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<document id="20755D8B647DDDDFC87FA156EBB7B747" ID-CLB-Dataset="21893" ID-DOI="10.5852/ejt.2016.221" ID-GBIF-Dataset="6994309f-b9aa-4a68-a050-f8fa43a63ada" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840279" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590171138210" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2016" docId="8D3887BCFFECFF93DD5D4DB88CFF2C22" docLanguage="en" docName="ejt-2016-221.pdf.imf" docOrigin="European Journal of Taxonomy 221" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Utiliverpa decapsulatrix Enghoff 2016, gen. et sp. nov." docType="treatment" docVersion="7" lastPageNumber="16" masterDocId="7101FFC4FFE1FF9CDF704E4D8E192816" masterDocTitle="A mountain of millipedes V: three new genera of Odontopygidae from the Udzungwa mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="14" updateTime="1698841056769" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AD78C29C4EFA73D2C52218918071E5C6">A mountain of millipedes V: three new genera of Odontopygidae from the Udzungwa mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:name id="12F57D073AAF08408295E2A1AC838CC2" type="personal">
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<mods:namePart id="F6849B230EF45911ED8D3FEEAB7E0D36">Enghoff, Henrik</mods:namePart>
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<mods:title id="04AD2492DC74F998350627437D47C159">European Journal of Taxonomy</mods:title>
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<mods:part id="8463C2004E040468D352128B08B44569">
<mods:date id="BF41CAF4E7FCD89A1D199CFB1FDE67E1">2016</mods:date>
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<mods:number id="64A3593F302A01011674B5BBC61F8E6C">2016-08-12</mods:number>
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<mods:number id="F0284CB501929EAED79B4021D95B7C21">221</mods:number>
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<mods:identifier id="A31A61573C8136215063652F246371FB" type="DOI">10.5852/ejt.2016.221</mods:identifier>
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<treatment id="8D3887BCFFECFF93DD5D4DB88CFF2C22" ID-DOI="http://doi.org/10.5281/zenodo.3853130" ID-GBIF-Taxon="164252615" ID-Zenodo-Dep="3853130" LSID="urn:lsid:plazi:treatment:8D3887BCFFECFF93DD5D4DB88CFF2C22" httpUri="http://treatment.plazi.org/id/8D3887BCFFECFF93DD5D4DB88CFF2C22" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="4D8B6521FFECFF91DD5D4DB88AD42C22" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="052E36AAFFECFF91DD5D4DB88A1E2C06" blockId="13.[358,1229,1013,1111]" box="[557,1031,1013,1040]" pageId="13" pageNumber="14">
<heading id="5E6681C6FFECFF91DD5D4DB88A1E2C06" box="[557,1031,1013,1040]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<taxonomicName id="C2914D29FFECFF91DD5D4DB88D492C06" ID-CoL="7F64D" authority="Enghoff, 2016" authorityName="Enghoff" authorityYear="2016" box="[557,848,1013,1040]" class="Diplopoda" family="Odontopygidae" genus="Utiliverpa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="decapsulatrix" status="gen. et sp. nov.">
