE 1. Dissomphalus sp. males. Patterns of metasomal tergal process in dorsal view. A. D. maryae sp. nov., paratype. B. D. punctatus. C. D. carolinae sp. nov., paratype. D. D. johannae sp. nov., paratype. E. D. deaneae sp. nov., paratype. F. D. margarethae sp. nov., paratype. Scale bars: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.14239041" httpUri="https://zenodo.org/record/14239041/files/figure.png" pageId="2" pageNumber="391">Fig. 1B</figureCitation>
)...... 
<taxonomicName id="4C2507A2FFB1BA7B3A1275E679F3FB7E" authority="Evans" authorityName="Evans" authorityYear="1979" box="[1294,1437,1255,1276]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="napo">
<emphasis id="B951A033FFB1BA7B3A1275E67932FB7E" box="[1294,1372,1255,1276]" italics="true" pageId="2" pageNumber="391">D. napo</emphasis>
Evans
</taxonomicName>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B740579F2FAB4" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
– Digitus short; posterior margin of basal cover plate completely incurved; apex of harpe narrow (
<figureCitation id="131E60A4FFB1BA7B3B60740578A8FA9B" box="[1148,1222,1283,1306]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">Fig. 5C</figureCitation>
) .................................................................................................. 
<taxonomicName id="4C2507A2FFB1BA7B3BDE74237924FAB4" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1218,1354,1313,1335]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="lispectorae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BDE74237924FAB4" box="[1218,1354,1313,1335]" italics="true" pageId="2" pageNumber="391">D. lispectorae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C742079F2FAB4" bold="true" box="[1360,1436,1313,1334]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C742079F2FAB4" box="[1360,1436,1313,1334]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B743F79F2FAF2" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B743F7CC0FAD1" bold="true" box="[151,174,1342,1363]" pageId="2" pageNumber="391">11</emphasis>
(7). Basal cover plate very narrow with posterior margin completely incurved (
<figureCitation id="131E60A4FFB1BA7B3CB7743F7F9AFAD1" box="[939,1012,1342,1364]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="2" pageNumber="391">Fig. 2F</figureCitation>
); harpe entirely wide; inner median filament directed posterad (
<figureCitation id="131E60A4FFB1BA7B3E8F745D7D8DFAF3" box="[403,483,1372,1394]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Figs 3G</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3EF1745A7E64FAF3" box="[493,522,1371,1393]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">5B</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3D09745D7E5FFAF0" box="[533,561,1372,1394]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="2" pageNumber="391">6E</figureCitation>
)...................................................................... 
<emphasis id="B951A033FFB1BA7B3A98745A79F2FAF2" bold="true" box="[1412,1436,1371,1392]" pageId="2" pageNumber="391">12</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B747879F2FA29" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
– Basal cover plate wide with posterior margin broadly incurved; harpe wide basally; inner median filament directed ventrad (
<figureCitation id="131E60A4FFB1BA7B3FF574977D5AFA29" box="[233,308,1430,1452]" captionStart="FIGURE 7" captionStartId="39.[151,250,1057,1081]" captionTargetBox="[157,1431,565,1032]" captionTargetId="figure-159@39.[151,1436,561,1034]" captionTargetPageId="39" captionText="FIGURE 7. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. sueliae sp. nov., holotype. B. D. vivianeae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239053" httpUri="https://zenodo.org/record/14239053/files/figure.png" pageId="2" pageNumber="391">Fig. 7A</figureCitation>
)............................................................................. 
<taxonomicName id="4C2507A2FFB1BA7B3BF474967924FA29" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1256,1354,1430,1452]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="sueliae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BF474967924FA29" box="[1256,1354,1430,1452]" italics="true" pageId="2" pageNumber="391">D. sueliae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C749779F2FA29" bold="true" box="[1360,1436,1430,1451]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C749779F2FA29" box="[1360,1436,1430,1451]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B74B279F2FA64" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B74B27CC1FA4A" bold="true" box="[151,175,1459,1480]" pageId="2" pageNumber="391">12</emphasis>
(11). Apex of harpe wide; abaxial margin of harpe not projected anterad; basivolsella and digitus sclerotized (
<figureCitation id="131E60A4FFB1BA7B3BD574B5797AFA4B" box="[1225,1300,1460,1482]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="2" pageNumber="391">Fig. 3H</figureCitation>
)................................................................................................ 
<taxonomicName id="4C2507A2FFB1BA7B3BE874D37924FA64" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1268,1354,1489,1511]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="elisae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BE874D37924FA64" box="[1268,1354,1489,1511]" italics="true" pageId="2" pageNumber="391">D. elisae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C74D079F2FA64" bold="true" box="[1360,1436,1489,1510]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C74D079F2FA64" box="[1360,1436,1489,1510]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B74EE79F2F9A3" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
– Apex of harpe narrow; abaxial margin of harpe projected anterad; basivolsella and digitus not sclerotized (
<figureCitation id="131E60A4FFB1BA7B3BFC74EF7949F981" box="[1248,1319,1518,1540]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="2" pageNumber="391">Fig. 2F</figureCitation>
)............................................................................................. 
<taxonomicName id="4C2507A2FFB1BA7B3BFC770D7924F9A3" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1248,1354,1548,1569]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="ameliae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BFC770D7924F9A3" box="[1248,1354,1548,1569]" italics="true" pageId="2" pageNumber="391">D. ameliae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C770D79F2F9A3" bold="true" box="[1360,1436,1548,1569]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C770D79F2F9A3" box="[1360,1436,1548,1569]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B772879F2F9BC" blockId="2.[151,1437,579,2039]" box="[151,1436,1577,1599]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B77287CC1F9BC" bold="true" box="[151,175,1577,1598]" pageId="2" pageNumber="391">13</emphasis>
(3). Aedeagal dorsal lobe very long (
<figureCitation id="131E60A4FFB1BA7B3D0677287E06F9BC" box="[538,616,1577,1599]" captionStart="FIGURE 1" captionStartId="12.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-18@12.[151,1435,181,1615]" captionTargetPageId="12" captionText="FIGURE 1. Dissomphalus sp. males. Patterns of metasomal tergal process in dorsal view. A. D. maryae sp. nov., paratype. B. D. punctatus. C. D. carolinae sp. nov., paratype. D. D. johannae sp. nov., paratype. E. D. deaneae sp. nov., paratype. F. D. margarethae sp. nov., paratype. Scale bars: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.14239041" httpUri="https://zenodo.org/record/14239041/files/figure.png" pageId="2" pageNumber="391">Figs 1C</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3D6F77287EFFF9BD" box="[627,657,1577,1599]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">5H</figureCitation>
)............................................................. 
<emphasis id="B951A033FFB1BA7B3A98772879F2F9BC" bold="true" box="[1412,1436,1577,1598]" pageId="2" pageNumber="391">14</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B774679F2F9D9" blockId="2.[151,1437,579,2039]" box="[151,1436,1606,1628]" pageId="2" pageNumber="391">
– Aedeagal dorsal lobe long............................................................................. 
