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<document id="159BC414D3C1012797A859BB51AFE301" ID-DOI="http://dx.doi.org/10.3897/zookeys.665.11792" ID-GBIF-Dataset="f49697cc-fc0c-4a14-bba1-edf1268f59f0" ID-PMC="PMC5523169" ID-Pensoft-Pub="1313-2970-665-93" ID-PubMed="28769629" ID-ZooBank="4D5230AC67CB47ADAF2F051CA10D9724" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-665-93" ModsDocOrigin="ZooKeys 665" ModsDocTitle="Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1491378304537" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Wild, Michael, Sumoked, Bob &amp; Balke, Michael" docDate="2017" docId="42402B3BFFAB28CE06C558F66FC53294" docLanguage="en" docName="ZooKeys 665: 93-120" docOrigin="ZooKeys 665" docSource="http://dx.doi.org/10.3897/zookeys.665.11792" docTitle="Exocelina ascendens Balke 1998" docType="treatment" docVersion="5" lastPageNumber="93" masterDocId="DC520217FF93FFF8FFDEFFCFFFA3777E" masterDocTitle="Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="120" masterPageNumber="93" pageNumber="93" updateTime="1701318038850" updateUser="plazi">
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<mods:title id="2BE80896C28EAA48C042DCE98843C615">Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="A94FA7DED997EC67F52B8CF231CDA75A">Shaverdo, Helena</mods:namePart>
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<mods:namePart id="3F3EB3C4A08AFA5B1B699CABA315F833">Wild, Michael</mods:namePart>
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<mods:namePart id="E8161998864F20AEB6DD57C166E151EC">Sumoked, Bob</mods:namePart>
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<mods:namePart id="CBDBEDEAA20BBF7B28152122CD15F334">Balke, Michael</mods:namePart>
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<treatment id="42402B3BFFAB28CE06C558F66FC53294" ID-GBIF-Taxon="128229134" LSID="urn:lsid:plazi:treatment:42402B3BFFAB28CE06C558F66FC53294" httpUri="http://treatment.plazi.org/id/42402B3BFFAB28CE06C558F66FC53294" lastPageNumber="93" pageId="0" pageNumber="93">
<subSubSection id="DD8FB4BF25F6A8EA4A021A2A295EF2F6" pageId="0" pageNumber="93" type="nomenclature">
<paragraph id="F1C453E8B27F88C38DA4B726D83B928D" pageId="0" pageNumber="93">
1.
<taxonomicName id="CDEF4474326C762E0C34B033D84A0557" ID-CoL="3DLNW" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ascendens" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="ascendens">Exocelina ascendens (Balke, 1998)</taxonomicName>
Figs 3, 11
</paragraph>
</subSubSection>
<subSubSection id="132F1964D38D81DB79E4959D156F094E" pageId="0" pageNumber="93" type="reference_group">
<paragraph id="A5F189DBEB2484EAE9E64CB1AA4BAF93" pageId="0" pageNumber="93">
<taxonomicName id="7723C6888E0740717E266E2378D55181" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) ascendens" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="ascendens" subGenus="Papuadytes">Copelatus (Papuadytes) ascendens</taxonomicName>
Balke, 1998: 322;
<bibRefCitation id="40EEFDF15AF73B3DBBF3E71DB3C4CFB2" author="Nilsson, AN" journalOrPublisher="Stenstrup, Apollo Books" pageId="12" pageNumber="105" title="Dytiscidae. World catalogue of insects. Vol. 3." year="2001">Nilsson 2001</bibRefCitation>
: 76 (catalogue).
</paragraph>
<paragraph id="0A806E7C8352E56A7CBA3B1BCC28CC59" pageId="0" pageNumber="93">
<taxonomicName id="CFC5EA2784F77ADC64F53519469A922B" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes ascendens" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="ascendens">Papuadytes ascendens</taxonomicName>
(Balke, 1998):
<bibRefCitation id="914A71D8FF43BBF6D7D118928FFE0E44" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="105" pagination="55 - 74" title="World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
: 56 (comb. n.).
</paragraph>
<paragraph id="64CB3928BC225809D7FBC3DB92CE83BE" pageId="0" pageNumber="93">
<taxonomicName id="F042CC15F47E96EA681D79492A7B1A83" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ascendens" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="ascendens">Exocelina ascendens</taxonomicName>
(Balke, 1998):
<bibRefCitation id="3379F5858CC5099EA647356C9CC967EA" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="12" pageNumber="105" pagination="33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
: 33 (comb. n.).
</paragraph>
</subSubSection>
<subSubSection id="B84261E1F480E0406BDF25BF02B9AA4E" pageId="0" pageNumber="93" type="type locality">
<paragraph id="A0B53263A494F450D74B608100F52113" pageId="0" pageNumber="93">Type locality.</paragraph>
<paragraph id="F5329FEAC97FE69846ECAB2225C76E4E" pageId="0" pageNumber="93">
Papua: Pegunungan Bintang Regency, trek between Aipomek and Diruemna,
<geoCoordinate id="1467B3124F0059419A2A9B18F426DAFB" direction="south" orientation="latitude" precision="925" value="-4.4166665">04°25'S</geoCoordinate>
;
<geoCoordinate id="C12ACF36059DC278B2D24DC07520339F" direction="east" orientation="longitude" precision="925" value="139.95">139°57'E</geoCoordinate>
, 2600 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C818A1BFE77C4C95357770D5E9E22E27" pageId="0" pageNumber="93" type="type material">
<paragraph id="8BEC3B6A2E5D8E214DF27CFCB62BE066" pageId="0" pageNumber="93">Type material studied.</paragraph>
<paragraph id="3E5D53042BE3208527829D02D776612E" pageId="0" pageNumber="93">
Holotype: male &quot;IRIAN JAYA, 3.9.1992 Aipomek - Diuremna [sic!]
