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<document id="8B4866C97C5A722B5340AF392897C99D" ID-CLB-Dataset="29410" ID-DOI="10.11646/zootaxa.4462.1.5" ID-GBIF-Dataset="bea9fe5a-e3aa-47c3-a268-766e1053e9b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441438" ID-ZooBank="FA506479-22AA-4875-9482-3391D2584A4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538402359286" checkinUser="plazi" docAuthor="Bota-Sierra, Cornelio Andrés, Sánchez-Herrera, Melissa &amp; Palacino-Rodríguez, Fredy" docDate="2018" docId="42270F6D5672FFB4FF27FE4D5A30FBF4" docLanguage="en" docName="zootaxa.4462.1.5.pdf" docOrigin="Zootaxa 4462 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cora verapax Bota-Sierra &amp; Sánchez-Herrera &amp; Palacino-Rodríguez 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="124" masterDocId="BE1E77155675FFBDFFB0FFB25807FFAC" masterDocTitle="Odonata from protected areas in Colombia with new records and description of Cora verapax sp. nov. (Zygoptera: Polythoridae)" masterLastPageNumber="131" masterPageNumber="115" pageNumber="122" updateTime="1698560320511" updateUser="ExternalLinkService">
<mods:mods id="12FCF0CD8C618AE813E4629C1DF3F9F2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="173C309FC68A4D0C446981375FC8E96F">
<mods:title id="66C6BDF80AD1CC77DB11CE2CE20BB794">Odonata from protected areas in Colombia with new records and description of Cora verapax sp. nov. (Zygoptera: Polythoridae)</mods:title>
</mods:titleInfo>
<mods:name id="57E2D49DB1BC36478AA8D1FC76D6AE6E" type="personal">
<mods:role id="E1584FB9A0DC2F78E31968A060DF9D44">
<mods:roleTerm id="7C24AF180875530458C383019290AA6D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D7CF4A8A9926F773CE1CDF3B395317A">Bota-Sierra, Cornelio Andrés</mods:namePart>
</mods:name>
<mods:name id="8A87606FCD005BE1051C451AE302D2E0" type="personal">
<mods:role id="0A6EF353E8D68861377D7E1B156DAA5C">
<mods:roleTerm id="26B98B522F9A02B3EF5CC6F7BF8839B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D215E0FE9348374370708783729F5B2">Sánchez-Herrera, Melissa</mods:namePart>
</mods:name>
<mods:name id="A13490DD83273D15613ABC06D5C28B1E" type="personal">
<mods:role id="EF89AF9FBACD91A524A12A5F70C2E29D">
<mods:roleTerm id="7F5BCE20218F88F17AFE96EF2D52A167">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E2C931D08052DC982CB0F1D30E736611">Palacino-Rodríguez, Fredy</mods:namePart>
</mods:name>
<mods:typeOfResource id="810A7BFB4C9DB41B80B328C0DD6F4852">text</mods:typeOfResource>
<mods:relatedItem id="19616EE87719B710C04D355A36183316" type="host">
<mods:titleInfo id="689992705885EFE41D1295B391528888">
<mods:title id="9E4C2F07673F15EE0EE4F1FB21DF1FC4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0A525047ACEA39B56F1C509F386E1462">
<mods:date id="4E0BC262A8E47AA96D0B605CAD531013">2018</mods:date>
<mods:detail id="685F4827698861B2F76D0E077F85AEF2" type="pubDate">
<mods:number id="26B2996A2B7AB8565D90D20971B35F5B">2018-08-24</mods:number>
</mods:detail>
<mods:detail id="E0261407BB6C385CBA3DDFD3EF8CA64B" type="volume">
<mods:number id="E3DC6BCE2C253C2731B0A7D7EEA001AD">4462</mods:number>
</mods:detail>
<mods:detail id="AE1F10082FB9AC372B1BEAA9CF60BA18" type="issue">
<mods:number id="CB9C6F6C8830C27A7591F2767C25D5DE">1</mods:number>
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<mods:start id="D15A601BD89AA1619DE48E1E0308E15F">115</mods:start>
<mods:end id="75FCB0951C8485AF0425D75275A1D2DA">131</mods:end>
</mods:extent>
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<mods:classification id="73AC5960C3B011BAABC0CA62B87B6EDD">journal article</mods:classification>
<mods:identifier id="C4EA254D1B16FD87356FBB491A7A49D6" type="CLB-Dataset">29410</mods:identifier>