<emphasis id="37E5EAB8FFECFF91DD5D4DB88D492C06" bold="true" box="[557,848,1013,1040]" italics="true" pageId="13" pageNumber="14">Utiliverpa decapsulatrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2CD657C3FFECFF91DC274DBB8A1E2C06" box="[855,1031,1014,1040]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="052E36AAFFECFF91DE164A548AD42C22" blockId="13.[358,1229,1013,1111]" box="[358,1229,1049,1076]" pageId="13" pageNumber="14">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="4D8B6521FFECFF91DD884A718D252C41" box="[760,828,1084,1111]" pageId="13" pageNumber="14" type="description">
<paragraph id="052E36AAFFECFF91DD884A718D252C41" blockId="13.[358,1229,1013,1111]" box="[760,828,1084,1111]" pageId="13" pageNumber="14">
<figureCitation id="9DAA2A2FFFECFF91DD884A718D252C41" box="[760,828,1084,1111]" captionStart="Fig" captionStartId="14.[189,230,1824,1850]" captionTargetBox="[217,1366,476,1797]" captionTargetId="figure@14.[217,1366,476,1797]" captionTargetPageId="14" captionText="Fig. 7. Utiliverpa decapsulatrix gen. et sp. nov., holotype. AD. Right gonopod coxa. A. Anterior view. B. Mesal view. C. Posterior view. D. Posterior-apical view. EH. Left gonopod telopodite. E. Anterior view. F. Posterior view. G. Mesal-anterior-apical view. H. Solenomere. I. Limbus. Abbreviations: All = apical lateral lobe of coxa, mbp = metaplical bifid process, mf = metaplical flange, mp = metaplica, msp = metaplical spine, prl = proplical lobe, pp = proplica, ps proximal solenomeral spine, slm = solenomere, tlm = telomere, tss = telomeral subapical spike, tt = torsotope. Scales: AH = 0.1 mm; I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3840295" httpUri="https://zenodo.org/record/3840295/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4D8B6521FFECFF91DFCD4A378C072CD5" box="[189,542,1146,1219]" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="052E36AAFFECFF91DFCD4A378F2F2C82" blockId="13.[189,310,1146,1172]" box="[189,310,1146,1172]" pageId="13" pageNumber="14">
<heading id="5E6681C6FFECFF91DFCD4A378F2F2C82" bold="true" box="[189,310,1146,1172]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="37E5EAB8FFECFF91DFCD4A378F2F2C82" bold="true" box="[189,310,1146,1172]" pageId="13" pageNumber="14">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="052E36AAFFECFF91DFCD4AE58C072CD5" blockId="13.[189,542,1192,1219]" box="[189,542,1192,1219]" pageId="13" pageNumber="14">
<emphasis id="37E5EAB8FFECFF91DFCD4AE58EDE2CD4" bold="true" box="[189,199,1192,1218]" pageId="13" pageNumber="14">(</emphasis>
redundant, genus monotypic)
</paragraph>
</subSubSection>
<subSubSection id="4D8B6521FFECFF93DFCD4AAB8C972B1B" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="etymology">
<paragraph id="052E36AAFFECFF91DFCD4AAB8F5D2D16" blockId="13.[189,324,1254,1280]" box="[189,324,1254,1280]" pageId="13" pageNumber="14">
<heading id="5E6681C6FFECFF91DFCD4AAB8F5D2D16" bold="true" box="[189,324,1254,1280]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="37E5EAB8FFECFF91DFCD4AAB8F5D2D16" bold="true" box="[189,324,1254,1280]" pageId="13" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="052E36AAFFECFF91DFCD4B588A9E2D39" blockId="13.[189,1159,1301,1327]" box="[189,1159,1301,1327]" pageId="13" pageNumber="14">The name is a Latin noun meaning “remover of capsules”, cf. etymology of genus.</paragraph>
<paragraph id="052E36AAFFECFF91DFCD4B1F8C0E2D7B" blockId="13.[189,535,1362,1389]" box="[189,535,1362,1389]" pageId="13" pageNumber="14">
<emphasis id="37E5EAB8FFECFF91DFCD4B1F8F952D7A" bold="true" box="[189,396,1362,1388]" pageId="13" pageNumber="14">Material studied</emphasis>
(total: 1 Ƌ)
</paragraph>
