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<mods:title id="FEF884F2445BF7D893526DC6D29B4F1E">A new species of the endemic genus Hemicyrthus Reiche (Coleoptera: Scarabaeidae: Dynastinae) from New Caledonia, with a revised key</mods:title>
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<mods:namePart id="6796142F9523C00CAF80980E5C94AC72">Krell, Frank-Thorsten</mods:namePart>
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<treatment id="E90DF050FFF28D30FF7629BF97B60A80" ID-DOI="http://doi.org/10.5281/zenodo.6113455" ID-GBIF-Taxon="119645602" ID-Zenodo-Dep="6113455" LSID="urn:lsid:plazi:treatment:E90DF050FFF28D30FF7629BF97B60A80" httpUri="http://treatment.plazi.org/id/E90DF050FFF28D30FF7629BF97B60A80" lastPageNumber="288" pageId="7" pageNumber="288">
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<paragraph id="611B4146FFF28D30FF7629BF919D0B17" blockId="7.[151,724,1516,1543]" box="[151,724,1516,1543]" pageId="7" pageNumber="288">
<heading id="3A53F62AFFF28D30FF7629BF919D0B17" bold="true" box="[151,724,1516,1543]" fontSize="11" level="1" pageId="7" pageNumber="288" reason="1">
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<taxonomicName id="A6A43AC5FFF28D30FF7629BF927A0B16" box="[151,307,1516,1542]" class="Insecta" family="Dynastidae" genus="Hemicyrthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="53D09D54FFF28D30FF7629BF927A0B16" bold="true" box="[151,307,1516,1542]" italics="true" pageId="7" pageNumber="288">Hemicyrthus</emphasis>
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, a genus of short-range endemics
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<paragraph id="611B4146FFF28D30FF762A6097900BEF" blockId="7.[151,1437,1587,1936]" pageId="7" pageNumber="288">
All seven
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<emphasis id="53D09D54FFF28D30FEEF2A6092D40B5C" box="[270,413,1587,1612]" italics="true" pageId="7" pageNumber="288">Hemicyrthus</emphasis>
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species are known from few specimens, sometimes only from the
<typeStatus id="BE1FFFE4FFF28D30FB472A6096410B5C" box="[1190,1288,1587,1612]" pageId="7" pageNumber="288" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="05353CB7FFF28D30FAF82A60939A0B60" author="Dechambre" pageId="7" pageNumber="288" refString="Dechambre, R. - P. (1982) Contribution a l'etude phylogenetique des Hexodontini (Coleoptera, Dynastidae). Etude systematique preliminaire du genre Hemicyrthus Reiche. Revue Francaise d'Entomologie, N. S., 4, 101 - 107." type="journal article" year="1982">Dechambre 1982</bibRefCitation>
;
<bibRefCitation id="05353CB7FFF28D30FF002A0492330B60" author="Paulian" box="[225,378,1623,1648]" pageId="7" pageNumber="288" refString="Paulian, R. (1991) Les Coleopteres Scarabaeoidea de Nouvelle-Caledonie (Faune Tropicale 29). Orstom, Paris, 166 pp." type="book" year="1991">Paulian 1991</bibRefCitation>
;
<bibRefCitation id="05353CB7FFF28D30FE682A04915A0B60" author="Thery" box="[393,531,1623,1648]" pageId="7" pageNumber="288" refString="Thery, T. (2008) Nouvelles captures concernant le genre Hemicyrthus Reiche, 1860 (Col., Dynastidae, Hexodontini). Bulletin de la Societe Entomologique de France, 113, 341 - 342." type="journal article" year="2008">Théry 2008</bibRefCitation>
). They all are known from either single locations or small areas (
<figureCitation id="F99F5DC3FFF28D30FAF12A0496110B60" box="[1296,1368,1623,1648]" captionStart="FIGURE 5" captionStartId="6.[151,250,1757,1779]" captionTargetBox="[207,1379,876,1735]" captionTargetId="figure@6.[207,1379,876,1736]" captionTargetPageId="6" captionText="FIGURE 5. Published collecting localities of Hemicyrthus species (Dechambre 1982; Paulian 1991; Théry 2008; present study). 