<mods:identifier id="FA1FC30631C21FC0BBB1EF2C5A124897" type="DOI">10.11646/zootaxa.4462.1.5</mods:identifier>
<mods:identifier id="48F1A3EE2CDD1270D25F0ABC53359454" type="GBIF-Dataset">bea9fe5a-e3aa-47c3-a268-766e1053e9b3</mods:identifier>
<mods:identifier id="F7F2D19751E72C24B4F82FD5A7208786" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="831E4ECED2B44C621C9A4FBB6716CFAA" type="Zenodo-Dep">1441438</mods:identifier>
<mods:identifier id="38CB5BBE9BD726B08E06DD661457B045" type="ZooBank">FA506479-22AA-4875-9482-3391D2584A4C</mods:identifier>
</mods:mods>
<treatment id="42270F6D5672FFB4FF27FE4D5A30FBF4" ID-DOI="http://doi.org/10.5281/zenodo.5986827" ID-GBIF-Taxon="148424419" ID-Zenodo-Dep="5986827" LSID="urn:lsid:plazi:treatment:42270F6D5672FFB4FF27FE4D5A30FBF4" httpUri="http://treatment.plazi.org/id/42270F6D5672FFB4FF27FE4D5A30FBF4" lastPageId="9" lastPageNumber="124" pageId="7" pageNumber="122">
<subSubSection id="8294EDF05672FFBAFF27FE4D5915FD97" pageId="7" pageNumber="122" type="nomenclature">
<paragraph id="CA31BE7B5672FFBAFF27FE4D59A7FDB6" blockId="7.[151,416,511,571]" box="[151,416,511,538]" pageId="7" pageNumber="122">
<heading id="917909175672FFBAFF27FE4D59A7FDB6" bold="true" box="[151,416,511,538]" fontSize="11" level="1" pageId="7" pageNumber="122" reason="1">
<emphasis id="F8FA62695672FFBAFF27FE4D59A7FDB6" bold="true" box="[151,416,511,538]" pageId="7" pageNumber="122">
<taxonomicName id="0D8EC5F85672FFBAFF27FE4D593DFDB5" ID-CoL="9WTQL" authority="Bota-Sierra &amp; Sánchez-Herrera &amp; Palacino-Rodríguez, 2018" authorityName="Bota-Sierra &amp; Sánchez-Herrera &amp; Palacino-Rodríguez" authorityYear="2018" box="[151,314,511,537]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="verapax" status="sp. nov.">
<emphasis id="F8FA62695672FFBAFF27FE4D593DFDB5" bold="true" box="[151,314,511,537]" italics="true" pageId="7" pageNumber="122">Cora verapax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3C9DF125672FFBAFEF2FDB259A7FDB6" box="[322,416,512,538]" pageId="7" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CA31BE7B5672FFBAFF27FD905915FD97" blockId="7.[151,416,511,571]" box="[151,274,546,571]" pageId="7" pageNumber="122">
(
<figureCitation id="52B5A2FE5672FFBAFF2FFD90590EFD97" box="[159,265,546,571]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a–e</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8294EDF05672FFBAFF27FDD85ADBFD67" pageId="7" pageNumber="122" type="materials_examined">
<paragraph id="CA31BE7B5672FFBAFF27FDD85ADBFD67" blockId="7.[151,1437,618,2011]" pageId="7" pageNumber="122">
<materialsCitation id="7AE6B4265672FFBAFF27FDD85ADBFD67" ID-GBIF-Occurrence="1913891585" collectingDate="2016-07-13" collectionCode="CEUA" collectorName="de Cuarzo &amp; Bota &amp; Ayala &amp; GarCia &amp; A. Orejuela" country="Colombia" county="National Natural Park Tatama" elevation="1480" latitude="5.22861" location="Monte Bello Township" longLatPrecision="1" longitude="-76.09806" municipality="Pueblo RiCo MuniCipality" pageId="7" pageNumber="122" specimenCode="CEUA 99826" specimenCount="1" specimenCount-male="1" stateProvince="Risaralda Department" typeStatus="holotype">
<emphasis id="F8FA62695672FFBAFF27FDD8590CFD2F" bold="true" box="[151,267,618,643]" pageId="7" pageNumber="122">
<typeStatus id="153500D95672FFBAFF27FDD85901FD2F" box="[151,262,618,643]" pageId="7" pageNumber="122" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="DC8875F25672FFBAFEA8FDD9592AFD2E" box="[280,301,619,642]" pageId="7" pageNumber="122" type="male">♂</specimenCount>
<collectingCountry id="B299FEEB5672FFBAFE8AFDD959D4FD2F" box="[314,467,619,643]" name="Colombia" pageId="7" pageNumber="122">COLOMBIA</collectingCountry>
.