<emphasis id="B951A033FFB1BA7B3A98774779F2F9D9" bold="true" box="[1412,1436,1606,1627]" pageId="2" pageNumber="391">15</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B776279F2F9F8" blockId="2.[151,1437,579,2039]" box="[151,1436,1635,1658]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B77627CC1F9FA" bold="true" box="[151,175,1635,1656]" pageId="2" pageNumber="391">14</emphasis>
(13). Basivolsella elongated with well delimited projection (
<figureCitation id="131E60A4FFB1BA7B3DED77657F54F9FB" box="[753,826,1636,1658]" captionStart="FIGURE 1" captionStartId="12.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-18@12.[151,1435,181,1615]" captionTargetPageId="12" captionText="FIGURE 1. Dissomphalus sp. males. Patterns of metasomal tergal process in dorsal view. A. D. maryae sp. nov., paratype. B. D. punctatus. C. D. carolinae sp. nov., paratype. D. D. johannae sp. nov., paratype. E. D. deaneae sp. nov., paratype. F. D. margarethae sp. nov., paratype. Scale bars: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.14239041" httpUri="https://zenodo.org/record/14239041/files/figure.png" pageId="2" pageNumber="391">Fig. 1C</figureCitation>
)............................... 
<taxonomicName id="4C2507A2FFB1BA7B3BDB776579F2F9F8" authority="(Kieffer)" baseAuthorityName="Kieffer" baseAuthorityYear="1910" box="[1223,1436,1636,1658]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B951A033FFB1BA7B3BDB7765792FF9FB" box="[1223,1345,1636,1657]" italics="true" pageId="2" pageNumber="391">D. punctatus</emphasis>
(Kieffer)
</taxonomicName>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B778079F2F914" blockId="2.[151,1437,579,2039]" box="[151,1436,1665,1687]" pageId="2" pageNumber="391">
– Basivolsella not elongated without such projection (
<figureCitation id="131E60A4FFB1BA7B3DC977807F4EF915" box="[725,800,1665,1687]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">Fig. 5H</figureCitation>
)..................................... 
<taxonomicName id="4C2507A2FFB1BA7B3BF477837924F915" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1256,1354,1666,1687]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="martae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BF477837924F915" box="[1256,1354,1666,1687]" italics="true" pageId="2" pageNumber="391">D. martae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C778079F2F914" bold="true" box="[1360,1436,1665,1686]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C778079F2F914" box="[1360,1436,1665,1686]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B779F79F2F952" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B779F7CC1F931" bold="true" box="[151,175,1694,1715]" pageId="2" pageNumber="391">15</emphasis>
(13). Basal part of basivolsella united to aedeagal ventral valve.................................................... 
<emphasis id="B951A033FFB1BA7B3A98779F79F2F931" bold="true" box="[1412,1436,1694,1715]" pageId="2" pageNumber="391">16</emphasis>
Basal part of basivolsella not united to aedeagal ventral valve................................................. 
<emphasis id="B951A033FFB1BA7B3A9877BA79F2F952" bold="true" box="[1412,1436,1723,1744]" pageId="2" pageNumber="391">17</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B77D879F2F889" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B77D87CC1F96C" bold="true" box="[151,175,1753,1774]" pageId="2" pageNumber="391">16</emphasis>
(14). Margin of aedeagal outer lobe sinuous; ventral margin of outer lobe with pointed projection upward (
<figureCitation id="131E60A4FFB1BA7B3BB277D87897F96D" box="[1198,1273,1753,1775]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="2" pageNumber="391">Fig. 6G</figureCitation>
)................................................................................................ 
<taxonomicName id="4C2507A2FFB1BA7B3BC677F67924F889" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1242,1354,1782,1804]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="silveirae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BC677F67924F889" box="[1242,1354,1782,1804]" italics="true" pageId="2" pageNumber="391">D. silveirae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C77F779F2F889" bold="true" box="[1360,1436,1782,1803]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C77F779F2F889" box="[1360,1436,1782,1803]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B761579F2F8C5" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
– Margin of aedeagal outer lobe rounded; ventral margin of outer lobe without pointed projection upward (see 
<bibRefCitation id="EFB401D0FFB1BA7B3A3B76157D00F8C4" author="Colombo, W. D. &amp; Alencar, I. D. C. C. &amp; Limeira-De-Oliveira, F. &amp; Azevedo, C. O." pageId="2" pageNumber="391" pagination="1 - 40" refId="ref43809" refString="Colombo, W. D., Alencar, I. D. C. C., Limeira-De-Oliveira, F. &amp; Azevedo, C. O. (2018) New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic forest from northeastern Brazil. Zootaxa, 4462 (1), 1 - 40. https: // doi. org / 10.11646 / zootaxa. 4462.1.1" type="journal article" year="2018">Colombo &amp; Azevedo 2018</bibRefCitation>
)................................................................ 
<taxonomicName id="4C2507A2FFB1BA7B3B9C763379F2F8C5" authority="Colombo &amp; Azevedo" authorityName="Colombo &amp; Azevedo" authorityYear="2018" box="[1152,1436,1841,1863]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="cuca">
<emphasis id="B951A033FFB1BA7B3B9C763378A4F8C5" box="[1152,1226,1842,1863]" italics="true" pageId="2" pageNumber="391">D. cuca</emphasis>
Colombo &amp; Azevedo
</taxonomicName>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B764F79F2F800" blockId="2.[151,1437,579,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B764F7CC1F8E1" bold="true" box="[151,175,1870,1891]" pageId="2" pageNumber="391">17</emphasis>
(15). Harpe with dorsal margin slightly narrower in anterior part (see 
<bibRefCitation id="EFB401D0FFB1BA7B3C57764F7837F8E6" author="Redighieri, E. S. &amp; Azevedo, C. O." box="[843,1113,1870,1892]" pageId="2" pageNumber="391" pagination="297 - 334" refId="ref44744" refString="Redighieri, E. S. &amp; Azevedo, C. O. (2006) Fauna de Dissomphalus Ashmead (Hymenoptera, Bethylidae) da Mata Atlantica BRAZILeira, com descricao de 23 especies novas. Revista Brasielira de Entomologia, 50, 297 - 334. https: // doi. org / 10.1590 / S 0085 - 56262006000300001" type="journal article" year="2006">Redighieri &amp; Azevedo 2006</bibRefCitation>
).................................................................................................. 
<taxonomicName id="4C2507A2FFB1BA7B3B5A766D79F2F800" authority="Redighieri &amp; Azevedo" authorityName="Redighieri &amp; Azevedo" authorityYear="2006" box="[1094,1436,1900,1922]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="bahiensis">
<emphasis id="B951A033FFB1BA7B3B5A766D78D1F803" box="[1094,1215,1900,1921]" italics="true" pageId="2" pageNumber="391">D. bahiensis</emphasis>
Redighieri &amp; Azevedo
</taxonomicName>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B768879F2F81C" blockId="2.[151,1437,579,2039]" box="[151,1436,1929,1951]" pageId="2" pageNumber="391">
– Harpe with dorsal margin not slightly narrower in posterior part............................................... 
<emphasis id="B951A033FFB1BA7B3A98768879F2F81C" bold="true" box="[1412,1436,1929,1950]" pageId="2" pageNumber="391">18</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B76A779F2F839" blockId="2.[151,1437,579,2039]" box="[151,1436,1958,1980]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B76A77CC1F839" bold="true" box="[151,175,1958,1979]" pageId="2" pageNumber="391">18</emphasis>
(17). Harpe color darker than other parts of genitalia (
<figureCitation id="131E60A4FFB1BA7B3DAF76A77E92F839" box="[691,764,1958,1980]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="2" pageNumber="391">Fig. 2E</figureCitation>
)....................................... 