<paragraph id="052E36AAFFECFF91DF9A4BCC8F452D8D" blockId="13.[189,1398,1409,1506]" box="[234,348,1409,1435]" pageId="13" pageNumber="14">
<emphasis id="37E5EAB8FFECFF91DF9A4BCC8F452D8D" bold="true" box="[234,348,1409,1435]" pageId="13" pageNumber="14">
<typeStatus id="DA2A8808FFECFF91DF9A4BCC8F452D8D" box="[234,348,1409,1435]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="052E36AAFFECFF91DFCD4BE98AB12DF7" blockId="13.[189,1398,1409,1506]" pageId="13" pageNumber="14">
<materialsCitation id="B5F93CF7FFECFF91DFCD4BE98AB12DF7" ID-GBIF-Occurrence="2625232301" collectionCode="VMNH" collectorName="K. M. Howell" country="Tanzania" county="Kilombero District" location="Mwanihana Forest Reserve" municipality="Udzungwa Mts" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Morogoro" typeStatus="holotype">
<collectingCountry id="7D86763AFFECFF91DFCD4BE98F4E2DA8" box="[189,343,1444,1470]" name="Tanzania" pageId="13" pageNumber="14">TANZANIA</collectingCountry>
: Ƌ,
<collectingRegion id="C755F848FFECFF91DEFB4BE88C7A2DA8" box="[395,611,1444,1471]" country="Tanzania" name="Morogoro" pageId="13" pageNumber="14">Morogoro Region</collectingRegion>
,
<collectingCounty id="EC4F4E26FFECFF91DD014BE98D4C2DA8" box="[625,853,1444,1470]" pageId="13" pageNumber="14">Kilombero District</collectingCounty>
,
<collectingMunicipality id="E54AACD0FFECFF91DC134BE98A022DA9" box="[867,1051,1444,1471]" pageId="13" pageNumber="14">Udzungwa Mts</collectingMunicipality>
,
<location id="004E6071FFECFF91DB5A4BE98B6B2DA9" LSID="urn:lsid:plazi:treatment:8D3887BCFFECFF93DD5D4DB88CFF2C22:004E6071FFECFF91DB5A4BE98B6B2DA9" box="[1066,1394,1444,1471]" country="Tanzania" county="Kilombero District" municipality="Udzungwa Mts" name="Mwanihana Forest Reserve" pageId="13" pageNumber="14" stateProvince="Morogoro">Mwanihana Forest Reserve</location>
, near
<location id="004E6071FFECFF91DF874B8A8F8B2DF7" LSID="urn:lsid:plazi:treatment:8D3887BCFFECFF93DD5D4DB88CFF2C22:004E6071FFECFF91DF874B8A8F8B2DF7" box="[247,402,1479,1505]" country="Tanzania" county="Kilombero District" municipality="Udzungwa Mts" name="Sanje village" pageId="13" pageNumber="14" stateProvince="Morogoro">Sanje village</location>
, under rotten log in forest,
<collectorName id="A864537CFFECFF91DDA04B8A8D7D2DF7" box="[720,868,1479,1505]" pageId="13" pageNumber="14">K.M.Howell</collectorName>
leg., KMH 1686 (
<collectionCode id="6380AE6FFFECFF91DB4C4B8A8A852DF7" box="[1084,1180,1479,1505]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="13" pageNumber="14">VMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="052E36AAFFECFF91DFCD48488FBB2E36" blockId="13.[189,418,1541,1568]" box="[189,418,1541,1568]" pageId="13" pageNumber="14">
<emphasis id="37E5EAB8FFECFF91DFCD48488F572E09" bold="true" box="[189,334,1541,1567]" pageId="13" pageNumber="14">Description</emphasis>
(male)
</paragraph>
<paragraph id="052E36AAFFECFF91DFCD48798AE52E5B" blockId="13.[189,1276,1587,1614]" box="[189,1276,1587,1614]" pageId="13" pageNumber="14">
SIZE. Length ca
<quantity id="C2699B4FFFECFF91DE08487E8FAC2E58" box="[376,437,1587,1614]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="14" unit="cm" value="6.0">6 cm</quantity>
, diameter
<quantity id="C2699B4FFFECFF91DD5E48798C922E58" box="[558,651,1588,1614]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="14" unit="mm" value="3.9">3.9 mm</quantity>
, 67 podous rings, no apodous rings in front of telson.