<collectingRegion id="084A70995672FFBAFE53FDD85AE4FD2E" box="[483,739,618,643]" country="Colombia" name="Risaralda" pageId="7" pageNumber="122">Risaralda Department</collectingRegion>
:
<collectingCounty id="2350C6F75672FFBAFD43FDD85C5FFD2F" box="[755,1112,618,643]" pageId="7" pageNumber="122">National Natural Park Tatamá</collectingCounty>
,
<collectingMunicipality id="2A5524015672FFBAFBD9FDD85D9FFD2F" box="[1129,1432,618,643]" pageId="7" pageNumber="122">Pueblo RiCo MuniCipality</collectingMunicipality>
,
<location id="CF51E8A05672FFBAFF27FD3D599CFD0B" LSID="urn:lsid:plazi:treatment:42270F6D5672FFB4FF27FE4D5A30FBF4:CF51E8A05672FFBAFF27FD3D599CFD0B" box="[151,411,654,679]" country="Colombia" county="National Natural Park Tatama" latitude="5.22861" longLatPrecision="1" longitude="-76.09806" municipality="Pueblo RiCo MuniCipality" name="Monte Bello Township" pageId="7" pageNumber="122" stateProvince="Risaralda Department">Monte Bello Township</location>
,
<location id="CF51E8A05672FFBAFE16FD3C5A62FD0B" LSID="urn:lsid:plazi:treatment:42270F6D5672FFB4FF27FE4D5A30FBF4:CF51E8A05672FFBAFE16FD3C5A62FD0B" box="[422,613,654,679]" country="Colombia" county="National Natural Park Tatama" latitude="5.22861" longLatPrecision="1" longitude="-76.09806" municipality="Pueblo RiCo MuniCipality" name="Minas de Cuarzo" pageId="7" pageNumber="122" stateProvince="Risaralda Department">
Minas
<collectorName id="677BDBAD5672FFBAFE43FD3C5A62FD0B" box="[499,613,654,679]" pageId="7" pageNumber="122">de Cuarzo</collectorName>
</location>
stream,
<geoCoordinate id="AFBAD8BC5672FFBAFD74FD3C5B38FD0B" box="[708,831,654,679]" direction="north" orientation="latitude" pageId="7" pageNumber="122" precision="1" value="5.22861">5.22861°N</geoCoordinate>
<geoCoordinate id="AFBAD8BC5672FFBAFCF6FD3C5BD3FD0B" box="[838,980,654,679]" direction="west" orientation="longitude" pageId="7" pageNumber="122" precision="1" value="-76.09806">76.09806°W</geoCoordinate>
<elevation id="41A359485672FFBAFC6BFD3D5C74FD0B" box="[987,1139,654,679]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="7" pageNumber="122" unit="m" value="1480.0">
<quantity id="0D76139E5672FFBAFC6BFD3D5C30FD0A" box="[987,1079,655,679]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="7" pageNumber="122" unit="m" value="1480.0">1,480 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="AE7461535672FFBAFB30FD3D5D14FD0B" box="[1152,1299,654,679]" pageId="7" pageNumber="122" value="2016-07-13">
<date id="BE3098BB5672FFBAFB30FD3D5D14FD0B" box="[1152,1299,654,679]" pageId="7" pageNumber="122" value="2016-07-13">13 July 2016</date>
</collectingDate>
, C.
<collectorName id="677BDBAD5672FFBAFA8EFD3D5D71FD0A" box="[1342,1398,655,678]" pageId="7" pageNumber="122">Bota</collectorName>
, D.
<collectorName id="677BDBAD5672FFBAFF27FD0058DDFD67" box="[151,218,690,715]" pageId="7" pageNumber="122">Ayala</collectorName>
, F.