1. Hemicyrthus serresii (Montrouzier, 1860), Iles Belep; 2. Hemicyrthus villersi (Montrouzier, 1860), Mont Mandjélia; 3. Hemicyrthus villersi (Montrouzier, 1860), Mont Panié; 4. Hemicyrthus costatus Dechambre, 1982, Mont Boulinda; 5. Hemicyrthus chazeaui Dechambre, 1982 and Hemicyrthus gutierrezi Dechambre, 1982, Mont Do; 6. Hemicyrthus elongatus Dechambre, 1982, Upper Ngoe River; 7. Hemicyrthus blaffarti new species, Parc Provincial de la Rivière Bleue. Dark grey area (elevation 700 1628 m), light grey area (elevation 300 700 m), white area (elevation under 300 m). Hemicyrthus reichei Dechambre, 1982, is not indicated in this map because it was described from a historical specimen labelled “ N. Caled. ” without further detail." httpUri="https://zenodo.org/record/232080/files/figure.png" pageId="7" pageNumber="288">Fig. 5</figureCitation>
). The species with the widest known range is
<taxonomicName id="A6A43AC5FFF28D30FDB22A2E91830B84" box="[595,714,1659,1684]" class="Insecta" family="Dynastidae" genus="Hemicyrthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="villiersi">
<emphasis id="53D09D54FFF28D30FDB22A2E91830B84" box="[595,714,1659,1684]" italics="true" pageId="7" pageNumber="288">H. villiersi</emphasis>
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from Mont Mandjelia to Mont Panié, which are just over
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apart and are connected by a continuous mountain forest. The next largest range is that of
<taxonomicName id="A6A43AC5FFF28D30FB672AF2964D0BA8" box="[1158,1284,1695,1720]" class="Insecta" family="Dynastidae" genus="Hemicyrthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="blaffarti">
<emphasis id="53D09D54FFF28D30FB672AF2964D0BA8" box="[1158,1284,1695,1720]" italics="true" pageId="7" pageNumber="288">H. blaffarti</emphasis>
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, with Refuge de la Rivière Bleue in the east and Grand Kaori in the west of its known range being just over
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apart. The other species are known from one locality only, with no available spatial data about the collecting area.
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<paragraph id="611B4146FFF28D30FF262B5897B60A80" blockId="7.[151,1437,1587,1936]" pageId="7" pageNumber="288">
Short-range endemism is common in
<collectingCountry id="19B301D6FFF28D30FD9C2B5F90670A34" box="[637,814,1803,1828]" name="New Caledonia" pageId="7" pageNumber="288">New Caledonia</collectingCountry>
with numerous examples in,
<emphasis id="53D09D54FFF28D30FB692B5E97F90A34" box="[1160,1200,1805,1828]" italics="true" pageId="7" pageNumber="288">e.g.</emphasis>
, reptiles, plants and insects (Murienne
<emphasis id="53D09D54FFF28D30FE872B6292D70A58" box="[358,414,1839,1864]" italics="true" pageId="7" pageNumber="288">et al.</emphasis>
2008), and is promoted by low dispersal capability and low ecological tolerance (
<bibRefCitation id="05353CB7FFF28D30FACF2B7C927B0A7C" author="Ponder" pageId="7" pageNumber="288" refString="Ponder, W. F. &amp; Colgan, D. J. (2002) What makes a narrow-range taxon? Insights from Australian freshwater snails. Invertebrate Systematics, 16, 571 - 582. http: // dx. doi. org / 10.1071 / IT 01043" type="journal article" year="2002">Ponder &amp; Colgan 2002</bibRefCitation>
). The ecological requirements of
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<emphasis id="53D09D54FFF28D30FD292B00901E0A7C" box="[712,855,1875,1900]" italics="true" pageId="7" pageNumber="288">Hemicyrthus</emphasis>
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species are unknown, but the flightlessness and substantial size of the
<taxonomicName id="A6A43AC5FFF28D30FE712B2491560A80" box="[400,543,1911,1936]" class="Insecta" family="Dynastidae" genus="Hemicyrthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="53D09D54FFF28D30FE712B2491560A80" box="[400,543,1911,1936]" italics="true" pageId="7" pageNumber="288">Hemicyrthus</emphasis>
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species greatly restrict their active and passive dispersal abilities.
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