<taxonomicName id="4C2507A2FFB1BA7B3BF676A67924F839" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1258,1354,1958,1980]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="alzirae" status="sp. nov.">
<emphasis id="B951A033FFB1BA7B3BF676A67924F839" box="[1258,1354,1958,1980]" italics="true" pageId="2" pageNumber="391">D. alzirae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB1BA7B3A4C76A779F2F839" bold="true" box="[1360,1436,1958,1979]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A2621D48FFB1BA7B3A4C76A779F2F839" box="[1360,1436,1958,1979]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B76C579F2F85A" blockId="2.[151,1437,579,2039]" box="[151,1436,1987,2010]" pageId="2" pageNumber="391">
– Harpe color not darker than other parts of genitalia.......................................................... 
<emphasis id="B951A033FFB1BA7B3A9876C279F2F85A" bold="true" box="[1412,1436,1987,2008]" pageId="2" pageNumber="391">19</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB1BA7B3F8B76E079F2F874" blockId="2.[151,1437,579,2039]" box="[151,1436,2017,2039]" pageId="2" pageNumber="391">
<emphasis id="B951A033FFB1BA7B3F8B76E07CC1F874" bold="true" box="[151,175,2017,2038]" pageId="2" pageNumber="391">19</emphasis>
(18). Harpe very wide with dorsal margin entirely straight; basal cover plate in inverted triangle shaped (
<figureCitation id="131E60A4FFB1BA7B3B8776E07885F875" box="[1179,1259,2017,2039]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="2" pageNumber="391">Figs 5D</figureCitation>
, 
<figureCitation id="131E60A4FFB1BA7B3BE976E0797EF875" box="[1269,1296,2017,2039]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="2" pageNumber="391">6F</figureCitation>
)........ 
<emphasis id="B951A033FFB1BA7B3A9876E079F2F874" bold="true" box="[1412,1436,2017,2038]" pageId="2" pageNumber="391">20</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B719979F2FF48" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Harpe not very wide with dorsal margin strongly or slightly incurved dorsally; basal cover plate not inverted triangle shaped. .................................................................................................. 
<emphasis id="B951A033FFB0BA7A3A9871B479F2FF48" bold="true" box="[1412,1436,181,202]" pageId="3" pageNumber="392">22</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B71D379F2FE86" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B71D37CC1FF65" bold="true" box="[151,175,210,231]" pageId="3" pageNumber="392">20</emphasis>
(19). Inner median filament anterior to aedeagal dorsal lobes (
<figureCitation id="131E60A4FFB0BA7A3C1871D37F3CFF6A" box="[772,850,210,232]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="3" pageNumber="392">Figs 5E</figureCitation>
, 
<figureCitation id="131E60A4FFB0BA7A3C4071D37F17FF6A" box="[860,889,210,232]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">6E</figureCitation>
, 
<figureCitation id="131E60A4FFB0BA7A3C9F71D37FCCFF65" box="[899,930,210,231]" captionStart="FIGURE 7" captionStartId="39.[151,250,1057,1081]" captionTargetBox="[157,1431,565,1032]" captionTargetId="figure-159@39.[151,1436,561,1034]" captionTargetPageId="39" captionText="FIGURE 7. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. sueliae sp. nov., holotype. B. D. vivianeae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239053" httpUri="https://zenodo.org/record/14239053/files/figure.png" pageId="3" pageNumber="392">7B</figureCitation>
); digitus narrow (
<figureCitation id="131E60A4FFB0BA7A3B5171D378F9FF65" box="[1101,1175,210,232]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="3" pageNumber="392">Figs 2F</figureCitation>
, 
<figureCitation id="131E60A4FFB0BA7A3BBD71D378AFFF65" box="[1185,1217,210,231]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="3" pageNumber="392">4H</figureCitation>
, 
<figureCitation id="131E60A4FFB0BA7A3BD771D37882FF6A" box="[1227,1260,210,232]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">6H</figureCitation>
); posterior margin of hypopygium straight or nearly so ...................................................................... 
<emphasis id="B951A033FFB0BA7A3A9871EE79F2FE86" bold="true" box="[1412,1436,239,260]" pageId="3" pageNumber="392">21</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B700C79F2FEBD" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Inner median filament aligned with aedeagal outer lobes; digitus wide; posterior margin of hypopygium weakly outcurved (
<figureCitation id="131E60A4FFB0BA7A3A69700C7C91FEC2" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="3" pageNumber="392">Fig. 5B</figureCitation>
)................................................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BEB702A7927FEBD" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1271,1353,298,320]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="limae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BEB702A7927FEBD" box="[1271,1353,298,320]" italics="true" pageId="3" pageNumber="392">D. limae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C702B79F2FEBD" bold="true" box="[1360,1436,298,319]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C702B79F2FEBD" box="[1360,1436,298,319]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B704679F2FEDF" blockId="3.[151,1437,151,2021]" box="[151,1436,327,349]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B70467CC1FEDE" bold="true" box="[151,175,327,348]" pageId="3" pageNumber="392">21</emphasis>
(20). Digitus very long and narrow; basivolsella with narrow protuberance (
<figureCitation id="131E60A4FFB0BA7A3C9B70467FA1FEDF" box="[903,975,327,349]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">Fig. 6F</figureCitation>
)........................ 
<taxonomicName id="4C2507A2FFB0BA7A3BE370497927FEDF" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1279,1353,328,349]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="ritae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BE370497927FEDF" box="[1279,1353,328,349]" italics="true" pageId="3" pageNumber="392">D. ritae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C704979F2FEDF" bold="true" box="[1360,1436,328,349]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C704979F2FEDF" box="[1360,1436,328,349]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B706479F2FEF8" blockId="3.[151,1437,151,2021]" box="[151,1436,357,379]" pageId="3" pageNumber="392">
– Digitus short and wide; basivolsella with wide protuberance (
<figureCitation id="131E60A4FFB0BA7A3C2C70647F15FEF9" box="[816,891,357,379]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="3" pageNumber="392">Fig. 5D</figureCitation>
).............................. 
<taxonomicName id="4C2507A2FFB0BA7A3BEF70647927FEF8" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1267,1353,357,378]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="luisae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BEF70647927FEF8" box="[1267,1353,357,378]" italics="true" pageId="3" pageNumber="392">D. luisae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C706479F2FEF8" bold="true" box="[1360,1436,357,378]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C706479F2FEF8" box="[1360,1436,357,378]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B708379F2FE15" blockId="3.[151,1437,151,2021]" box="[151,1436,386,408]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B70837CC1FE15" bold="true" box="[151,175,386,407]" pageId="3" pageNumber="392">22</emphasis>
(19). Aedeagal dorsal lobes wide (
<figureCitation id="131E60A4FFB0BA7A3EEB70837E29FE15" box="[503,583,386,408]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="3" pageNumber="392">Figs 3D</figureCitation>
, 
<figureCitation id="131E60A4FFB0BA7A3D4D70837E05FE1A" box="[593,619,386,408]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="3" pageNumber="392">5F</figureCitation>
, 
<figureCitation id="131E60A4FFB0BA7A3D6970837EFCFE1A" box="[629,658,386,408]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">6D</figureCitation>
) .............................................................. 
