treatments-xml/data/03/A1/87/03A187FA0D2EBF67FF42DFDAFBD0F82B.xml

<document id="E15A004A8FC7AF148C14B34F6D6FA699" ID-CLB-Dataset="53507" ID-DOI="10.11646/phytotaxa.576.2.8" ID-GBIF-Dataset="e6fb33da-dd97-4631-b047-c31f8359efd8" ID-ISSN="1179-3163" ID-Zenodo-Dep="7461375" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671522154790" checkinUser="plazi" docAuthor="Hammel, Barry &amp; Arias, Salvador" docDate="2022" docId="03A187FA0D2EBF67FF42DFDAFBD0F82B" docLanguage="en" docName="phytotaxa.576.2.8.pdf" docOrigin="Phytotaxa 576 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.576.2.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Deamia funis Hammel &amp; S. Arias 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="225" masterDocId="FF98FF820D2FBF62FFCADE0EFF80FFAA" masterDocTitle="A new species of Deamia (Cactaceae) from Nicaragua" masterLastPageNumber="226" masterPageNumber="220" pageNumber="221" updateTime="1699413736207" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="594825781209CC2606E61DA1956BBA55">A new species of Deamia (Cactaceae) from Nicaragua</mods:title>
</mods:titleInfo>
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<mods:namePart id="06C7C537C9F44E0762F4D78B47F53846">Hammel, Barry</mods:namePart>
<mods:affiliation id="25F2977308892DD8573A05642399DEFC">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110 U. S. A.</mods:affiliation>
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<mods:namePart id="BBCB0CB1460ED8BEEC54DFD0F7A64D13">Arias, Salvador</mods:namePart>
<mods:nameIdentifier id="6D36F71674CC2D02729F1B56DC981F70" type="ORCID">0000-0002-7674-7050</mods:nameIdentifier>
<mods:affiliation id="C5C08E3648A6AE893403F1B7A7B8F5BF">Jardín Botánico, Instituto de Biología, Universidad Nacional Autónoma de México, 04510 México D. F., México sarias @ ib. unam. mx; https: // orcid. org / 0000 - 0002 - 7674 - 7050</mods:affiliation>
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<mods:title id="D9AB5D863DE661EDC7C5CD7C399B4A9C">Phytotaxa</mods:title>
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<mods:date id="0D45888F431C02DA0C20A4DC6023E7A0">2022</mods:date>
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<treatment id="03A187FA0D2EBF67FF42DFDAFBD0F82B" ID-DOI="http://doi.org/10.5281/zenodo.7461377" ID-GBIF-Taxon="206453783" ID-Zenodo-Dep="7461377" LSID="urn:lsid:plazi:treatment:03A187FA0D2EBF67FF42DFDAFBD0F82B" httpUri="http://treatment.plazi.org/id/03A187FA0D2EBF67FF42DFDAFBD0F82B" lastPageId="5" lastPageNumber="225" pageId="1" pageNumber="221">
<subSubSection id="C31265670D2EBF63FF42DFDAFD5FFE44" box="[136,735,468,494]" pageId="1" pageNumber="221" type="nomenclature">
<paragraph id="8BB736EC0D2EBF63FF42DFDAFD5FFE44" blockId="1.[136,735,468,494]" box="[136,735,468,494]" pageId="1" pageNumber="221">
<heading id="D0FF81800D2EBF63FF42DFDAFD5FFE44" box="[136,735,468,494]" fontSize="11" level="2" pageId="1" pageNumber="221" reason="3">
<taxonomicName id="4C084D6F0D2EBF63FF42DFDAFD89FE44" ID-CoL="B3Q8R" authority="Hammel &amp; S. Arias" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[136,521,468,494]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="221" phylum="Tracheophyta" rank="species" species="funis" status="sp. nov.">
<emphasis id="B97CEAFE0D2EBF63FF42DFDAFEA4FE44" bold="true" box="[136,292,468,494]" italics="true" pageId="1" pageNumber="221">Deamia funis</emphasis>
Hammel &amp; S. Arias
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<taxonomicNameLabel id="A24F57850D2EBF63FDC5DFDBFDE4FE44" box="[527,612,469,494]" pageId="1" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13332A690D2EBF63FDB8DFDAFD39FE44" box="[626,697,468,494]" captionStart="FIGURE 1" captionStartId="2.[136,229,2028,2050]" captionTargetBox="[184,1408,199,1998]" captionTargetId="figure-19@2.[173,1414,190,2004]" captionTargetPageId="2" captionText="FIGURE 1. Deamia funis, vegetative characters.A–C. Habitat and habit (Stevens &amp; Montiel 33938). D. Close-up of stem showing cracks in epidermal wax (Stevens &amp; Montiel 39242). Photo credits: all Montiel." figureDoi="http://doi.org/10.5281/zenodo.7461379" httpUri="https://zenodo.org/record/7461379/files/figure.png" pageId="1" pageNumber="221">Figs 1</figureCitation>
, 
<figureCitation id="13332A690D2EBF63FD0DDFDAFD56FE44" box="[711,726,468,494]" captionStart="FIGURE 2" captionStartId="3.[136,229,1992,2014]" captionTargetBox="[216,1371,205,1967]" captionTargetId="figure-20@3.[208,1380,190,1968]" captionTargetPageId="3" captionText="FIGURE 2. Deamia funis, reproductive characters (A–D from Stevens &amp; Montiel 39242, E from Stevens &amp; Calero 44070). A. Side-view of flower on specimen. B. Side-view of flower in life. C. Front-view of flower in life (arrow at stigma). D. Close-up of base of flower in life. E. Submature fruit, in life. Photo credits: A, Hammel; B–D, Montiel; E, Chamorro." figureDoi="http://doi.org/10.5281/zenodo.7461383" httpUri="https://zenodo.org/record/7461383/files/figure.png" pageId="1" pageNumber="221">2</figureCitation>
)
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</paragraph>
</subSubSection>
<subSubSection id="C31265670D2EBF63FF42DFF2FE7BFDF3" pageId="1" pageNumber="221" type="materials_examined">
<paragraph id="8BB736EC0D2EBF63FF42DFF2FE7BFDF3" blockId="1.[136,1452,508,745]" pageId="1" pageNumber="221">
<materialsCitation id="3B603CB10D2EBF63FF42DFF2FE77FDF3" ID-GBIF-Occurrence="3999262305" collectingDate="2017-03-15" collectionCode="N, MO, HULE" collectorName="Stevens &amp; Montiel" country="Nicaragua" county="Type" elevation="423" latitude="12.898055" location="Tierra Blanca" longLatPrecision="21" longitude="-85.504166" municipality="Matiguas" pageId="1" pageNumber="221" specimenCount="1" stateProvince="Matagalpa" typeStatus="holotype">
<collectingCounty id="62D64E600D2EBF63FF42DFF2FF3BFDBB" box="[136,187,508,529]" pageId="1" pageNumber="221">Type</collectingCounty>
:— 
<collectingCountry id="F31F767C0D2EBF63FF1DDFF2FEEAFDBB" box="[215,362,508,529]" name="Nicaragua" pageId="1" pageNumber="221">NICARAGUA</collectingCountry>
. 