</paragraph>
<paragraph id="052E36AAFFECFF91DFCD483C8DEB2EB8" blockId="13.[189,1398,1649,1711]" pageId="13" pageNumber="14">COLOUR. Specimen faded, traces of a broad, light dorsal stripe. Colour according to the collectors field notes: “dark, shiny black with brown stripe length of back; legs pale”.</paragraph>
<paragraph id="052E36AAFFECFF91DFCD489F8C112EFA" blockId="13.[189,520,1746,1772]" box="[189,520,1746,1772]" pageId="13" pageNumber="14">HEAD. Without peculiarities.</paragraph>
<paragraph id="052E36AAFFECFF91DFCD495D8D392F3F" blockId="13.[189,800,1807,1834]" box="[189,800,1807,1834]" pageId="13" pageNumber="14">COLLUM. With a marginal and a submarginal furrow.</paragraph>
<paragraph id="052E36AAFFECFF91DFCD49038F152F9D" blockId="13.[189,1398,1869,1931]" pageId="13" pageNumber="14">BODY RINGS. Almost perfect cylinders, not vaulted; suture straight; ozopores ca three diameters behind suture.</paragraph>
<paragraph id="052E36AAFFECFF91DFCD49E38CCB2FFD" blockId="13.[189,1398,1966,2027]" pageId="13" pageNumber="14">ANAL VALVES. Each with a short, curved dorsal spine and a small ventral denticle; marginal rim slightly raised, setiferous tubercles hardly protruding.</paragraph>
<paragraph id="052E36AAFFEFFF92DFCD4F468C342933" blockId="14.[189,557,267,294]" box="[189,557,267,294]" pageId="14" pageNumber="15">OZOPORES. Starting from ring 6.</paragraph>
<paragraph id="052E36AAFFEFFF92DFCD4F078B362975" blockId="14.[189,1327,329,356]" box="[189,1327,329,356]" pageId="14" pageNumber="15">
LIMBUS (
<figureCitation id="9DAA2A2FFFEFFF92DE514F048F6B2972" box="[289,370,329,356]" captionStart="Fig" captionStartId="14.[189,230,1824,1850]" captionTargetBox="[217,1366,476,1797]" captionTargetId="figure@14.[217,1366,476,1797]" captionTargetPageId="14" captionText="Fig. 7. Utiliverpa decapsulatrix gen. et sp. nov., holotype. AD. Right gonopod coxa. A. Anterior view. B. Mesal view. C. Posterior view. D. Posterior-apical view. EH. Left gonopod telopodite. E. Anterior view. F. Posterior view. G. Mesal-anterior-apical view. H. Solenomere. I. Limbus. Abbreviations: All = apical lateral lobe of coxa, mbp = metaplical bifid process, mf = metaplical flange, mp = metaplica, msp = metaplical spine, prl = proplical lobe, pp = proplica, ps proximal solenomeral spine, slm = solenomere, tlm = telomere, tss = telomeral subapical spike, tt = torsotope. Scales: AH = 0.1 mm; I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3840295" httpUri="https://zenodo.org/record/3840295/files/figure.png" pageId="14" pageNumber="15">Fig. 7I</figureCitation>
). With simple, long-triangular lobes. Surface of lobes longitudinally microstriate.
</paragraph>
<paragraph id="052E36AAFFEFFF92DFCD4FCA8D8F29D2" blockId="14.[189,1398,391,452]" pageId="14" pageNumber="15">MALE LEGS. With postfemoral and tibial ventral pads on legs, except a few anteriormost and posteriormost pairs, pads gradually decreasing in size towards posterior end.</paragraph>
<caption id="51EE6622FFEFFF92DFCD496D8B542FFD" ID-DOI="http://doi.org/10.5281/zenodo.3840295" ID-Zenodo-Dep="3840295" httpUri="https://zenodo.org/record/3840295/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,230,1824,1850]" targetBox="[217,1366,476,1797]" targetPageId="14">
<paragraph id="052E36AAFFEFFF92DFCD496D8B542FFD" blockId="14.[189,1399,1824,2027]" pageId="14" pageNumber="15">
<emphasis id="37E5EAB8FFEFFF92DFCD496D8F102F2D" bold="true" box="[189,265,1824,1851]" pageId="14" pageNumber="15">Fig. 7.</emphasis>
<taxonomicName id="C2914D29FFEFFF92DE60496D8C312F2C" authority="Enghoff, 2016" authorityName="Enghoff" authorityYear="2016" box="[272,552,1824,1850]" class="Diplopoda" family="Odontopygidae" genus="Utiliverpa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="decapsulatrix" status="gen. et sp. nov.">
<emphasis id="37E5EAB8FFEFFF92DE60496D8C312F2C" box="[272,552,1824,1850]" italics="true" pageId="14" pageNumber="15">Utiliverpa decapsulatrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2CD657C3FFEFFF92DD40496C8CFC2F2D" box="[560,741,1825,1851]" pageId="14" pageNumber="15" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="37E5EAB8FFEFFF92DC13496D8D852F2C" bold="true" box="[867,924,1824,1850]" pageId="14" pageNumber="15">AD</emphasis>
. Right gonopod coxa. A. Anterior view. B. Mesal view. C. Posterior view. D. Posterior-apical view.