<emphasis id="B951A033FFB0BA7A3A98708379F2FE15" bold="true" box="[1412,1436,386,407]" pageId="3" pageNumber="392">23</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B70A179F2FE36" blockId="3.[151,1437,151,2021]" box="[151,1436,415,437]" pageId="3" pageNumber="392">
– Aedeagal outer lobes narrow........................................................................... 
<emphasis id="B951A033FFB0BA7A3A98709E79F2FE36" bold="true" box="[1412,1436,415,436]" pageId="3" pageNumber="392">25</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B70BC79F2FE50" blockId="3.[151,1437,151,2021]" box="[151,1436,445,467]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B70BC7CC1FE50" bold="true" box="[151,175,445,466]" pageId="3" pageNumber="392">23</emphasis>
(22). Harpe with dorsal margin deeply incurved posteriorly with apex narrow........................................ 
<emphasis id="B951A033FFB0BA7A3A9870BC79F2FE50" bold="true" box="[1412,1436,445,466]" pageId="3" pageNumber="392">24</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B70DA79F2FE6D" blockId="3.[151,1437,151,2021]" box="[151,1436,474,496]" pageId="3" pageNumber="392">
– Harpe with dorsal margin not deeply incurved posteriorly with apex wide (
<figureCitation id="131E60A4FFB0BA7A3CAC70DB7F95FE6D" box="[944,1019,474,496]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="3" pageNumber="392">Fig. 3D</figureCitation>
)................ 
<taxonomicName id="4C2507A2FFB0BA7A3BD270DA7924FE72" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1230,1354,475,496]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="carmenae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BD270DA7924FE72" box="[1230,1354,475,496]" italics="true" pageId="3" pageNumber="392">D. carmenae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C70DB79F2FE6D" bold="true" box="[1360,1436,474,495]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C70DB79F2FE6D" box="[1360,1436,474,495]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B70F679F2FDA8" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B70F67CC1FD8E" bold="true" box="[151,175,503,524]" pageId="3" pageNumber="392">24</emphasis>
(23). Margin of aedeagal outer lobes rounded; basal cover plate with ventrolateral corner very narrow; apex of aedeagal ventral valve parallel (
<figureCitation id="131E60A4FFB0BA7A3E2B73147DECFDA9" box="[311,386,533,555]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">Fig. 6D</figureCitation>
)........................................................................ 
<taxonomicName id="4C2507A2FFB0BA7A3BEF73147927FDA8" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1267,1353,533,554]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="nisiae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BEF73147927FDA8" box="[1267,1353,533,554]" italics="true" pageId="3" pageNumber="392">D. nisiae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C731479F2FDA8" bold="true" box="[1360,1436,533,554]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C731479F2FDA8" box="[1360,1436,533,554]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B733279F2FDE7" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Margin of aedeagal outer lobes almost straight with pointed protuberance posterad; basal cover plate entirely wide; apex of aedeagal ventral valve curved laterad (
<figureCitation id="131E60A4FFB0BA7A3D48734E7EF5FDE7" box="[596,667,591,613]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="3" pageNumber="392">Fig. 5F</figureCitation>
)................................................ 
<taxonomicName id="4C2507A2FFB0BA7A3BEF73517927FDE6" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1267,1353,591,613]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="lutzae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BEF73517927FDE6" box="[1267,1353,591,613]" italics="true" pageId="3" pageNumber="392">D. lutzae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C735179F2FDE7" bold="true" box="[1360,1436,592,613]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C735179F2FDE7" box="[1360,1436,592,613]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B736C79F2FD00" blockId="3.[151,1437,151,2021]" box="[151,1436,621,643]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B736C7CC1FD00" bold="true" box="[151,175,621,642]" pageId="3" pageNumber="392">25</emphasis>
(22). Inner median filament curved laterad.................................................................... 
<emphasis id="B951A033FFB0BA7A3A98736C79F2FD00" bold="true" box="[1412,1436,621,642]" pageId="3" pageNumber="392">26</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B738A79F2FD1D" blockId="3.[151,1437,151,2021]" box="[151,1436,650,672]" pageId="3" pageNumber="392">
– Inner median filament curved posterad, dorsad or ventrad.................................................... 
<emphasis id="B951A033FFB0BA7A3A98738B79F2FD1D" bold="true" box="[1412,1436,650,671]" pageId="3" pageNumber="392">27</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B73A679F2FD58" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B73A67CC1FD3E" bold="true" box="[151,175,679,700]" pageId="3" pageNumber="392">26</emphasis>
(25). Posterior margin of basal cover plate incurved medially; basivolsellar protuberance poorly delimited; posterior margin of harpe outcurved (
<figureCitation id="131E60A4FFB0BA7A3E5373C47DF6FD58" box="[335,408,709,731]" captionStart="FIGURE 7" captionStartId="39.[151,250,1057,1081]" captionTargetBox="[157,1431,565,1032]" captionTargetId="figure-159@39.[151,1436,561,1034]" captionTargetPageId="39" captionText="FIGURE 7. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. sueliae sp. nov., holotype. B. D. vivianeae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239053" httpUri="https://zenodo.org/record/14239053/files/figure.png" pageId="3" pageNumber="392">Fig. 7B</figureCitation>
)................................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BCC73C47924FD58" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1232,1354,709,730]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="vivianeae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BCC73C47924FD58" box="[1232,1354,709,730]" italics="true" pageId="3" pageNumber="392">D. vivianeae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C73C479F2FD58" bold="true" box="[1360,1436,709,730]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C73C479F2FD58" box="[1360,1436,709,730]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B73E279F2FC97" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Posterior margin of basal cover plate not incurved dorsally; basivolsellar protuberance well delimited; posterior margin of harpe slightly incurved (
<figureCitation id="131E60A4FFB0BA7A3E8D73FE7DB2FC97" box="[401,476,767,789]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="3" pageNumber="392">Fig. 2H</figureCitation>
)............................................................. 
<taxonomicName id="4C2507A2FFB0BA7A3BDB72017927FC97" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1223,1353,768,789]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="antonietae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BDB72017927FC97" box="[1223,1353,768,789]" italics="true" pageId="3" pageNumber="392">D. antonietae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C720179F2FC97" bold="true" box="[1360,1436,768,789]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C720179F2FC97" box="[1360,1436,768,789]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B721C79F2FCB0" blockId="3.[151,1437,151,2021]" box="[151,1436,797,819]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B721C7CC1FCB0" bold="true" box="[151,175,797,818]" pageId="3" pageNumber="392">27</emphasis>
(25). Gonostipe is about two times shorter than harpe (
<figureCitation id="131E60A4FFB0BA7A3DA0721C7F6AFCB1" box="[700,772,797,819]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="3" pageNumber="392">Fig. 5E</figureCitation>
)....................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BEE721C7924FCB0" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1266,1354,797,818]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="lumae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BEE721C7924FCB0" box="[1266,1354,797,818]" italics="true" pageId="3" pageNumber="392">D. lumae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C721C79F2FCB0" bold="true" box="[1360,1436,797,818]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C721C79F2FCB0" box="[1360,1436,797,818]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B723A79F2FCCD" blockId="3.[151,1437,151,2021]" box="[151,1436,826,848]" pageId="3" pageNumber="392">
– Gonostipe as long as harpe or nearly so................................................................... 