<collectingRegion id="49CCF80E0D2EBF63FEBCDFF2FE5FFDB8" box="[374,479,508,530]" country="Nicaragua" name="Matagalpa" pageId="1" pageNumber="221">Matagalpa</collectingRegion>
: 
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<collectionCode id="ED19AE290D2EBF63FDF0DFF2FDCBFDBB" box="[570,587,508,529]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="221" type="Herbarium">N</collectionCode>
of 
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(highway) along road through 
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, 
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<collectionCode id="ED19AE290D2EBF63FB23DFF2FB7AFDBB" box="[1257,1274,508,529]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="221" type="Herbarium">N</collectionCode>
of 
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, deciduous forest, rocky and heavily grazed, 
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, 
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, 
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, 
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(fl), 
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&amp; 
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39242
</emphasis>
(
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, 
<collectionCode id="ED19AE290D2EBF63FEEFDC4AFED3FDF3" box="[293,339,580,601]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="221" type="Herbarium">MO</collectionCode>
!; isotype 
<collectionCode id="ED19AE290D2EBF63FE66DC4AFE70FDF3" box="[428,496,580,601]" country="Nicaragua" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15172" lsid="urn:lsid:biocol.org:col:15172" name="Universidad Nacional Autónoma de Nicaragua" pageId="1" pageNumber="221" type="Herbarium">HULE</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="C31265670D2EBF63FF42DC69FB23FD43" pageId="1" pageNumber="221" type="diagnosis">
<paragraph id="8BB736EC0D2EBF63FF42DC69FB23FD43" blockId="1.[136,1452,508,745]" pageId="1" pageNumber="221">
<emphasis id="B97CEAFE0D2EBF63FF42DC69FF6BFDD7" bold="true" box="[136,235,615,637]" pageId="1" pageNumber="221">Diagnosis</emphasis>
:— 
<emphasis id="B97CEAFE0D2EBF63FEC3DC66FB23FD43" italics="true" pageId="1" pageNumber="221">
In its flowers 
<taxonomicName id="4C084D6F0D2EBF63FE45DC66FD90FDD7" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[399,528,616,637]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="221" phylum="Tracheophyta" rank="species" species="funis">Deamia funis</taxonomicName>
is outwardly similar to 
<taxonomicName id="4C084D6F0D2EBF63FD32DC66FC10FDD7" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[760,912,616,637]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="221" phylum="Tracheophyta" rank="species" species="chontalensis">D. chontalensis</taxonomicName>
, but has stamens (1.5–3 vs. 
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long) and styles (6–8 vs. 
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long) much longer, and with the stigma exerted among or slightly beyond the anthers (vs. the brevistylous flowers of the latter), furthermore, its stems are less than half the diameter of those of the latter, and with more ribs; the stems are similar to those of 
<taxonomicName id="4C084D6F0D2EBF63FEC5DCDAFE1EFD43" box="[271,414,724,745]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="221" phylum="Tracheophyta" rank="species" species="montalvoae">D. montalvoae</taxonomicName>
, but the flowers are smaller, less than 1/2 (mostly 1/3) as long as in that species.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31265670D2EBF63FF42DD16FA1CFA11" pageId="1" pageNumber="221" type="description">
<paragraph id="8BB736EC0D2EBF63FF42DD16FA1CFA11" blockId="1.[136,1452,792,2007]" pageId="1" pageNumber="221">
<emphasis id="B97CEAFE0D2EBF63FF42DD16FE92FC98" bold="true" box="[136,274,792,818]" pageId="1" pageNumber="221">Description</emphasis>
:—Root-climbing plants or trailing epiphytes, with straight spines; stems ± straight and usually pendulous, irregularly branched, the internodes from ca. 0.12 up to ca. 
<quantity id="4CF09B090D2EBF63FCE2DD32FCE2FCFD" box="[808,866,828,855]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="221" unit="m" value="10.0">10 m</quantity>
long, 
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in diameter, ± cylindrical with ca. (6–)8 relatively rounded ribs, with a brittle, transparent, varnish like epidermal coating and abundant adventitious roots, monomorphic; areoles 
<quantity id="4CF09B090D2EBF63FE13DD8AFDC4FC35" box="[473,580,900,927]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="221" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
apart, with 4–18 orange to black, persistent spines, to ca. 
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long, the trichome-spines lacking or a few persistent. Leaves scalelike, to ca. 
<quantity id="4CF09B090D2EBF63FC6BDDA6FC7EFC69" box="[929,1022,936,963]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="221" unit="cm" value="0.04">0.04 cm</quantity>
long, deltate, acute, soon deciduous. Flowers mostly restricted to the distal, new growth of pendulous stems, nocturnal, 
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long, actinomorphic, funnel-shaped, the pericarpel ca. 
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long, with ca. 8 series of indistinct pustules (podaria) bearing the areoles with the bracteole dark reddish brown, ca. 
<quantity id="4CF09B090D2EBF63FDE6DA1AFD02FB85" box="[556,642,1044,1071]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="221" unit="mm" value="0.5">0.5 mm</quantity>
long, deltate, with dense and conspicuous felt and with few or no spines but abundant trichome-spines (hairs); receptacular tube pale green to greenish yellow, sometimes tinged pink, 
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long, funnel shaped (the narrow portion of the tube up to ca. 3 × 
<quantity id="4CF09B090D2EBF63FC95DA52FC2FFBDD" box="[863,943,1116,1143]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="221" unit="cm" value="0.7">0.7 cm</quantity>
, the wider portion up to ca. 3.5 × 
<quantity id="4CF09B090D2EBF63FAE1DA52FAE6FBDC" box="[1323,1382,1116,1142]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="221" unit="cm" value="3.0">3 cm</quantity>
), with numerous scattered bracteoles (sometimes intergrading with or positioned upon the outer tepals), up to ca. 
<quantity id="4CF09B090D2EBF63FAE6DA8EFAECFB31" box="[1324,1388,1152,1179]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="221" unit="mm" value="2.0">2 mm</quantity>
long, the areoles sometimes with a few spines (in the proximal 1/3 to 1/2 of the tube) and with abundant, white to orange trichome-spines to ca. 
<quantity id="4CF09B090D2EBF63FE41DAC6FE59FB49" box="[395,473,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="221" unit="cm" value="1.5">1.5 cm</quantity>
long; perianth with numerous, slightly differentiated tepals, up to 
<quantity id="4CF09B090D2EBF63FB1EDAC6FABEFB49" box="[1236,1342,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="221" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
long, the outer tepals pale yellow green to white, sometimes apically pale brown, ascending, ± oblong, obtuse to acute, the inner tepals white, ascending, oblong, acute; stamens numerous, 
<quantity id="4CF09B090D2EBF63FCD4DB1EFC06FA81" box="[798,902,1296,1323]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="221" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
long, attached in two bands, one in the distal 1/3 of the narrow portion of the tube, to ca. 
<quantity id="4CF09B090D2EBF63FDF7DB3AFD08FAE5" box="[573,648,1332,1359]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="221" unit="cm" value="1.5">1.5 cm</quantity>
wide, the other uniseriate and born at the mouth of the tube (at its widest point), the filaments white, included (within the cup of the limb), the anthers 1–1.4 × ca. 
<quantity id="4CF09B090D2EBF63FBB8DB56FB4BFAD9" box="[1138,1227,1368,1395]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="221" unit="mm" value="0.8">0.8 mm</quantity>
; style to ca. 
<quantity id="4CF09B090D2EBF63FA92DB56FA2BFAD9" box="[1368,1451,1368,1395]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="221" unit="cm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 cm</quantity>
long; stigma white to cream, with ca. 8 simple lobes. Submature fruit yellowish green, ca. 2 × 
<quantity id="4CF09B090D2EBF63FB60DB72FB79FA3D" box="[1194,1273,1404,1431]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="221" unit="cm" value="2.5">2.5 cm</quantity>
, ovoid, smooth, the areoles with ca. 20 spines, to ca. 
<quantity id="4CF09B090D2EBF63FDE8DBAEFDEEFA11" box="[546,622,1440,1467]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="221" unit="cm" value="0.7">0.7 cm</quantity>
long, not completely covering the fruit surface; pulp and seeds not seen.
</paragraph>
</subSubSection>
<subSubSection id="C31265670D2EBF63FF77DBCAFC56F91C" pageId="1" pageNumber="221" type="materials_examined">
<paragraph id="8BB736EC0D2EBF63FF77DBCAFC56F91C" blockId="1.[136,1452,792,2007]" pageId="1" pageNumber="221">
<emphasis id="B97CEAFE0D2EBF63FF77DBCAFDB9FA74" bold="true" box="[189,569,1476,1502]" pageId="1" pageNumber="221">Additional specimens examined</emphasis>
:— 
<materialsCitation id="3B603CB10D2EBF63FD90DBCAFF42F98C" ID-GBIF-Occurrence="3999262304" collectingDate="2009-01-20" collectionCode="NE, HULE, MO" collectorName="Stevens &amp; Montiel" country="Nicaragua" elevation="355" latitude="12.280833" location="Cerro Oluma" longLatPrecision="21" longitude="-85.37417" pageId="1" pageNumber="221" specimenCount="1" stateProvince="Chontales">
<collectingCountry id="F31F767C0D2EBF63FD90DBCAFC89FA74" box="[602,777,1476,1502]" name="Nicaragua" pageId="1" pageNumber="221">NICARAGUA</collectingCountry>
. 