<emphasis id="37E5EAB8FFEFFF92DC0E49098DAE2F48" bold="true" box="[894,951,1860,1886]" pageId="14" pageNumber="15">EH</emphasis>
. Left gonopod telopodite. E. Anterior view. F. Posterior view. G. Mesal-anterior-apical view. H. Solenomere.
<emphasis id="37E5EAB8FFEFFF92DB75492A8A092F97" bold="true" box="[1029,1040,1895,1921]" pageId="14" pageNumber="15">I</emphasis>
. Limbus. Abbreviations:
<emphasis id="37E5EAB8FFEFFF92DA4C492A8B472F97" box="[1340,1374,1895,1921]" italics="true" pageId="14" pageNumber="15">All</emphasis>
= apical lateral lobe of coxa,
<emphasis id="37E5EAB8FFEFFF92DE8349C78C3F2FB2" box="[499,550,1930,1956]" italics="true" pageId="14" pageNumber="15">mbp</emphasis>
= metaplical bifid process,
<emphasis id="37E5EAB8FFEFFF92DC1249C78D662FB2" box="[866,895,1930,1956]" italics="true" pageId="14" pageNumber="15">mf</emphasis>
= metaplical flange,
<emphasis id="37E5EAB8FFEFFF92DB0049C68A8D2FB2" box="[1136,1172,1931,1956]" italics="true" pageId="14" pageNumber="15">mp</emphasis>
= metaplica, msp = metaplical spine,
<emphasis id="37E5EAB8FFEFFF92DEFA49E08FB52FD1" box="[394,428,1965,1991]" italics="true" pageId="14" pageNumber="15">prl</emphasis>
= proplical lobe,
<emphasis id="37E5EAB8FFEFFF92DD0B49E38C812FD1" box="[635,664,1966,1991]" italics="true" pageId="14" pageNumber="15">pp</emphasis>
= proplica,
<emphasis id="37E5EAB8FFEFFF92DC5549E38D262FD1" box="[805,831,1966,1991]" italics="true" pageId="14" pageNumber="15">ps</emphasis>
proximal solenomeral spine,
<emphasis id="37E5EAB8FFEFFF92DBD149E08AD32FD1" box="[1185,1226,1965,1991]" italics="true" pageId="14" pageNumber="15">slm</emphasis>
= solenomere,
<emphasis id="37E5EAB8FFEFFF92DFCD499D8EFB2FFC" box="[189,226,2000,2026]" italics="true" pageId="14" pageNumber="15">tlm</emphasis>
= telomere,
<emphasis id="37E5EAB8FFEFFF92DE07499C8F8F2FFC" box="[375,406,2001,2026]" italics="true" pageId="14" pageNumber="15">tss</emphasis>
= telomeral subapical spike,
<emphasis id="37E5EAB8FFEFFF92DD9E499C8CE72FFC" box="[750,766,2001,2026]" italics="true" pageId="14" pageNumber="15">tt</emphasis>
= torsotope. Scales: AH = 0.1 mm; I = 0.01 mm.