<emphasis id="B951A033FFB0BA7A3A98723B79F2FCCD" bold="true" box="[1412,1436,826,847]" pageId="3" pageNumber="392">28</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B725679F2FC09" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B72567CC1FCEE" bold="true" box="[151,175,855,876]" pageId="3" pageNumber="392">28</emphasis>
(27). Inner median filament posterior to aedeagal dorsal lobes; apex of harpe very narrow with abaxial margin projected anterad (see Colombo &amp; 
<bibRefCitation id="EFB401D0FFB0BA7A3E4572747D8BFC09" author="Azevedo, C. O. &amp; Colombo, W. D. &amp; Alencar, I. D. C. C. &amp; Brito, C. D. de &amp; Waichert, C." box="[345,485,885,907]" pageId="3" pageNumber="392" refId="ref43617" refString="Azevedo, C. O., Colombo, W. D., Alencar, I. D. C. C., Brito, C. D. de &amp; Waichert, C. (2016) Couples in phoretic copulation, a tool for male-female association in highly dimorphic insects of the wasp genus Dissomphalus Ashmead (Hymenoptera: Bethylidae). Zoologia, 33 (6), e 20160076. https: // doi. org / 10.1590 / S 1984 - 4689 zool- 20160076." type="journal volume" year="2016">Azevedo 2016</bibRefCitation>
).................................................. 
<taxonomicName id="4C2507A2FFB0BA7A3B51727479F2FC09" authority="Colombo &amp; Azevedo" authorityName="Colombo &amp; Azevedo" authorityYear="2016" box="[1101,1436,885,907]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="tupinikum">
<emphasis id="B951A033FFB0BA7A3B51727478A4FC08" box="[1101,1226,885,906]" italics="true" pageId="3" pageNumber="392">D. tupinikum</emphasis>
Colombo &amp; Azevedo
</taxonomicName>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B729279F2FC25" blockId="3.[151,1437,151,2021]" box="[151,1436,914,936]" pageId="3" pageNumber="392">
– Inner median filament aligned or anterior to apex of aedeagal outer lobes; apex of harpe wide or very wide............. 
<emphasis id="B951A033FFB0BA7A3A98729379F2FC25" bold="true" box="[1412,1436,914,935]" pageId="3" pageNumber="392">29</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B72AE79F2FC46" blockId="3.[151,1437,151,2021]" box="[151,1436,943,965]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B72AE7CC1FC46" bold="true" box="[151,175,943,964]" pageId="3" pageNumber="392">29</emphasis>
(28). Inner median filament aligned with apex of aedeagal lobes................................................... 
<emphasis id="B951A033FFB0BA7A3A9872AE79F2FC46" bold="true" box="[1412,1436,943,964]" pageId="3" pageNumber="392">30</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B72CC79F2FC60" blockId="3.[151,1437,151,2021]" box="[151,1436,973,995]" pageId="3" pageNumber="392">
– Inner median filament anterior to apex of aedeagal lobes..................................................... 
<emphasis id="B951A033FFB0BA7A3A9872CC79F2FC60" bold="true" box="[1412,1436,973,994]" pageId="3" pageNumber="392">31</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B72EB79F2FB9F" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B72EB7CC1FC7D" bold="true" box="[151,175,1002,1023]" pageId="3" pageNumber="392">30</emphasis>
(29). Aedeagal dorsal valve in bottle shaped; basal cover plate narrow with posterior margin straight; basivolsellar protuberance well delimited (
<figureCitation id="131E60A4FFB0BA7A3E5675067DFDFB9F" box="[330,403,1031,1053]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="3" pageNumber="392">Fig. 2A</figureCitation>
).................................................................. 
<taxonomicName id="4C2507A2FFB0BA7A3BDB750979F2FB9F" authority="(Kieffer)" baseAuthorityName="Kieffer" baseAuthorityYear="1910" box="[1223,1436,1031,1053]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B951A033FFB0BA7A3BDB7509792FFB9F" box="[1223,1345,1032,1053]" italics="true" pageId="3" pageNumber="392">D. montanus</emphasis>
(Kieffer)
</taxonomicName>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B752479F2FBD5" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Aedeagal dorsal margin almost in bottle or nearly so; basal cover plate wide with posterior margin slightly incurved; basivolsellar protuberance poorly delimited (
<figureCitation id="131E60A4FFB0BA7A3D0E75437E35FBD5" box="[530,603,1090,1112]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="3" pageNumber="392">Fig. 3C</figureCitation>
)...................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BF275427924FBD5" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1262,1354,1090,1112]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="boriae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BF275427924FBD5" box="[1262,1354,1090,1112]" italics="true" pageId="3" pageNumber="392">D. boriae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C754379F2FBD5" bold="true" box="[1360,1436,1090,1111]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C754379F2FBD5" box="[1360,1436,1090,1111]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B755E79F2FBF6" blockId="3.[151,1437,151,2021]" box="[151,1436,1119,1141]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B755E7CC1FBF6" bold="true" box="[151,175,1119,1140]" pageId="3" pageNumber="392">31</emphasis>
(29). Basal cover plate very wide medially in inverted triangle shape............................................... 
<emphasis id="B951A033FFB0BA7A3A98755E79F2FBF6" bold="true" box="[1412,1436,1119,1140]" pageId="3" pageNumber="392">32</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B757C79F2FB10" blockId="3.[151,1437,151,2021]" box="[151,1436,1149,1171]" pageId="3" pageNumber="392">
– Basal cover plate is completely narrow, not inverted triangle shape............................................. 
<emphasis id="B951A033FFB0BA7A3A98757C79F2FB10" bold="true" box="[1412,1436,1149,1170]" pageId="3" pageNumber="392">33</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B759B79F2FB4F" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B759B7CC1FB2D" bold="true" box="[151,175,1178,1199]" pageId="3" pageNumber="392">32</emphasis>
(31). Inner median filament directed posterad; basal cover plate with sclerotized oblique structure; apodeme far beyond genital ring (
<figureCitation id="131E60A4FFB0BA7A3FF575B67D5BFB4F" box="[233,309,1207,1229]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">Fig. 6H</figureCitation>
)............................................................................. 
<taxonomicName id="4C2507A2FFB0BA7A3BF275B97924FB4F" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1262,1354,1208,1229]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="soniae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BF275B97924FB4F" box="[1262,1354,1208,1229]" italics="true" pageId="3" pageNumber="392">D. soniae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C75B979F2FB4F" bold="true" box="[1360,1436,1208,1229]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C75B979F2FB4F" box="[1360,1436,1208,1229]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B75D479F2FA85" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Inner median filament directed dorsad; basal cover plate without such structure; apodeme not far beyond genital ring (
<figureCitation id="131E60A4FFB0BA7A3A5675D479FBFB68" box="[1354,1429,1237,1259]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="3" pageNumber="392">Fig. 2G</figureCitation>
)....................................................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BEE75F27924FA8A" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1266,1354,1267,1288]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="annae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BEE75F27924FA8A" box="[1266,1354,1267,1288]" italics="true" pageId="3" pageNumber="392">D. annae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C75F379F2FA85" bold="true" box="[1360,1436,1266,1287]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C75F379F2FA85" box="[1360,1436,1266,1287]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B740E79F2FAA6" blockId="3.[151,1437,151,2021]" box="[151,1436,1295,1317]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B740E7CC1FAA6" bold="true" box="[151,175,1295,1316]" pageId="3" pageNumber="392">33</emphasis>
(31). Apodeme far beyond genital ring; dorsal margin of harpe entirely narrow........................................ 