<collectingRegion id="49CCF80E0D2EBF63FCDEDBCAFC08FA74" box="[788,904,1476,1502]" country="Nicaragua" name="Chontales" pageId="1" pageNumber="221">Chontales</collectingRegion>
: 
<locationDeviation id="9A052FCE0D2EBF63FC5EDBCAFBEBFA74" box="[916,1131,1476,1503]" location="Cuapa" pageId="1" pageNumber="221">
<quantity id="4CF09B090D2EBF63FC5EDBCAFC4EFA74" box="[916,974,1476,1502]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="221" unit="km" value="2.0">2 km</quantity>
<collectionCode id="ED19AE290D2EBF63FC1CDBCBFC7BFA75" box="[982,1019,1477,1503]" country="Australia" lsid="urn:lsid:biocol.org:col:14575" name="University of New England" pageId="1" pageNumber="221" type="Herbarium">NE</collectionCode>
of Cuapa
</locationDeviation>
along road to 
<location id="8ED760370D2EBF63FADADBCAFA27FA74" LSID="urn:lsid:plazi:treatment:03A187FA0D2EBF67FF42DFDAFBD0F82B:8ED760370D2EBF63FADADBCAFA27FA74" box="[1296,1447,1476,1502]" country="Nicaragua" latitude="12.280833" longLatPrecision="21" longitude="-85.37417" name="Cerro Oluma" pageId="1" pageNumber="221" stateProvince="Chontales">Cerro Oluma</location>
, dry forest patch on hillside, 
<geoCoordinate id="EE3C502B0D2EBF63FE76DBE6FDC8F9A9" box="[444,584,1512,1539]" degrees="12" direction="north" minutes="16" orientation="latitude" pageId="1" pageNumber="221" precision="15" seconds="51" value="12.280833">12°16’51”N</geoCoordinate>
, 
<geoCoordinate id="EE3C502B0D2EBF63FD98DBE6FD62F9A9" box="[594,738,1512,1539]" degrees="85" direction="west" minutes="22" orientation="longitude" pageId="1" pageNumber="221" precision="15" seconds="27" value="-85.37417">85°22’27”W</geoCoordinate>
, 
<quantity id="4CF09B090D2EBF63FD26DBE6FCB4F9A9" box="[748,820,1512,1539]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="1" pageNumber="221" unit="m" value="355.0">
<elevation id="0025D1DF0D2EBF63FD26DBE6FCB4F9A9" box="[748,820,1512,1539]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="1" pageNumber="221" unit="m" value="355.0">355 m</elevation>
</quantity>
, 
<date id="FFB6102C0D2EBF63FCF4DBE6FC75F9A8" box="[830,1013,1512,1539]" pageId="1" pageNumber="221" value="2009-01-20">
<collectingDate id="EFF2E9C40D2EBF63FCF4DBE6FC75F9A8" box="[830,1013,1512,1539]" pageId="1" pageNumber="221" value="2009-01-20">20 January 2009</collectingDate>
</date>
(fl.), 
<emphasis id="B97CEAFE0D2EBF63FBF8DBE6FACBF9A8" box="[1074,1355,1512,1538]" italics="true" pageId="1" pageNumber="221">
<collectorName id="26FD533A0D2EBF63FBF8DBE6FB06F9A8" box="[1074,1158,1512,1538]" pageId="1" pageNumber="221">Stevens</collectorName>
&amp; 
<collectorName id="26FD533A0D2EBF63FB62DBE6FB7FF9A8" box="[1192,1279,1512,1538]" pageId="1" pageNumber="221">Montiel</collectorName>
27909
</emphasis>
(
<collectionCode id="ED19AE290D2EBF63FA92DBE6FA27F9A8" box="[1368,1447,1512,1538]" country="Nicaragua" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15172" lsid="urn:lsid:biocol.org:col:15172" name="Universidad Nacional Autónoma de Nicaragua" pageId="1" pageNumber="221" type="Herbarium">HULE</collectionCode>
, 
<collectionCode id="ED19AE290D2EBF63FF42D802FF3AF98C" box="[136,186,1548,1574]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="221" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B603CB10D2EBF63FF07D802FBE0F9E0" ID-GBIF-Occurrence="3999262306" collectingDate="2018-04-03" collectionCode="N, HULE, MO" collectorName="Stevens" country="Nicaragua" elevation="322" latitude="12.739722" location="San Dionysio" longLatPrecision="21" longitude="-85.78778" municipality="Puente Kaulapa" pageId="1" pageNumber="221" specimenCount="1" stateProvince="Matagalpa">
<collectingRegion id="49CCF80E0D2EBF63FF07D802FEC8F98C" box="[205,328,1548,1574]" country="Nicaragua" name="Matagalpa" pageId="1" pageNumber="221">Matagalpa</collectingRegion>
: 
<locationDeviation id="9A052FCE0D2EBF63FE99D802FDECF98C" box="[339,620,1548,1575]" location="Esquipulas" pageId="1" pageNumber="221">
<quantity id="4CF09B090D2EBF63FE99D802FE2FF98C" box="[339,431,1548,1574]" metricMagnitude="4" metricUnit="m" metricValue="1.19" pageId="1" pageNumber="221" unit="km" value="11.9">11.9 km</quantity>
<collectionCode id="ED19AE290D2EBF63FE7CD803FE4AF98D" box="[438,458,1549,1575]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="221" type="Herbarium">N</collectionCode>
of Esquipulas
</locationDeviation>
(at 
<collectingMunicipality id="6BD3AC960D2EBF63FD53D803FCCCF98C" box="[665,844,1548,1575]" pageId="1" pageNumber="221">Puente Kaulapa</collectingMunicipality>
) along road to 
<location id="8ED760370D2EBF63FC32D802FB14F98C" LSID="urn:lsid:plazi:treatment:03A187FA0D2EBF67FF42DFDAFBD0F82B:8ED760370D2EBF63FC32D802FB14F98C" box="[1016,1172,1548,1574]" country="Nicaragua" latitude="12.739722" longLatPrecision="21" longitude="-85.78778" municipality="Puente Kaulapa" name="San Dionysio" pageId="1" pageNumber="221" stateProvince="Matagalpa">San Dionysio</location>
; dry forest, 
<geoCoordinate id="EE3C502B0D2EBF63FAD1D802FA27F98C" box="[1307,1447,1548,1574]" degrees="12" direction="north" minutes="44" orientation="latitude" pageId="1" pageNumber="221" precision="15" seconds="23" value="12.739722">12°44’23”N</geoCoordinate>
, 
<geoCoordinate id="EE3C502B0D2EBF63FF42D83EFE97F9E1" box="[136,279,1584,1611]" degrees="85" direction="west" minutes="47" orientation="longitude" pageId="1" pageNumber="221" precision="15" seconds="16" value="-85.78778">85°47’16”W</geoCoordinate>
, 
<quantity id="4CF09B090D2EBF63FEECD83EFEF1F9E1" box="[294,369,1584,1611]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="1" pageNumber="221" unit="m" value="322.0">
<elevation id="0025D1DF0D2EBF63FEECD83EFEF1F9E1" box="[294,369,1584,1611]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="1" pageNumber="221" unit="m" value="322.0">322 m</elevation>
</quantity>
, 
<date id="FFB6102C0D2EBF63FE4AD83EFD93F9E0" box="[384,531,1584,1610]" pageId="1" pageNumber="221" value="2018-04-03">
<collectingDate id="EFF2E9C40D2EBF63FE4AD83EFD93F9E0" box="[384,531,1584,1610]" pageId="1" pageNumber="221" value="2018-04-03">3 April 2018</collectingDate>
</date>
(fl., immature fruit.), 
<emphasis id="B97CEAFE0D2EBF63FCDFD83EFC38F9E0" box="[789,952,1584,1610]" italics="true" pageId="1" pageNumber="221">
<collectorName id="26FD533A0D2EBF63FCDFD83EFCE9F9E0" box="[789,873,1584,1610]" pageId="1" pageNumber="221">Stevens</collectorName>
40324
</emphasis>
(
<collectionCode id="ED19AE290D2EBF63FC00D83EFB99F9E0" box="[970,1049,1584,1610]" country="Nicaragua" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15172" lsid="urn:lsid:biocol.org:col:15172" name="Universidad Nacional Autónoma de Nicaragua" pageId="1" pageNumber="221" type="Herbarium">HULE</collectionCode>
, 
<collectionCode id="ED19AE290D2EBF63FBEDD83EFBD9F9E0" box="[1063,1113,1584,1610]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="221" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
; ibd., 
<materialsCitation id="3B603CB10D2EBF63FB60D83EFDFDF9C4" ID-GBIF-Occurrence="3999262303" collectionCode="N" country="Nicaragua" location="Tierra Blanca" pageId="1" pageNumber="221" specimenCount="1" stateProvince="Matagalpa">
<locationDeviation id="9A052FCE0D2EBF63FB60D83EFA2BF9E0" box="[1194,1451,1584,1611]" location="Matiguas" pageId="1" pageNumber="221">