</paragraph>
</caption>
<paragraph id="052E36AAFFEEFF93DFCD4F468B7F29A4" blockId="15.[189,1399,265,435]" pageId="15" pageNumber="16">
GONOPOD COXA (
<figureCitation id="9DAA2A2FFFEEFF93DE0C4F468FD82930" box="[380,449,267,294]" captionStart="Fig" captionStartId="14.[189,230,1824,1850]" captionTargetBox="[217,1366,476,1797]" captionTargetId="figure@14.[217,1366,476,1797]" captionTargetPageId="14" captionText="Fig. 7. Utiliverpa decapsulatrix gen. et sp. nov., holotype. AD. Right gonopod coxa. A. Anterior view. B. Mesal view. C. Posterior view. D. Posterior-apical view. EH. Left gonopod telopodite. E. Anterior view. F. Posterior view. G. Mesal-anterior-apical view. H. Solenomere. I. Limbus. Abbreviations: All = apical lateral lobe of coxa, mbp = metaplical bifid process, mf = metaplical flange, mp = metaplica, msp = metaplical spine, prl = proplical lobe, pp = proplica, ps proximal solenomeral spine, slm = solenomere, tlm = telomere, tss = telomeral subapical spike, tt = torsotope. Scales: AH = 0.1 mm; I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3840295" httpUri="https://zenodo.org/record/3840295/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
AD). In anterior view parallel-sided, ca 4½ times as long as broad, subapically with a lateral triangular incision delimiting an apical lateral lobe (
<emphasis id="37E5EAB8FFEEFF93DCD44F628DDA295F" box="[932,963,303,329]" italics="true" pageId="15" pageNumber="16">all</emphasis>
). Proplica (
<emphasis id="37E5EAB8FFEEFF93DB394F628A7F295E" box="[1097,1126,303,328]" italics="true" pageId="15" pageNumber="16">pp</emphasis>
) simple, parallel-sided; proplical lobe (
<emphasis id="37E5EAB8FFEEFF93DE024F1F8F8D297A" box="[370,404,338,364]" italics="true" pageId="15" pageNumber="16">prl</emphasis>
) in anterior view hidden behind metaplical spine. Metaplica (
<emphasis id="37E5EAB8FFEEFF93DB044F1E8A81297A" box="[1140,1176,339,364]" italics="true" pageId="15" pageNumber="16">mp</emphasis>
) apically rounded, with a large mesal flange (
<emphasis id="37E5EAB8FFEEFF93DE844F388C082999" box="[500,529,373,399]" italics="true" pageId="15" pageNumber="16">mf</emphasis>
) closing basal part of space between pro- and metaplica, subapically with a long basad spine (
<emphasis id="37E5EAB8FFEEFF93DED64FD48FCC29A4" box="[422,469,409,434]" italics="true" pageId="15" pageNumber="16">msp</emphasis>
) covering proplical lobe and a stout, bifid process (
<emphasis id="37E5EAB8FFEEFF93DB424FD58A7C29A4" box="[1074,1125,408,434]" italics="true" pageId="15" pageNumber="16">mbp</emphasis>
) on posterior surface.
</paragraph>
<paragraph id="052E36AAFFEEFF93DFCD4F978C972B1B" blockId="15.[189,1399,473,781]" pageId="15" pageNumber="16">
GONOPOD TELOPODITE (
<figureCitation id="9DAA2A2FFFEEFF93DEBA4F948C0D29E2" box="[458,532,473,500]" captionStart="Fig" captionStartId="14.[189,230,1824,1850]" captionTargetBox="[217,1366,476,1797]" captionTargetId="figure@14.[217,1366,476,1797]" captionTargetPageId="14" captionText="Fig. 7. Utiliverpa decapsulatrix gen. et sp. nov., holotype. AD. Right gonopod coxa. A. Anterior view. B. Mesal view. C. Posterior view. D. Posterior-apical view. EH. Left gonopod telopodite. E. Anterior view. F. Posterior view. G. Mesal-anterior-apical view. H. Solenomere. I. Limbus. Abbreviations: All = apical lateral lobe of coxa, mbp = metaplical bifid process, mf = metaplical flange, mp = metaplica, msp = metaplical spine, prl = proplical lobe, pp = proplica, ps proximal solenomeral spine, slm = solenomere, tlm = telomere, tss = telomeral subapical spike, tt = torsotope. Scales: AH = 0.1 mm; I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3840295" httpUri="https://zenodo.org/record/3840295/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
EH). Arculus 90°. Torsotope (
<emphasis id="37E5EAB8FFEEFF93DCEC4F978DB529E5" box="[924,940,474,499]" italics="true" pageId="15" pageNumber="16">tt</emphasis>
) simple, compact, without processes (