<emphasis id="B951A033FFB0BA7A3A98740E79F2FAA6" bold="true" box="[1412,1436,1295,1316]" pageId="3" pageNumber="392">34</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B742C79F2FAC0" blockId="3.[151,1437,151,2021]" box="[151,1436,1325,1347]" pageId="3" pageNumber="392">
– Apodeme not far beyond genital ring; dorsal margin of harpe not entirely narrow.................................. 
<emphasis id="B951A033FFB0BA7A3A98742C79F2FAC0" bold="true" box="[1412,1436,1325,1346]" pageId="3" pageNumber="392">37</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B744B79F2FAFF" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B744B7CC1FADD" bold="true" box="[151,175,1354,1375]" pageId="3" pageNumber="392">34</emphasis>
(33). Median region of aedeagal dorsal valve slightly incurved medially; posterior margin of basal cover plate incurved medially (
<figureCitation id="131E60A4FFB0BA7A3FF574667D5AFAFF" box="[233,308,1383,1405]" captionStart="FIGURE 5" captionStartId="28.[151,250,1909,1933]" captionTargetBox="[239,1352,189,1875]" captionTargetId="figure-18@28.[229,1359,181,1884]" captionTargetPageId="28" captionText="FIGURE 5. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. leliae sp. nov., paratype. B. D. limae sp. nov., holotype. C. D. lispectorae sp. nov., holotype. D. D. luisae sp. nov., holotype. E. D. lumae sp. nov., holotype. F. D. lutzae sp. nov., paratype. G. D. margarethae sp. nov., holotype. H. D. martae sp. nov., paratype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239049" httpUri="https://zenodo.org/record/14239049/files/figure.png" pageId="3" pageNumber="392">Fig. 5A</figureCitation>
).............................................................................. 
<taxonomicName id="4C2507A2FFB0BA7A3BEB74697927FAFE" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1271,1353,1383,1405]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="leliae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BEB74697927FAFE" box="[1271,1353,1383,1405]" italics="true" pageId="3" pageNumber="392">D. leliae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C746979F2FAFF" bold="true" box="[1360,1436,1384,1405]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C746979F2FAFF" box="[1360,1436,1384,1405]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B748479F2FA18" blockId="3.[151,1437,151,2021]" box="[151,1436,1413,1435]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B74847CCDFA18" bold="true" box="[151,163,1413,1434]" pageId="3" pageNumber="392">–</emphasis>
Median region of aedeagal dorsal valve outcurved; posterior margin of basal cover plate not incurved medially.......... 
<emphasis id="B951A033FFB0BA7A3A98748479F2FA18" bold="true" box="[1412,1436,1413,1434]" pageId="3" pageNumber="392">35</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B74A379F2FA57" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B74A37CC1FA35" bold="true" box="[151,175,1442,1463]" pageId="3" pageNumber="392">35</emphasis>
(34).Apex of aedeagal dorsal valve not abruptly narrow; basal cover plate entirely narrow with posterior margin straight; basivolsellar protuberance well demarcated and wide; ventral margin of harpe with small projection (
<figureCitation id="131E60A4FFB0BA7A3B3274BE7816FA57" box="[1070,1144,1471,1493]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">Fig. 6C</figureCitation>
)...... 
<taxonomicName id="4C2507A2FFB0BA7A3BCC74C17924FA56" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1232,1354,1471,1493]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="mirandae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BCC74C17924FA56" box="[1232,1354,1471,1493]" italics="true" pageId="3" pageNumber="392">D. mirandae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C74C179F2FA57" bold="true" box="[1360,1436,1472,1493]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C74C179F2FA57" box="[1360,1436,1472,1493]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B74DC79F2F98D" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Apex of aedeagal dorsal valve abruptly narrow; basal cover plate entirely narrow with posterior margin broadly rounded and narrow in dorsolateral margin; basivolsella without protuberance (
<figureCitation id="131E60A4FFB0BA7A3C4174FB7FCAF98D" box="[861,932,1530,1552]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="3" pageNumber="392">Fig. 4F</figureCitation>
)....................... 
<taxonomicName id="4C2507A2FFB0BA7A3BCF74FA7927F98D" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1235,1353,1530,1552]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="guidonae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BCF74FA7927F98D" box="[1235,1353,1530,1552]" italics="true" pageId="3" pageNumber="392">D. guidonae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C74FB79F2F98D" bold="true" box="[1360,1436,1530,1551]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C74FB79F2F98D" box="[1360,1436,1530,1551]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B771679F2F9AE" blockId="3.[151,1437,151,2021]" box="[151,1436,1559,1581]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B77167CC1F9AE" bold="true" box="[151,175,1559,1580]" pageId="3" pageNumber="392">37</emphasis>
(33). Basivolsella without protuberance....................................................................... 
<emphasis id="B951A033FFB0BA7A3A98771679F2F9AE" bold="true" box="[1412,1436,1559,1580]" pageId="3" pageNumber="392">38</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B773479F2F9C8" blockId="3.[151,1437,151,2021]" box="[151,1436,1589,1611]" pageId="3" pageNumber="392">
– Basivolsella with protuberance......................................................................... 
<emphasis id="B951A033FFB0BA7A3A98773479F2F9C8" bold="true" box="[1412,1436,1589,1610]" pageId="3" pageNumber="392">40</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B775379F2F907" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B77537CC1F9E5" bold="true" box="[151,175,1618,1639]" pageId="3" pageNumber="392">38</emphasis>
(37). Apex of aedeagal outer lobes directed laterad; dorsal margin of harpe slightly incurved posteriorly; aedeagal ventral valve parallel (
<figureCitation id="131E60A4FFB0BA7A3E2B776E7DEEF907" box="[311,384,1647,1669]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="3" pageNumber="392">Fig. 3B</figureCitation>
)..................................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BCF77717927F906" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1235,1353,1647,1669]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="beatrizae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BCF77717927F906" box="[1235,1353,1647,1669]" italics="true" pageId="3" pageNumber="392">D. beatrizae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C777179F2F907" bold="true" box="[1360,1436,1648,1669]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C777179F2F907" box="[1360,1436,1648,1669]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B778C79F2F93D" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Apex of aedeagal outer lobes directed posterad; dorsal margin of harpe strongly incurved posteriorly; aedeagal ventral valve angled laterad....................................................................................... 