<quantity id="4CF09B090D2EBF63FB60D83EFB7BF9E0" box="[1194,1275,1584,1610]" metricMagnitude="3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="221" unit="km" value="9.1">9.1 km</quantity>
<collectionCode id="ED19AE290D2EBF63FACED83FFA98F9E1" box="[1284,1304,1585,1611]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="221" type="Herbarium">N</collectionCode>
of Matiguás
</locationDeviation>
(highway) along road through 
<location id="8ED760370D2EBF63FE2BD85AFDFDF9C4" LSID="urn:lsid:plazi:treatment:03A187FA0D2EBF67FF42DFDAFBD0F82B:8ED760370D2EBF63FE2BD85AFDFDF9C4" box="[481,637,1620,1646]" country="Nicaragua" name="Tierra Blanca" pageId="1" pageNumber="221" stateProvince="Matagalpa">Tierra Blanca</location>
</materialsCitation>
, 
<materialsCitation id="3B603CB10D2EBF63FD42D85AFB58F938" ID-GBIF-Occurrence="3999262301" collectingDate="2013-01-04" collectionCode="N, HULE, MO" collectorName="Stevens &amp; Montiel" country="Nicaragua" elevation="423" latitude="12.898055" location="Tierra Blanca" longLatPrecision="21" longitude="-85.504166" pageId="1" pageNumber="221" specimenCount="1" stateProvince="Matagalpa">
<quantity id="4CF09B090D2EBF63FD42D85AFD57F9C4" box="[648,727,1620,1646]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="221" unit="km" value="2.9">2.9 km</quantity>
<collectionCode id="ED19AE290D2EBF63FD14D85BFD72F9C5" box="[734,754,1621,1647]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="221" type="Herbarium">N</collectionCode>
of 
<location id="8ED760370D2EBF63FCD3D85AFC35F9C4" LSID="urn:lsid:plazi:treatment:03A187FA0D2EBF67FF42DFDAFBD0F82B:8ED760370D2EBF63FCD3D85AFC35F9C4" box="[793,949,1620,1646]" country="Nicaragua" latitude="12.898055" longLatPrecision="21" longitude="-85.504166" name="Tierra Blanca" pageId="1" pageNumber="221" stateProvince="Matagalpa">Tierra Blanca</location>
, deciduous forest, rocky and heavily grazed, 
<geoCoordinate id="EE3C502B0D2EBF63FF42D876FE94F939" box="[136,276,1656,1683]" degrees="12" direction="north" minutes="53" orientation="latitude" pageId="1" pageNumber="221" precision="15" seconds="53" value="12.898055">12°53’53”N</geoCoordinate>
, 
<geoCoordinate id="EE3C502B0D2EBF63FEE8D876FE31F939" box="[290,433,1656,1683]" degrees="85" direction="west" minutes="30" orientation="longitude" pageId="1" pageNumber="221" precision="15" seconds="15" value="-85.504166">85°30’15”W</geoCoordinate>
, 
<quantity id="4CF09B090D2EBF63FE75D876FD8AF939" box="[447,522,1656,1683]" metricMagnitude="2" metricUnit="m" metricValue="4.2299999999999995" pageId="1" pageNumber="221" unit="m" value="423.0">
<elevation id="0025D1DF0D2EBF63FE75D876FD8AF939" box="[447,522,1656,1683]" metricMagnitude="2" metricUnit="m" metricValue="4.2299999999999995" pageId="1" pageNumber="221" unit="m" value="423.0">423 m</elevation>
</quantity>
, 
<date id="FFB6102C0D2EBF63FDD3D876FD48F938" box="[537,712,1656,1683]" pageId="1" pageNumber="221" value="2013-01-04">
<collectingDate id="EFF2E9C40D2EBF63FDD3D876FD48F938" box="[537,712,1656,1683]" pageId="1" pageNumber="221" value="2013-01-04">4 January 2013</collectingDate>
</date>
(fl.), 
<emphasis id="B97CEAFE0D2EBF63FCC7D876FBB0F938" box="[781,1072,1656,1682]" italics="true" pageId="1" pageNumber="221">
<collectorName id="26FD533A0D2EBF63FCC7D876FCE1F938" box="[781,865,1656,1682]" pageId="1" pageNumber="221">Stevens</collectorName>
&amp; 
<collectorName id="26FD533A0D2EBF63FC40D876FC61F938" box="[906,993,1656,1682]" pageId="1" pageNumber="221">Montiel</collectorName>
33938
</emphasis>
(
<collectionCode id="ED19AE290D2EBF63FB8BD876FB10F938" box="[1089,1168,1656,1682]" country="Nicaragua" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15172" lsid="urn:lsid:biocol.org:col:15172" name="Universidad Nacional Autónoma de Nicaragua" pageId="1" pageNumber="221" type="Herbarium">HULE</collectionCode>
, 
<collectionCode id="ED19AE290D2EBF63FB54D876FB50F938" box="[1182,1232,1656,1682]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="221" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B603CB10D2EBF63FB2CD876FC51F91C" ID-GBIF-Occurrence="3999262302" collectingDate="2022-02-25" collectionCode="HULE, MO" collectorName="Stevens &amp; Calero" country="Nicaragua" pageId="1" pageNumber="221" specimenCount="1" stateProvince="Matagalpa">
ibd., 
<date id="FFB6102C0D2EBF63FAEBD876FF40F91C" pageId="1" pageNumber="221" value="2022-02-25">
<collectingDate id="EFF2E9C40D2EBF63FAEBD876FF40F91C" pageId="1" pageNumber="221" value="2022-02-25">25 February 2022</collectingDate>
</date>
(flower bud, immature fruits), 
<emphasis id="B97CEAFE0D2EBF63FDD7D892FCAFF91C" box="[541,815,1692,1718]" italics="true" pageId="1" pageNumber="221">
<collectorName id="26FD533A0D2EBF63FDD7D892FDF1F91C" box="[541,625,1692,1718]" pageId="1" pageNumber="221">Stevens</collectorName>
&amp; 
<collectorName id="26FD533A0D2EBF63FD5FD892FD62F91C" box="[661,738,1692,1718]" pageId="1" pageNumber="221">Calero</collectorName>
44070
</emphasis>
(
<collectionCode id="ED19AE290D2EBF63FCF7D892FC0CF91C" box="[829,908,1692,1718]" country="Nicaragua" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15172" lsid="urn:lsid:biocol.org:col:15172" name="Universidad Nacional Autónoma de Nicaragua" pageId="1" pageNumber="221" type="Herbarium">HULE</collectionCode>
, 
<collectionCode id="ED19AE290D2EBF63FC52D892FC4AF91C" box="[920,970,1692,1718]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="221" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31265670D2EBF63FF77D8CEFE16F818" pageId="1" pageNumber="221" type="distribution">
<paragraph id="8BB736EC0D2EBF63FF77D8CEFE16F818" blockId="1.[136,1452,792,2007]" pageId="1" pageNumber="221">
<emphasis id="B97CEAFE0D2EBF63FF77D8CEFE62F970" bold="true" box="[189,482,1728,1754]" pageId="1" pageNumber="221">Distribution and habitat</emphasis>
:— 
<taxonomicName id="4C084D6F0D2EBF63FDCFD8CFFD1CF970" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[517,668,1728,1754]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="221" phylum="Tracheophyta" rank="species" species="funis">
<emphasis id="B97CEAFE0D2EBF63FDCFD8CFFD1CF970" box="[517,668,1728,1754]" italics="true" pageId="1" pageNumber="221">Deamia funis</emphasis>
</taxonomicName>
is known only from low to mid elevations (
<quantity id="4CF09B090D2EBF63FB46D8CEFA8EF970" box="[1164,1294,1728,1754]" metricMagnitude="2" metricUnit="m" metricValue="3.725" metricValueMax="4.2299999999999995" metricValueMin="3.22" pageId="1" pageNumber="221" unit="m" value="372.5" valueMax="423.0" valueMin="322.0">322–423 m</quantity>
) of fog-zones of dry, seasonal forests on the Atlantic watershed, from central 
<collectingCountry id="F31F767C0D2EBF63FC85D8EAFC46F954" box="[847,966,1764,1790]" name="Nicaragua" pageId="1" pageNumber="221">Nicaragua</collectingCountry>
. Although presently endemic to 
<collectingCountry id="F31F767C0D2EBF63FAFBD8EAFA27F954" box="[1329,1447,1764,1790]" name="Nicaragua" pageId="1" pageNumber="221">Nicaragua</collectingCountry>
, the species should be sought in similar habitat just to the north, in 
<collectingCountry id="F31F767C0D2EBF63FCACD906FC57F888" box="[870,983,1800,1826]" name="Honduras" pageId="1" pageNumber="221">Honduras</collectingCountry>
. 