<figureCitation id="9DAA2A2FFFEEFF93DFB64FB08F392A01" box="[198,288,509,535]" captionStart="Fig" captionStartId="7.[189,230,1683,1709]" captionTargetBox="[279,1298,264,1646]" captionTargetId="figure@7.[279,1298,263,1648]" captionTargetPageId="7" captionText="Fig. 4. Casuariverpa scarpa gen. et sp. nov., paratype from Udzungwa Scarp Forest Reserve above Chita village, left gonopod telopodite. A. Apical view, coxal parts in orange colour. B. Basal view, purple oval roughly corresponds to Fig. 4D, coxal parts in orange colour, the green line indicates the distal margin of the telomere. C.Tip of solenomere. D. Origin of proximal solenomeral spine (ps) between two basal lobe of telomere (tlm), image roughly corresponding to purple oval on Fig. 4B. E. Micro-spinose field on basal external side of telomere basis, image roughly corresponding to area in yellow circle on Fig. 3G. F. Micro-spinose field on distal part of telomere concavity, image roughly corresponding to area in red oval on Fig. 3H. Abbreviations: eg = efferent groove, pn = post-torsal narrowing, ps = proximal solenomeral spine, slm = solenomere, tf = telomeral furrow, tlm = telomere, tr = telomeral ridge tt = torsotope. Scales: AB = 0.1 mm; CF = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3840289" httpUri="https://zenodo.org/record/3840289/files/figure.png" pageId="15" pageNumber="16">Fig. 4B</figureCitation>
). Posttorsal narrowing without processes or spines. Telopodite just distal to posttorsal narrowing dividing into slender solenomere and broader telomere. Solenomere (
<emphasis id="37E5EAB8FFEEFF93DC894C6D8A3B2A2C" box="[1017,1058,544,570]" italics="true" pageId="15" pageNumber="16">slm</emphasis>
) ca as long as and normally resting within concavity of telomere (projecting perpendicularly on
<figureCitation id="9DAA2A2FFFEEFF93DC834C0E8A542A48" box="[1011,1101,579,606]" captionStart="Fig" captionStartId="14.[189,230,1824,1850]" captionTargetBox="[217,1366,476,1797]" captionTargetId="figure@14.[217,1366,476,1797]" captionTargetPageId="14" captionText="Fig. 7. Utiliverpa decapsulatrix gen. et sp. nov., holotype. AD. Right gonopod coxa. A. Anterior view. B. Mesal view. C. Posterior view. D. Posterior-apical view. EH. Left gonopod telopodite. E. Anterior view. F. Posterior view. G. Mesal-anterior-apical view. H. Solenomere. I. Limbus. Abbreviations: All = apical lateral lobe of coxa, mbp = metaplical bifid process, mf = metaplical flange, mp = metaplica, msp = metaplical spine, prl = proplical lobe, pp = proplica, ps proximal solenomeral spine, slm = solenomere, tlm = telomere, tss = telomeral subapical spike, tt = torsotope. Scales: AH = 0.1 mm; I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3840295" httpUri="https://zenodo.org/record/3840295/files/figure.png" pageId="15" pageNumber="16">Fig. 7E</figureCitation>
due to shrinkage during preparation for SEM), simple, apically with a stout hook and a subapical pointed lobe, in profile strongly reminding of a kind of bottle-opener (e.g., http://www.barleypop.com/best-beer-bottle-opener-period/). A long, slender, curved spine (
<emphasis id="37E5EAB8FFEEFF93DD524CE38C252AD1" box="[546,572,686,711]" italics="true" pageId="15" pageNumber="16">ps</emphasis>
) arising near base of solenomere. Telomere forming a curved, roughly parallel-sided trough, vaguely boat- or pod-shaped, with a subapical spike (
<emphasis id="37E5EAB8FFEEFF93DB5C4C9D8A4C2AFC" box="[1068,1109,720,746]" italics="true" pageId="15" pageNumber="16">tss)</emphasis>
and several complicated lamellae inside the concavity (