<emphasis id="B951A033FFB0BA7A3A9877AB79F2F93D" bold="true" box="[1412,1436,1706,1727]" pageId="3" pageNumber="392">39</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B77C679F2F978" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B77C67CC1F95E" bold="true" box="[151,175,1735,1756]" pageId="3" pageNumber="392">39</emphasis>
(38). Tergal process with dense tuft of setae; dorsal margin of harpe strongly incurved posteriorly; basal cover plate entirely narrow (
<figureCitation id="131E60A4FFB0BA7A3FF577E47D5AF978" box="[233,308,1765,1787]" captionStart="FIGURE 2" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[217,1367,190,1916]" captionTargetId="figure-18@14.[209,1378,181,1919]" captionTargetPageId="14" captionText="FIGURE 2. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. montanus, holotype. B. D. napo, holotype. C. D. punctatus, holotype. D. D. alexandrinae sp. nov., holotype. E. D. alzirae sp. nov., holotype. F. D. ameliae sp. nov., holotype. G. D. annae sp. nov., holotype. H. D. antonietae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239043" httpUri="https://zenodo.org/record/14239043/files/figure.png" pageId="3" pageNumber="392">Fig. 2D</figureCitation>
)......................................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BB277E47924F978" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1198,1354,1765,1786]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="alexandrinae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BB277E47924F978" box="[1198,1354,1765,1786]" italics="true" pageId="3" pageNumber="392">D. alexandrinae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C77E479F2F978" bold="true" box="[1360,1436,1765,1786]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C77E479F2F978" box="[1360,1436,1765,1786]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B760279F2F8B7" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Tergal process without dense tuft of setae; dorsal margin of harpe not strongly incurved posteriorly; basal cover plate very narrow in dorsolateral corner (
<figureCitation id="131E60A4FFB0BA7A3D1A761E7E3CF8B7" box="[518,594,1823,1845]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="3" pageNumber="392">Fig. 4D</figureCitation>
)................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BD476217924F8B7" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1224,1354,1824,1845]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="evaristoae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BD476217924F8B7" box="[1224,1354,1824,1845]" italics="true" pageId="3" pageNumber="392">D. evaristoae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C762179F2F8B7" bold="true" box="[1360,1436,1824,1845]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C762179F2F8B7" box="[1360,1436,1824,1845]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B763C79F2F8D0" blockId="3.[151,1437,151,2021]" box="[151,1436,1853,1875]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B763C7CC1F8D0" bold="true" box="[151,175,1853,1874]" pageId="3" pageNumber="392">40</emphasis>
(37). Basivolsellar protuberance narrow...................................................................... 
<emphasis id="B951A033FFB0BA7A3A98763C79F2F8D0" bold="true" box="[1412,1436,1853,1874]" pageId="3" pageNumber="392">41</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B765A79F2F8ED" blockId="3.[151,1437,151,2021]" box="[151,1436,1882,1904]" pageId="3" pageNumber="392">
– Basivolsellar protuberance wide........................................................................ 
<emphasis id="B951A033FFB0BA7A3A98765B79F2F8ED" bold="true" box="[1412,1436,1882,1903]" pageId="3" pageNumber="392">42</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B767679F2F828" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
<emphasis id="B951A033FFB0BA7A3F8B76767CC1F80E" bold="true" box="[151,175,1911,1932]" pageId="3" pageNumber="392">41</emphasis>
(40). Aedeagal dorsal valve robust; aedeagal ventral valve posterior to aedeagal dorsal valve; dorsal margin of harpe strongly incurved posteriorly (
<figureCitation id="131E60A4FFB0BA7A3EAD76947D93F829" box="[433,509,1941,1963]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="3" pageNumber="392">Fig. 6A</figureCitation>
)............................................................. 
<taxonomicName id="4C2507A2FFB0BA7A3BF876947924F828" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1252,1354,1941,1962]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BF876947924F828" box="[1252,1354,1941,1962]" italics="true" pageId="3" pageNumber="392">D. maryae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C769479F2F828" bold="true" box="[1360,1436,1941,1962]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C769479F2F828" box="[1360,1436,1941,1962]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB0BA7A3F8B76B279F2F867" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="392">
– Aedeagal dorsal valve narrow; aedeagal ventral valve aligned to aedeagal dorsal valve; dorsal margin of harpe not strongly posteriorly (
<figureCitation id="131E60A4FFB0BA7A3E4B76CE7DCFF867" box="[343,417,1999,2021]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="3" pageNumber="392">Fig. 4C</figureCitation>
).................................................................... 
<taxonomicName id="4C2507A2FFB0BA7A3BC376D17927F866" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1247,1353,1999,2021]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="392" phylum="Arthropoda" rank="species" species="eulaliae" status="sp. nov.">
<emphasis id="B951A033FFB0BA7A3BC376D17927F866" box="[1247,1353,1999,2021]" italics="true" pageId="3" pageNumber="392">D. eulaliae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB0BA7A3A4C76D179F2F867" bold="true" box="[1360,1436,2000,2021]" pageId="3" pageNumber="392">
<taxonomicNameLabel id="A2621D48FFB0BA7A3A4C76D179F2F867" box="[1360,1436,2000,2021]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB7BA7D3F8B719679F2FF6A" blockId="4.[151,1437,151,584]" pageId="4" pageNumber="393">
<emphasis id="B951A033FFB7BA7D3F8B71967CC1FF2E" bold="true" box="[151,175,151,172]" pageId="4" pageNumber="393">42</emphasis>
(40). Basal cover plate narrow; apex of harpe truncate and very wide................................................ 
<emphasis id="B951A033FFB7BA7D3A98719679F2FF2E" bold="true" box="[1412,1436,151,172]" pageId="4" pageNumber="393">43</emphasis>
– Basal cover plate wide; apex of harpe sinuous and not very wide ............................................... 
<emphasis id="B951A033FFB7BA7D3A9871B47CC1FF65" bold="true" pageId="4" pageNumber="393">44 43</emphasis>
(42). Basidorsal corner of digitus strongly projected; posterior hypopygeal margin straight or nearly so; aedeagal dorsal valve narrow
</paragraph>
<paragraph id="8B9A7C21FFB7BA7D3FFE71EE79FBFEDF" blockId="4.[151,1437,151,584]" pageId="4" pageNumber="393">
(
<figureCitation id="131E60A4FFB7BA7D3FF571EE7D5AFE87" box="[233,308,239,261]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="4" pageNumber="393">Fig. 4G</figureCitation>
)......................................................................... 
<taxonomicName id="4C2507A2FFB7BA7D3BA071F17924FE86" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1212,1354,239,261]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="jaquelineae" status="sp. nov.">
<emphasis id="B951A033FFB7BA7D3BA071F17924FE86" box="[1212,1354,239,261]" italics="true" pageId="4" pageNumber="393">D. jaquelineae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB7BA7D3A4C71F179F2FE87" bold="true" box="[1360,1436,240,261]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A2621D48FFB7BA7D3A4C71F179F2FE87" box="[1360,1436,240,261]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
– Basidorsal corner of digitus poorly projected; posterior hypopygeal margin sinuous; aedeagal dorsal valve in bottle shape or nearly so (
<figureCitation id="131E60A4FFB7BA7D3E5A702B7DE0FEC2" box="[326,398,298,320]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="4" pageNumber="393">Fig. 6E</figureCitation>
)...................................................................... 