<taxonomicName id="4C084D6F0D2EBF63FC2AD907FB10F888" authorityName="Britton &amp; Rose" authorityYear="1920" baseAuthorityName="Karw. ex Zuccarini" baseAuthorityYear="1837" box="[992,1168,1800,1826]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="221" phylum="Tracheophyta" rank="species" species="testudo">
<emphasis id="B97CEAFE0D2EBF63FC2AD907FB10F888" box="[992,1168,1800,1826]" italics="true" pageId="1" pageNumber="221">Deamia testudo</emphasis>
</taxonomicName>
, the only other species in the genus known from 
<collectingCountry id="F31F767C0D2EBF63FE46D922FE7FF8EC" box="[396,511,1836,1862]" name="Nicaragua" pageId="1" pageNumber="221">Nicaragua</collectingCountry>
is more widespread, and is known to occur sympatrically, at least as close as ca. 
<quantity id="4CF09B090D2EBF63FA45D922FF2CF8C0" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="221" unit="km" value="25.0">25 km</quantity>
(
<figureCitation id="13332A690D2EBF63FF70D95EFF7EF8C0" box="[186,254,1872,1898]" captionStart="FIGURE 3" captionStartId="4.[136,229,1294,1316]" captionTargetBox="[157,1430,203,1267]" captionTargetId="figure-19@4.[151,1436,190,1270]" captionTargetPageId="4" captionText="FIGURE 3. Deamia funis and D. testudo, distribution map. Total distribution of D. funis (red balloons); Nicaraguan distribution of D. testudo (yellow balloons). Map generated from specimens on Tropicos (Google Maps)." figureDoi="http://doi.org/10.5281/zenodo.7461387" httpUri="https://zenodo.org/record/7461387/files/figure.png" pageId="1" pageNumber="221">Fig. 3</figureCitation>
). Several sterile collections from limestone outcrops in a mixture of moist forest and savana of the 
<collectingRegion id="49CCF80E0D2EBF63FA89D95EFE98F824" country="Nicaragua" name="Atlantico Norte" pageId="1" pageNumber="221">Atlántico Norte region</collectingRegion>
(northeastern 
<collectingCountry id="F31F767C0D2EBF63FE77D97AFDB5F824" box="[445,565,1908,1934]" name="Nicaragua" pageId="1" pageNumber="221">Nicaragua</collectingCountry>
), may belong to this species, but lacking fertile collections from that area, we have not included them.
</paragraph>
</subSubSection>
<subSubSection id="C31265670D2EBF63FF77D9B2FA12F87D" box="[189,1426,1980,2007]" pageId="1" pageNumber="221" type="biology_ecology">
<paragraph id="8BB736EC0D2EBF63FF77D9B2FA12F87D" blockId="1.[136,1452,792,2007]" box="[189,1426,1980,2007]" pageId="1" pageNumber="221">
<emphasis id="B97CEAFE0D2EBF63FF77D9B2FEB9F87C" bold="true" box="[189,313,1980,2006]" pageId="1" pageNumber="221">Phenology</emphasis>
:—Flowering plants have been found only in the months of January to April, during the dry season.
</paragraph>
</subSubSection>
<caption id="DF7766640D2DBF60FF42D9E2FCC0F78D" ID-DOI="http://doi.org/10.5281/zenodo.7461379" ID-Zenodo-Dep="7461379" httpUri="https://zenodo.org/record/7461379/files/figure.png" pageId="2" pageNumber="222" startId="2.[136,229,2028,2050]" targetBox="[184,1408,199,1998]" targetPageId="2">
<paragraph id="8BB736EC0D2DBF60FF42D9E2FCC0F78D" blockId="2.[136,1451,2028,2087]" pageId="2" pageNumber="222">
<emphasis id="B97CEAFE0D2DBF60FF42D9E2FF77F7AB" bold="true" box="[136,247,2028,2050]" pageId="2" pageNumber="222">FIGURE 1</emphasis>
. 
<taxonomicName id="4C084D6F0D2DBF60FEC8D9E3FE03F7A8" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[258,387,2029,2050]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="222" phylum="Tracheophyta" rank="species" species="funis">
<emphasis id="B97CEAFE0D2DBF60FEC8D9E3FE03F7A8" box="[258,387,2029,2050]" italics="true" pageId="2" pageNumber="222">Deamia funis</emphasis>
</taxonomicName>
, vegetative characters. A–C. Habitat and habit (
<emphasis id="B97CEAFE0D2DBF60FC86D9E3FBBEF7A8" box="[844,1086,2029,2050]" italics="true" pageId="2" pageNumber="222">Stevens &amp; Montiel 33938</emphasis>
). D. Close-up of stem showing cracks in epidermal wax (
<emphasis id="B97CEAFE0D2DBF60FEF7D61FFDB0F78C" box="[317,560,2065,2086]" italics="true" pageId="2" pageNumber="222">Stevens &amp; Montiel 39242</emphasis>
). Photo credits: all Montiel.
</paragraph>
</caption>
<caption id="DF7766640D2CBF61FF42D9C6FC52F78C" ID-DOI="http://doi.org/10.5281/zenodo.7461383" ID-Zenodo-Dep="7461383" httpUri="https://zenodo.org/record/7461383/files/figure.png" pageId="3" pageNumber="223" startId="3.[136,229,1992,2014]" targetBox="[216,1371,205,1967]" targetPageId="3">
<paragraph id="8BB736EC0D2CBF61FF42D9C6FC52F78C" blockId="3.[136,1452,1992,2086]" pageId="3" pageNumber="223">
<emphasis id="B97CEAFE0D2CBF61FF42D9C6FF77F877" bold="true" box="[136,247,1992,2014]" pageId="3" pageNumber="223">FIGURE 2</emphasis>
. 