<geoCoordinate id="8B4A9555D22A746CB457094AB0D47EC2" direction="east" orientation="longitude" precision="925" value="139.95">139°57'E</geoCoordinate>
<geoCoordinate id="CD57D6876D5C389096807A3FD18AE7E6" direction="south" orientation="latitude" precision="925" value="-4.4166665">04°25'S</geoCoordinate>
2600m, leg.Balke (35)&quot;,
<normalizedToken id="2EFC91B67A51BCC3327B4C6501DD548C" originalValue="“HOLOTYPUS”">&quot;HOLOTYPUS&quot;</normalizedToken>
[red], &quot;
<taxonomicName id="F1087FBA7E82FA12563571D13B228D36" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
<taxonomicName id="414E601C053A223DC0B8B3865DACACD7" lsidName="ascendens" pageId="0" pageNumber="93" rank="species" species="ascendens">ascendens</taxonomicName>
Balke des. 1997&quot; [red] (NHMW). Paratypes: 4 males, 2 females with the same label as the holotype and additionally with red labels &quot;Paratypus
<taxonomicName id="4623BD8318B0E46679FBC7A39CF6DEAE" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
<taxonomicName id="EAD030497A00884B478F0D0CE5EBDD9E" lsidName="ascendens" pageId="0" pageNumber="93" rank="species" species="ascendens">ascendens</taxonomicName>
Balke des. 1997&quot;, one of the males with an additional label &quot;M.Balke 3282&quot; [green], another male with two additional labels &quot;M.Balke 3283&quot; [green] and &quot;M.Balke 6409&quot; [green text] (NHMW). 1 male, 2 females &quot;IRIAN JAYA, 9.9.1992 Kono - Angguruk
<geoCoordinate id="45A71622D566E6A1632BCFF5B0E38FC7" direction="east" orientation="longitude" precision="925" value="139.78334">139°47'E</geoCoordinate>
<geoCoordinate id="CDCD949CB88EDECACC8DB9100B4C3F30" direction="south" orientation="latitude" precision="925" value="-4.3166666">04°19'S</geoCoordinate>
2600m, leg.Balke (44)&quot;, &quot;Paratypus
<taxonomicName id="635C0438FC1BFAF3533197487E46F62C" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
<taxonomicName id="EADF63EE0351E075E047BE5F63BEFEE4" lsidName="ascendens" pageId="0" pageNumber="93" rank="species" species="ascendens">ascendens</taxonomicName>
Balke des. 1997&quot; [red] (NHMW), note: the original description says &quot;2 males and 1 female&quot;, it is probably a mistake in the sex identification or type error. 4 males, 3 females &quot;IRIAN JAYA, 24.-26.9.1993 Eipomek [sic!] Gebiet Eipomek [sic!] - Diruemna&quot;, &quot;ca.
<geoCoordinate id="25DC1DD4EBC399700BB4ABD3BBFABFF7" direction="east" orientation="longitude" precision="925" value="140.01666">140°01'E</geoCoordinate>
<geoCoordinate id="4C683BC0F63DFDA1A0453B662A67AD21" direction="south" orientation="latitude" precision="925" value="-4.45">04°27'S</geoCoordinate>
1800-2600m, leg. M. Balke (21 [crossed out] -22)&quot;, &quot;Paratypus
<taxonomicName id="E8C1C2907B20295076178D05FC49163A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
<taxonomicName id="F70A1444A73E756AF3EF97B7FC1737B5" lsidName="ascendens" pageId="0" pageNumber="93" rank="species" species="ascendens">ascendens</taxonomicName>
Balke des. 1997&quot; [red] (NHMW, CGW).
</paragraph>
</subSubSection>
<subSubSection id="618EDFE5F68DE3DC6A7BD765B9F195DD" pageId="0" pageNumber="93" type="diagnosis">
<paragraph id="CE3727CC664F5F1C6C394312BCC0324F" pageId="0" pageNumber="93">Diagnosis.</paragraph>
<paragraph id="4DCE5F8538DF6F33749CD44D36894CBF" pageId="0" pageNumber="93">
Beetle large (TL-H 5.3-5.75 mm), elongate; piceous, with dark brown pronotal sites and head anteriorly; submatt, with fine but evident punctation and rather strongly impressed microreticulation; pronotum with distinct lateral bead; male antennae simple, slender (Fig. 3); male protarsomere 4 with small (slightly larger than more laterally situated large seta), weakly curved anterolateral hook-like seta; male protarsomere 5 ventrally with anterior row of 23 and posterior row of 8 short, strong, spine-like setae (Fig. 11D); median lobe evenly curved, pointed in lateral view and evenly tapering, with broadly pointed apex in ventral view, on both lateral sides with numerous fine setae situated linearly on anterior half of distal part of median lobe; paramere robust, with notch on dorsal side and very dense, strong setae on subdistal part; proximal setae sparse and fine (Fig. 11
<normalizedToken id="9202823B89DE9895E545257159205056" originalValue="AC">A-C</normalizedToken>
). For complete description, see
<bibRefCitation id="CBF3E84EBC641228BFFE89C24D71592B" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="11" pageNumber="104" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I" volume="100 B" year="1998">Balke (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E2D0F46780FA266BA66D521752F1ABC" pageId="0" pageNumber="93" type="distribution">
<paragraph id="34CA43F60E958F0CA42D90412F7131BF" pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph id="F9854C3FF6B003DEE8E9B823DEBC69CF" pageId="0" pageNumber="93">Papua: Pegunungan Bintang Regency. The species is known only from the type material (Fig. 19).</paragraph>
</subSubSection>
</treatment>
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