<taxonomicName id="4C2507A2FFB7BA7D3BFB702A7927FEBD" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1255,1353,298,320]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="penhae" status="sp. nov.">
<emphasis id="B951A033FFB7BA7D3BFB702A7927FEBD" box="[1255,1353,298,320]" italics="true" pageId="4" pageNumber="393">D. penhae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB7BA7D3A4C702B7CC1FEDE" bold="true" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A2621D48FFB7BA7D3A4C702B79F2FEBD" box="[1360,1436,298,319]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
44
</emphasis>
(42). Posterior margin of basal cover plate incurved medially; basivolsellar protuberance poorly delimited (
<figureCitation id="131E60A4FFB7BA7D3BA870467893FEDF" box="[1204,1277,327,349]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="4" pageNumber="393">Fig. 4E</figureCitation>
)............
</paragraph>
<paragraph id="8B9A7C21FFB7BA7D3FF7706479F2FE37" blockId="4.[151,1437,151,584]" pageId="4" pageNumber="393">
................................................................................... 
<taxonomicName id="4C2507A2FFB7BA7D3BCF70647927FEF8" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1235,1353,357,378]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="grazielae" status="sp. nov.">
<emphasis id="B951A033FFB7BA7D3BCF70647927FEF8" box="[1235,1353,357,378]" italics="true" pageId="4" pageNumber="393">D. grazielae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB7BA7D3A4C706479F2FEF8" bold="true" box="[1360,1436,357,378]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A2621D48FFB7BA7D3A4C706479F2FEF8" box="[1360,1436,357,378]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
– Posterior margin of basal cover plate broadly incurved; basivolsellar protuberance well projected..................... 
<emphasis id="B951A033FFB7BA7D3A9870837CC1FE36" bold="true" pageId="4" pageNumber="393">45 45</emphasis>
(44). Dorsal margin of harpe wide; basidorsal corner of digitus poorly projected (
<figureCitation id="131E60A4FFB7BA7D3CA6709E786CFE37" box="[954,1026,415,437]" captionStart="FIGURE 3" captionStartId="15.[151,250,1909,1933]" captionTargetBox="[222,1367,188,1873]" captionTargetId="figure-17@15.[211,1376,181,1884]" captionTargetPageId="15" captionText="FIGURE 3. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. ashauerae sp. nov., paratype. B. D. beatrizae sp. nov., holotype. C. D. boriae sp. nov., holotype. D. D. carmenae sp. nov., paratype. E. D. carolinae sp. nov., paratype. F. D. dandarae sp. nov., paratype. G. D. deaneae sp. nov., holotype. H. D. elisae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239045" httpUri="https://zenodo.org/record/14239045/files/figure.png" pageId="4" pageNumber="393">Fig. 3E</figureCitation>
)............... 
<taxonomicName id="4C2507A2FFB7BA7D3BCD70A17927FE36" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1233,1353,415,437]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B951A033FFB7BA7D3BCD70A17927FE36" box="[1233,1353,415,437]" italics="true" pageId="4" pageNumber="393">D. carolinae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB7BA7D3A4C70A179F2FE37" bold="true" box="[1360,1436,416,437]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A2621D48FFB7BA7D3A4C70A179F2FE37" box="[1360,1436,416,437]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A7C21FFB7BA7D3FFE70BC79F3FE72" blockId="4.[151,1437,151,584]" pageId="4" pageNumber="393">
Dorsal margin of harpe narrow; basidorsal corner of digitus strongly projected.................................... 
<emphasis id="B951A033FFB7BA7D3A9870BC7CC1FE6D" bold="true" pageId="4" pageNumber="393">46 46</emphasis>
(45). Tergal process with dense tuft of setae; inner median filament directed ventrad; ventral margin of harpe without projection (
<figureCitation id="131E60A4FFB7BA7D3A6970DB79F3FE72" box="[1397,1437,474,496]" captionStart="FIGURE 6" captionStartId="36.[151,250,1905,1929]" captionTargetBox="[225,1364,190,1872]" captionTargetId="figure-18@36.[214,1373,181,1881]" captionTargetPageId="36" captionText="FIGURE 6. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. maryae sp. nov., holotype. B. D. meirelesae sp. nov., holotype. C. D. mirandae sp. nov., holotype. D. D. nisiae sp. nov., holotype. E. D. penhae sp. nov., holotype. F. D. ritae sp. nov. holotype. G. D. silveirae sp. nov., paratype. H. D. soniae sp. nov., holotype. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239051" httpUri="https://zenodo.org/record/14239051/files/figure.png" pageId="4" pageNumber="393">Fig.</figureCitation>
</paragraph>
<paragraph id="8B9A7C21FFB7BA7D3FFE70F679F2FDA9" blockId="4.[151,1437,151,584]" pageId="4" pageNumber="393">
6B)............................................................................... 
<taxonomicName id="4C2507A2FFB7BA7D3BD970F97927FD8E" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1221,1353,503,525]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="meirelesae" status="sp. nov.">
<emphasis id="B951A033FFB7BA7D3BD970F97927FD8E" box="[1221,1353,503,525]" italics="true" pageId="4" pageNumber="393">D. meirelesae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB7BA7D3A4C70F979F2FD8F" bold="true" box="[1360,1436,504,525]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A2621D48FFB7BA7D3A4C70F979F2FD8F" box="[1360,1436,504,525]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
– Tergal process with tuft of little setae; inner median filament directed posterad; ventral margin of harpe with small projection
</paragraph>
<paragraph id="8B9A7C21FFB7BA7D3FFE733379F2FDC5" blockId="4.[151,1437,151,584]" box="[226,1436,562,584]" pageId="4" pageNumber="393">
(
<figureCitation id="131E60A4FFB7BA7D3FF573337D5DFDC5" box="[233,307,562,584]" captionStart="FIGURE 4" captionStartId="23.[151,250,1909,1933]" captionTargetBox="[228,1366,192,1872]" captionTargetId="figure-17@23.[214,1374,181,1884]" captionTargetPageId="23" captionText="FIGURE 4. Genitalia of Dissomphalus sp. males in ventral (in the left) and dorsal view (in the right). A. D. elzae sp. nov., holotype. B. D. enedinae sp. nov., paratype. C. D. eulaliae sp. nov., holotype. D. D. evaristoae sp. nov., holotype. E. D. grazielae sp. nov., holotype. F. D. guidonae sp. nov., holotype. G. D. jaquelineae sp. nov., holotype. H. D. johannae sp. nov., holotype. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.14239047" httpUri="https://zenodo.org/record/14239047/files/figure.png" pageId="4" pageNumber="393">Fig. 4B</figureCitation>
)............................................................................ 
<taxonomicName id="4C2507A2FFB7BA7D3BCA73327924FDC5" authorityName="De Brito &amp; Azevedo" authorityYear="2024" box="[1238,1354,562,584]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="enedinae" status="sp. nov.">
<emphasis id="B951A033FFB7BA7D3BCA73327924FDC5" box="[1238,1354,562,584]" italics="true" pageId="4" pageNumber="393">D. enedinae</emphasis>
</taxonomicName>
<emphasis id="B951A033FFB7BA7D3A4C733379F2FDC5" bold="true" box="[1360,1436,562,583]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="A2621D48FFB7BA7D3A4C733379F2FDC5" box="[1360,1436,562,583]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>