<taxonomicName id="4C084D6F0D2CBF61FEC9D9C7FE04F877" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[259,388,1992,2014]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="223" phylum="Tracheophyta" rank="species" species="funis">
<emphasis id="B97CEAFE0D2CBF61FEC9D9C7FE04F877" box="[259,388,1992,2014]" italics="true" pageId="3" pageNumber="223">Deamia funis</emphasis>
</taxonomicName>
, reproductive characters (A–D from 
<emphasis id="B97CEAFE0D2CBF61FD22D9C7FC5BF877" box="[744,987,1992,2014]" italics="true" pageId="3" pageNumber="223">Stevens &amp; Montiel 39242</emphasis>
, E from 
<emphasis id="B97CEAFE0D2CBF61FBFAD9C7FA9AF877" box="[1072,1306,1992,2014]" italics="true" pageId="3" pageNumber="223">Stevens &amp; Calero 44070</emphasis>
). A. Side-view of flower on specimen. B. Side-view of flower in life. C. Front-view of flower in life (arrow at stigma). D. Close-up of base of flower in life. E. Submature fruit, in life. Photo credits: A, Hammel; B–D, Montiel; E, Chamorro.
</paragraph>
</caption>
<caption id="DF7766640D2BBF66FF42DB00FC4DFAED" ID-DOI="http://doi.org/10.5281/zenodo.7461387" ID-Zenodo-Dep="7461387" httpUri="https://zenodo.org/record/7461387/files/figure.png" pageId="4" pageNumber="224" startId="4.[136,229,1294,1316]" targetBox="[157,1430,203,1267]" targetPageId="4">
<paragraph id="8BB736EC0D2BBF66FF42DB00FC4DFAED" blockId="4.[136,1452,1294,1352]" pageId="4" pageNumber="224">
<emphasis id="B97CEAFE0D2BBF66FF42DB00FF79FA89" bold="true" box="[136,249,1294,1316]" pageId="4" pageNumber="224">FIGURE 3</emphasis>
. 
<taxonomicName id="4C084D6F0D2BBF66FECCDB01FE09FA89" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[262,393,1294,1316]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="funis">
<emphasis id="B97CEAFE0D2BBF66FECCDB01FE09FA89" box="[262,393,1294,1316]" italics="true" pageId="4" pageNumber="224">Deamia funis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C084D6F0D2BBF66FE70DB01FD9EFA89" authorityName="Britton &amp; Rose" authorityYear="1920" baseAuthorityName="Karw. ex Zuccarini" baseAuthorityYear="1837" box="[442,542,1294,1316]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="testudo">
<emphasis id="B97CEAFE0D2BBF66FE70DB01FD9EFA89" box="[442,542,1294,1316]" italics="true" pageId="4" pageNumber="224">D. testudo</emphasis>
</taxonomicName>
, distribution map. Total distribution of 
<taxonomicName id="4C084D6F0D2BBF66FC68DB01FC70FA89" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[930,1008,1294,1316]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="funis">
<emphasis id="B97CEAFE0D2BBF66FC68DB01FC70FA89" box="[930,1008,1294,1316]" italics="true" pageId="4" pageNumber="224">D. funis</emphasis>
</taxonomicName>
(red balloons); Nicaraguan distribution of 
<taxonomicName id="4C084D6F0D2BBF66FA5EDB01FF4DFAED" authorityName="Britton &amp; Rose" authorityYear="1920" baseAuthorityName="Karw. ex Zuccarini" baseAuthorityYear="1837" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="testudo">
<emphasis id="B97CEAFE0D2BBF66FA5EDB01FF4DFAED" italics="true" pageId="4" pageNumber="224">D. testudo</emphasis>
</taxonomicName>
(yellow balloons). Map generated from specimens on Tropicos (Google Maps).
</paragraph>
</caption>
<subSubSection id="C31265670D2BBF66FF77DB79FDCDFA1F" pageId="4" pageNumber="224" type="etymology">
<paragraph id="8BB736EC0D2BBF66FF77DB79FDCDFA1F" blockId="4.[136,1452,1398,2037]" pageId="4" pageNumber="224">
<emphasis id="B97CEAFE0D2BBF66FF77DB79FEBEFA3B" bold="true" box="[189,318,1399,1425]" pageId="4" pageNumber="224">Etymology</emphasis>
:—The epithet refers to the long, dangling, rope-like stems common in this species when found growing on horizontal branches of trees.
</paragraph>
</subSubSection>
<subSubSection id="C31265670D2BBF66FF77DBB0FD28F952" pageId="4" pageNumber="224" type="conservation">
<paragraph id="8BB736EC0D2BBF66FF77DBB0FD28F952" blockId="4.[136,1452,1398,2037]" pageId="4" pageNumber="224">
<emphasis id="B97CEAFE0D2BBF66FF77DBB0FDBDFA73" bold="true" box="[189,573,1470,1497]" pageId="4" pageNumber="224">Preliminary conservation status</emphasis>
:—The new species is not known for sure to occur within any protected area, but the 
<typeStatus id="54B3884E0D2BBF66FF14DBEDFE8EFA57" box="[222,270,1507,1533]" pageId="4" pageNumber="224">type</typeStatus>
and two other collections were found within a few km of the Reserva Natural Sierra Quirragua and one other collection from very near the Reserva Natural Sierra Amerrisque. The species has been collected from only three different localities and we estimate that a polygon encompassing all three of them covers an area of no more than 
<quantity id="4CF09B090D2BBF66FF42D840FF6CF9C2" box="[136,236,1614,1641]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="224" unit="km" value="5000.0">5000 km</quantity>
<superScript id="7C7D9BA40D2BBF66FF26D840FF74F9F6" attach="left" box="[236,244,1614,1628]" fontSize="6" pageId="4" pageNumber="224">2</superScript>
. We consider that the area where this plant is known to occur, being subject to cattle farming, suffers from continuing habitat destruction. Thus, by the guidelines for using the IUCN Red List categories and criteria (2022), table 2.1, B1 a and b, this species ought to be considered endangered (EN). In any case, all species of plants everywhere, except common weeds and cultivated plants should be considered endangered. Human caused habitat destruction and climate change are not making things any better.