<figureCitation id="9DAA2A2FFFEEFF93DD524CBE8C982B1B" box="[546,641,755,781]" captionStart="Fig" captionStartId="14.[189,230,1824,1850]" captionTargetBox="[217,1366,476,1797]" captionTargetId="figure@14.[217,1366,476,1797]" captionTargetPageId="14" captionText="Fig. 7. Utiliverpa decapsulatrix gen. et sp. nov., holotype. AD. Right gonopod coxa. A. Anterior view. B. Mesal view. C. Posterior view. D. Posterior-apical view. EH. Left gonopod telopodite. E. Anterior view. F. Posterior view. G. Mesal-anterior-apical view. H. Solenomere. I. Limbus. Abbreviations: All = apical lateral lobe of coxa, mbp = metaplical bifid process, mf = metaplical flange, mp = metaplica, msp = metaplical spine, prl = proplical lobe, pp = proplica, ps proximal solenomeral spine, slm = solenomere, tlm = telomere, tss = telomeral subapical spike, tt = torsotope. Scales: AH = 0.1 mm; I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3840295" httpUri="https://zenodo.org/record/3840295/files/figure.png" pageId="15" pageNumber="16">Fig. 7G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4D8B6521FFEEFF93DFCD4D798CFF2C22" pageId="15" pageNumber="16" type="distribution">
<paragraph id="052E36AAFFEEFF93DFCD4D798FF72B58" blockId="15.[189,494,820,847]" box="[189,494,820,847]" pageId="15" pageNumber="16">
<heading id="5E6681C6FFEEFF93DFCD4D798FF72B58" bold="true" box="[189,494,820,847]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="37E5EAB8FFEEFF93DFCD4D798FF72B58" bold="true" box="[189,494,820,847]" pageId="15" pageNumber="16">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="052E36AAFFEEFF93DFCD4D298D822B68" blockId="15.[189,923,867,894]" box="[189,923,867,894]" pageId="15" pageNumber="16">
Known only from the
<typeStatus id="DA2A8808FFEEFF93DEB24D298FEC2B68" box="[450,501,868,894]" pageId="15" pageNumber="16">type</typeStatus>
locality. Habitat: forest (under log).
</paragraph>
<paragraph id="052E36AAFFEEFF93DFCD4DE98F872BA8" blockId="15.[189,414,932,958]" box="[189,414,932,958]" pageId="15" pageNumber="16">
<heading id="5E6681C6FFEEFF93DFCD4DE98F872BA8" bold="true" box="[189,414,932,958]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="37E5EAB8FFEEFF93DFCD4DE98F872BA8" bold="true" box="[189,414,932,958]" pageId="15" pageNumber="16">Coexisting species</emphasis>
</heading>
</paragraph>
<paragraph id="052E36AAFFEEFF93DFCD4D998CFF2C22" blockId="15.[189,1399,979,1076]" pageId="15" pageNumber="16">
No other odontopygid species were found in the same sample as the unique
<typeStatus id="DA2A8808FFEEFF93DB154D9E8AD62BFB" box="[1125,1231,979,1005]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, but three are known from Mwanihana Forest Reserve:
<taxonomicName id="C2914D29FFEEFF93DDD14DBB8AB12C06" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[673,1192,1014,1040]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mwanihanensis">
<emphasis id="37E5EAB8FFEEFF93DDD14DBB8DEF2C06" box="[673,1014,1014,1040]" italics="true" pageId="15" pageNumber="16">Chaleponcus mwanihanensis</emphasis>
<bibRefCitation id="61004B5BFFEEFF93DC8D4DBB8AB12C06" author="Enghoff H." box="[1021,1192,1014,1040]" pageId="15" pageNumber="16" pagination="1 - 75" refId="ref5948" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C2914D29FFEEFF93DBC34DBA8F7F2C22" authority="Enghoff, 2015" authorityName="Enghoff" authorityYear="2015" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="major">
<emphasis id="37E5EAB8FFEEFF93DBC34DBA8B6F2C06" box="[1203,1398,1015,1041]" italics="true" pageId="15" pageNumber="16">Aquattuor major</emphasis>
<bibRefCitation id="61004B5BFFEEFF93DFCD4A548F7F2C22" author="Enghoff H. &amp; Frederiksen S. B." box="[189,358,1049,1076]" pageId="15" pageNumber="16" pagination="1 - 25" refId="ref6158" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="C2914D29FFEEFF93DEEF4A578CFB2C22" authority="Enghoff, 2015" authorityName="Enghoff" authorityYear="2015" box="[415,738,1049,1076]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="37E5EAB8FFEEFF93DEEF4A578C362C22" box="[415,559,1050,1076]" italics="true" pageId="15" pageNumber="16">A. submajor</emphasis>
<bibRefCitation id="61004B5BFFEEFF93DD464A548CFB2C22" author="Enghoff H. &amp; Frederiksen S. B." box="[566,738,1049,1076]" pageId="15" pageNumber="16" pagination="1 - 25" refId="ref6158" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>