</paragraph>
</subSubSection>
<subSubSection id="C31265670D2BBF67FF77D90CFBD0F82B" lastPageId="5" lastPageNumber="225" pageId="4" pageNumber="224" type="discussion">
<paragraph id="8BB736EC0D2BBF66FF77D90CFEE3F85F" blockId="4.[136,1452,1398,2037]" pageId="4" pageNumber="224">
<emphasis id="B97CEAFE0D2BBF66FF77D90CFEBBF8B6" bold="true" box="[189,315,1794,1820]" pageId="4" pageNumber="224">Discussion</emphasis>
:—By its stems 
<quantity id="4CF09B090D2BBF66FE3BD90CFDEFF8B7" box="[497,623,1794,1821]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="4" pageNumber="224" unit="cm" value="0.85" valueMax="1.1" valueMin="0.6">0.6–1.1 cm</quantity>
in diameter with ca. (6–)8 ribs (
<emphasis id="B97CEAFE0D2BBF66FC15D90DFC7DF8B6" box="[991,1021,1795,1820]" italics="true" pageId="4" pageNumber="224">vs.</emphasis>
stems 
<quantity id="4CF09B090D2BBF66FB86D90DFB20F8B7" box="[1100,1184,1795,1821]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="224" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
in diameter with 7 or 8 ribs) this new species overlaps somewhat with those of 
<emphasis id="B97CEAFE0D2BBF66FD25D929FC57F8EA" box="[751,983,1830,1856]" italics="true" pageId="4" pageNumber="224">
<taxonomicName id="4C084D6F0D2BBF66FD25D929FC54F8EA" box="[751,980,1830,1856]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="montalvoae">Deamia montalvoae</taxonomicName>
,
</emphasis>
but in floral characters it differs markedly from that species by its much shorter flowers [8–10 
<emphasis id="B97CEAFE0D2BBF66FD19D945FD71F8CE" box="[723,753,1867,1892]" italics="true" pageId="4" pageNumber="224">vs.</emphasis>
(23–) 
<quantity id="4CF09B090D2BBF66FCF9D945FC25F8CF" box="[819,933,1867,1893]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="4" pageNumber="224" unit="cm" value="28.5" valueMax="30.0" valueMin="27.0">27–30 cm</quantity>
long in 
<taxonomicName id="4C084D6F0D2BBF66FBC8D945FB28F8CE" box="[1026,1192,1866,1892]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="montalvoae">
<emphasis id="B97CEAFE0D2BBF66FBC8D945FB28F8CE" box="[1026,1192,1866,1892]" italics="true" pageId="4" pageNumber="224">D. montalvoae</emphasis>
</taxonomicName>
]. However, in size and overall 
<taxonomicName id="4C084D6F0D2BBF66FF14D960FEBEF822" box="[222,318,1902,1928]" form="the" pageId="4" pageNumber="224" rank="form">form the</taxonomicName>
funnel-shaped flowers of the present species are almost identical to those of 
<taxonomicName id="4C084D6F0D2BBF66FB63D961FAD8F822" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[1193,1368,1902,1928]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2BBF66FB63D961FAD8F822" box="[1193,1368,1902,1928]" italics="true" pageId="4" pageNumber="224">D. chontalensis</emphasis>
</taxonomicName>
, which however, has stems 
<quantity id="4CF09B090D2BBF66FEACD99CFE63F807" box="[358,483,1938,1965]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="4" pageNumber="224" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 cm</quantity>
in diameter with only 5 or 6(–7) acute ribs. The fruits of 
<taxonomicName id="4C084D6F0D2BBF66FB91D99DFB32F807" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[1115,1202,1939,1965]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="funis">
<emphasis id="B97CEAFE0D2BBF66FB91D99DFB32F807" box="[1115,1202,1939,1965]" italics="true" pageId="4" pageNumber="224">D. funis</emphasis>
</taxonomicName>
differ from both latter two species by the surface being hardly at all covered by the areole spines, and in that respect are more similar to the fruits of 
<taxonomicName id="4C084D6F0D2BBF66FF22D9D5FEDCF85E" authorityName="Britton &amp; Rose" authorityYear="1920" baseAuthorityName="Karw. ex Zuccarini" baseAuthorityYear="1837" box="[232,348,2010,2036]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="224" phylum="Tracheophyta" rank="species" species="testudo">
<emphasis id="B97CEAFE0D2BBF66FF22D9D5FEDCF85E" box="[232,348,2010,2036]" italics="true" pageId="4" pageNumber="224">D. testudo</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB736EC0D2ABF67FF77DE90FC83FCEA" blockId="5.[136,1452,158,977]" pageId="5" pageNumber="225">
The flowers of 
<taxonomicName id="4C084D6F0D2ABF67FEA3DE91FDD4FF12" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[361,596,158,184]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2ABF67FEA3DE91FDD4FF12" box="[361,596,158,184]" italics="true" pageId="5" pageNumber="225">Deamia chontalensis</emphasis>
</taxonomicName>
itself are unique among species of the genus, and at least uncommon among all 
<taxonomicName id="4C084D6F0D2ABF67FF60DECCFE9FFF76" box="[170,287,194,220]" class="Magnoliopsida" family="Cactaceae" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="family">Cactaceae</taxonomicName>
.An illustration showing the three main floral types in 
<taxonomicName id="4C084D6F0D2ABF67FCB8DECCFC63FF76" box="[882,995,194,220]" class="Magnoliopsida" family="Cactaceae" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="family">Cactaceae</taxonomicName>
in the review by 
<bibRefCitation id="EF994B1D0D2ABF67FB55DECCFA2CFF76" author="Mandujano, M. &amp; Carrillo-Angeles, I. &amp; Martinez-Peralta, C. &amp; Golubov, J." box="[1183,1452,194,221]" pageId="5" pageNumber="225" pagination="197 - 230" refId="ref3523" refString="Mandujano, M. d. C., Carrillo-Angeles, I., Martinez-Peralta, C. &amp; Golubov, J. (2010) Reproductive biology of Cactaceae. In: Ramawat, K. (Ed.) Desert plants: biology and biotechnology. Springer, Berlin, Heidelberg, pp. 197 - 230. https: // doi. org / 10.1007 / 978 - 3 - 642 - 02550 - 1 _ 10" type="book chapter" year="2010">
Mandujano 
<emphasis id="B97CEAFE0D2ABF67FAEFDECDFAD4FF76" box="[1317,1364,194,220]" italics="true" pageId="5" pageNumber="225">et al</emphasis>
. (2010)
</bibRefCitation>
shows only styles with the stigma exerted among or slightly beyond the anthers (i.e., longistylous). They mention only one study of one species (
<taxonomicName id="4C084D6F0D2ABF67FE72DF05FC0DFE8E" authority="H. L. Wendl. ex Pfeiffer" authorityName="Pfeiffer" box="[440,909,266,293]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="robusta">
<emphasis id="B97CEAFE0D2ABF67FE72DF05FDF3FE8E" box="[440,627,266,293]" italics="true" pageId="5" pageNumber="225">Opuntia robusta</emphasis>
H.L. Wendl. ex Pfeiffer
</taxonomicName>
) with a brevistylous morph. The flowers of 
<taxonomicName id="4C084D6F0D2ABF67FA5ADF05FE94FEE2" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2ABF67FA5ADF05FE94FEE2" italics="true" pageId="5" pageNumber="225">D. chontalensis</emphasis>
</taxonomicName>
, however, were originally described as unusual for having an “extremely short style” (
<bibRefCitation id="EF994B1D0D2ABF67FB2ADF20FA1FFEE3" author="Alexander, E. J." box="[1248,1439,302,329]" pageId="5" pageNumber="225" pagination="131 - 133" refId="ref3217" refString="Alexander, E. J. (1950) A new Nyctocereus from southern Mexico. Cactus and Succulent Journal (Los Angeles) 22: 131 - 133." type="journal article" year="1950">Alexander, 1950</bibRefCitation>
), and were illustrated (
<bibRefCitation id="EF994B1D0D2ABF67FEB2DF5CFDCEFEC6" author="Bravo-Hollis, H." box="[376,590,338,364]" pageId="5" pageNumber="225" refId="ref3247" refString="Bravo-Hollis, H. (1978) Las cactaceas de Mexico. 2 nd ed., vol. 1. Universidad Nacional Autonoma de Mexico, Mexico City, 743 pp." type="book" year="1978">Bravo-Hollis 1978</bibRefCitation>
) with the stigma as being positioned well below the level of the anthers, in fact, below the insertion of the short stamens (i.e., brevistylous). Furthermore, the protologue includes a curious note attributed to the original collector “Mr. MacDougall notes that the style extends 
<quantity id="4CF09B090D2ABF67FBC0DF95FBCBFE1F" box="[1034,1099,411,437]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="225" unit="mm" value="7.0">7 mm</quantity>
beyond the stamens at anthesis but is back in place below the stamens when the flower closes.” This juxtaposition, so contrary to the usual—where the stigma is exerted slightly beyond or among the anthers—could appear to be some sort of error in the description (the original collection having been lost and the 
<typeStatus id="54B3884E0D2ABF67FDAEDC08FD4EFD8A" box="[612,718,518,544]" pageId="5" pageNumber="225" type="lectotype">lectotype</typeStatus>
, a photo that reveals nothing in this respect). Nonetheless, the short style (
<quantity id="4CF09B090D2ABF67FF07DC24FEB8FDEF" box="[205,312,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="5" pageNumber="225" unit="cm" value="2.1" valueMax="2.2" valueMin="2.0">2–2.2 cm</quantity>
long) with the stigma positioned well below the anthers, has been confirmed with recent collections of 
<taxonomicName id="4C084D6F0D2ABF67FF6CDC41FED4FDC2" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[166,340,590,616]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2ABF67FF6CDC41FED4FDC2" box="[166,340,590,616]" italics="true" pageId="5" pageNumber="225">D. chontalensis</emphasis>
</taxonomicName>
(
<figureCitation id="13332A690D2ABF67FEA9DC40FE27FDC2" box="[355,423,590,616]" captionStart="FIGURE 4" captionStartId="5.[136,229,1791,1813]" captionTargetBox="[158,1427,1012,1763]" captionTargetId="figure-445@5.[151,1436,1001,1766]" captionTargetPageId="5" captionText="FIGURE 4. Deamia chontalensis, long section of flower showing brevistyly, with the stigma positioned below the anthers (Yañez &amp; Arias 004, MEXU). Photo credits: Arias." figureDoi="http://doi.org/10.5281/zenodo.7461389" httpUri="https://zenodo.org/record/7461389/files/figure.png" pageId="5" pageNumber="225">Fig. 4</figureCitation>
). No species of 
<taxonomicName id="4C084D6F0D2ABF67FD93DC41FD32FDC2" authorityName="Britton &amp; Rose" authorityYear="1920" box="[601,690,591,616]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="genus">
<emphasis id="B97CEAFE0D2ABF67FD93DC41FD32FDC2" box="[601,690,591,616]" italics="true" pageId="5" pageNumber="225">Deamia</emphasis>
</taxonomicName>
with more than one flower morph is known. The suggestion that the style stretches out and then shrinks back has not been confirmed. The stamens of 
<taxonomicName id="4C084D6F0D2ABF67FBEDDC7DFB57FD26" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[1063,1239,626,652]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2ABF67FBEDDC7DFB57FD26" box="[1063,1239,626,652]" italics="true" pageId="5" pageNumber="225">D. chontalensis</emphasis>
</taxonomicName>
are also unusually short for a flower of this size, ranging from only 
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long; stamen length in the other species of 
<taxonomicName id="4C084D6F0D2ABF67FA99DC99FA2CFD1A" authorityName="Britton &amp; Rose" authorityYear="1920" box="[1363,1452,663,688]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="genus">
<emphasis id="B97CEAFE0D2ABF67FA99DC99FA2CFD1A" box="[1363,1452,663,688]" italics="true" pageId="5" pageNumber="225">Deamia</emphasis>
</taxonomicName>
varies from 
<quantity id="4CF09B090D2ABF67FEDBDCB4FE08FD7F" box="[273,392,698,725]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="2.0" metricValueMin="0.15" pageId="5" pageNumber="225" unit="cm" value="10.75" valueMax="20.0" valueMin="1.5">1.5–20 cm</quantity>
long. These characters in the floral morphology of 
<taxonomicName id="4C084D6F0D2ABF67FC1BDCB5FB00FD7E" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[977,1152,698,724]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2ABF67FC1BDCB5FB00FD7E" box="[977,1152,698,724]" italics="true" pageId="5" pageNumber="225">D. chontalensis</emphasis>
</taxonomicName>
are strongly suggestive of a particular pollination syndrome worthy of investigation. Thus, although the flowers of 
<taxonomicName id="4C084D6F0D2ABF67FBA6DCD1FA9BFD52" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[1132,1307,734,760]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2ABF67FBA6DCD1FA9BFD52" box="[1132,1307,734,760]" italics="true" pageId="5" pageNumber="225">D. chontalensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C084D6F0D2ABF67FA98DCD1FA2BFD52" authorityName="Hammel &amp; S. Arias" authorityYear="2022" box="[1362,1451,734,760]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="funis">
<emphasis id="B97CEAFE0D2ABF67FA98DCD1FA2BFD52" box="[1362,1451,734,760]" italics="true" pageId="5" pageNumber="225">D. funis</emphasis>
</taxonomicName>
are outwardly similar, the latter is further distinguished by its longer style (
<quantity id="4CF09B090D2ABF67FC0DDD0CFB9AFCB7" box="[967,1050,770,797]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="225" unit="cm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 cm</quantity>
long) and stamens (
<quantity id="4CF09B090D2ABF67FB30DD0CFAE3FCB7" box="[1274,1379,770,797]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="5" pageNumber="225" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
long), with the stigma among or slightly exceeding the anthers.
</paragraph>
<paragraph id="8BB736EC0D2ABF67FF77DD44FCDAFC7B" blockId="5.[136,1452,158,977]" pageId="5" pageNumber="225">
The submature fruits are similar to those of other species in the genus, in having a smooth surface with spiny areoles, the spines (as in the fruits of 
<taxonomicName id="4C084D6F0D2ABF67FDFBDD61FD63FC22" authorityName="Britton &amp; Rose" authorityYear="1920" baseAuthorityName="Karw. ex Zuccarini" baseAuthorityYear="1837" box="[561,739,878,904]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="testudo">
<emphasis id="B97CEAFE0D2ABF67FDFBDD61FD63FC22" box="[561,739,878,904]" italics="true" pageId="5" pageNumber="225">Deamia testudo</emphasis>
</taxonomicName>
) not completely covering the surface; we are confident that all characters of the new species, especially that of the unique epidermal wax of the stems, often with a sort of cob-web pattern of cracking, are congruent with its placement in 
<taxonomicName id="4C084D6F0D2ABF67FD31DDB9FCD4FC7A" authorityName="Britton &amp; Rose" authorityYear="1920" box="[763,852,951,976]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="genus">
<emphasis id="B97CEAFE0D2ABF67FD31DDB9FCD4FC7A" box="[763,852,951,976]" italics="true" pageId="5" pageNumber="225">Deamia</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF7766640D2ABF67FF42D8F1FE57F893" ID-DOI="http://doi.org/10.5281/zenodo.7461389" ID-Zenodo-Dep="7461389" httpUri="https://zenodo.org/record/7461389/files/figure.png" pageId="5" pageNumber="225" startId="5.[136,229,1791,1813]" targetBox="[158,1427,1012,1763]" targetPageId="5">
<paragraph id="8BB736EC0D2ABF67FF42D8F1FE57F893" blockId="5.[136,1451,1791,1849]" pageId="5" pageNumber="225">
<emphasis id="B97CEAFE0D2ABF67FF42D8F1FF77F8BE" bold="true" box="[136,247,1791,1813]" pageId="5" pageNumber="225">FIGURE 4</emphasis>
. 
<taxonomicName id="4C084D6F0D2ABF67FEC8D8F1FE4BF8BE" authorityName="Doweld" authorityYear="2002" baseAuthorityName="Alexander" baseAuthorityYear="1950" box="[258,459,1791,1812]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="species" species="chontalensis">
<emphasis id="B97CEAFE0D2ABF67FEC8D8F1FE4BF8BE" box="[258,459,1791,1812]" italics="true" pageId="5" pageNumber="225">Deamia chontalensis</emphasis>
</taxonomicName>
, long section of flower showing brevistyly, with the stigma positioned below the anthers (
<emphasis id="B97CEAFE0D2ABF67FAE9D8F1FF2CF892" italics="true" pageId="5" pageNumber="225">Yañez &amp; Arias 004</emphasis>
, MEXU). Photo credits: Arias.
</paragraph>
</caption>
<paragraph id="8BB736EC0D2ABF67FF77D966FBD0F82B" blockId="5.[189,1104,1895,1922]" box="[189,1104,1895,1922]" pageId="5" pageNumber="225">
To facilitate identification of the four known species of 
<taxonomicName id="4C084D6F0D2ABF67FCE7D966FC06F82B" authorityName="Britton &amp; Rose" authorityYear="1920" box="[813,902,1896,1921]" class="Magnoliopsida" family="Cactaceae" genus="Deamia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="225" phylum="Tracheophyta" rank="genus">
<emphasis id="B97CEAFE0D2ABF67FCE7D966FC06F82B" box="[813,902,1896,1921]" italics="true" pageId="5" pageNumber="225">Deamia</emphasis>
</taxonomicName>
we include a key.
</paragraph>
</subSubSection>
</treatment>
</document>