<collectorName id="677BDBAD5672FFBAFEB0FD01594CFD67" box="[256,331,691,715]" pageId="7" pageNumber="122">GarCía</collectorName>
&amp;
<collectorName id="677BDBAD5672FFBAFEDFFD0159F6FD67" box="[367,497,690,715]" pageId="7" pageNumber="122">A. Orejuela</collectorName>
leg. (
<specimenCode id="9A2816005672FFBAFD80FD015AD7FD67" box="[560,720,690,715]" collectionCode="CEUA" pageId="7" pageNumber="122">CEUA 99826</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8294EDF05672FFBAFF77FD645AE0FCD3" pageId="7" pageNumber="122" type="etymology">
<paragraph id="CA31BE7B5672FFBAFF77FD645AE0FCD3" blockId="7.[151,1437,618,2011]" pageId="7" pageNumber="122">
<emphasis id="F8FA62695672FFBAFF77FD645949FD43" bold="true" box="[199,334,726,751]" pageId="7" pageNumber="122">Etymology.</emphasis>
This name (f., noun in apposition) Comes from the Latin words,
<emphasis id="F8FA62695672FFBAFBE0FD655C94FD42" box="[1104,1171,727,750]" italics="true" pageId="7" pageNumber="122">verus,</emphasis>
meaning true and
<emphasis id="F8FA62695672FFBAFAC4FD655D9BFD42" box="[1396,1436,727,750]" italics="true" pageId="7" pageNumber="122">pax</emphasis>
meaning peaCe. It reCalls the Claim of the Colombian people for the end of the longest Civil war, whiCh has devastated the Country for deCades (i.e. 63 years). Moreover, it expresses the ConCern of the soCiety with the preservation of natural resourCes that were unreaChable and threatened due to this war in our territories; and Calls for our biggest hope: to aChieve a true state of peaCe.
</paragraph>
</subSubSection>
<subSubSection id="8294EDF05672FFBAFF77FC385CC2FA4F" pageId="7" pageNumber="122" type="description">
<paragraph id="CA31BE7B5672FFBAFF77FC385D28FC6B" blockId="7.[151,1437,618,2011]" pageId="7" pageNumber="122">
<emphasis id="F8FA62695672FFBAFF77FC3859E9FC0E" bold="true" box="[199,494,906,931]" pageId="7" pageNumber="122">
Description of
<typeStatus id="153500D95672FFBAFE31FC3859EFFC0F" box="[385,488,906,931]" pageId="7" pageNumber="122" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="F8FA62695672FFBAFE4BFC385A30FC0F" box="[507,567,906,931]" italics="true" pageId="7" pageNumber="122">Head</emphasis>
. Yellow, with greyish-yellow vertex, and two laterodorsal blaCk spots on labrum, posterior half of frons blaCk with two yellow spots behind lateral oCelli (
<figureCitation id="52B5A2FE5672FFBAFBAFFC1C5C76FC6B" box="[1055,1137,942,967]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a</figureCitation>
). Frons rounded.
</paragraph>
<paragraph id="CA31BE7B5672FFBAFF77FC605AF5FB4B" blockId="7.[151,1437,618,2011]" pageId="7" pageNumber="122">
<emphasis id="F8FA62695672FFBAFF77FC60591FFC47" box="[199,280,978,1003]" italics="true" pageId="7" pageNumber="122">Thorax</emphasis>
. Prothorax: pronotum blue with middorsal blaCk stripe, posterior portion of median lobe blaCk, and posterior lobe blaCk margined blue (
<figureCitation id="52B5A2FE5672FFBAFD9DFC445A9EFBA3" box="[557,665,1014,1039]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a, e</figureCitation>
); propleuron yellow (
<figureCitation id="52B5A2FE5672FFBAFC3CFC445BFFFBA3" box="[908,1016,1014,1039]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a, e</figureCitation>
). Pterothorax: blue with blaCk stripes in middorsal Carina, mesepisternum and antehumeral suture, the latter expanding at its proximal portion surrounding the mesoCoxae (
<figureCitation id="52B5A2FE5672FFBAFE45FB8C5A74FBFB" box="[501,627,1086,1111]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a, e</figureCitation>
); thin blaCk lines between mesepimeron and metepimeron and in metapleural suture, this one gets wider towards the posterior margin and toward its proximal portion surrounding the metaCoxae (
<figureCitation id="52B5A2FE5672FFBAFEFBFB3459BDFB33" box="[331,442,1158,1183]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a, e</figureCitation>
); blaCk spot surrounding the metepimeral spiraCle. Venter gray with a tuberCle Close to abdomen base, posteriorly Covered with golden setae. Legs: Coxae grayish blue (Covered with pruinesCenCe when alive,
<figureCitation id="52B5A2FE5672FFBAFF6BFB7C5942FB4B" box="[219,325,1230,1255]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a, e</figureCitation>
), armature and legs blaCk (
<figureCitation id="52B5A2FE5672FFBAFDC4FB7C5AE2FB4B" box="[628,741,1230,1255]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Figs 4a, e</figureCitation>
).
</paragraph>
<paragraph id="CA31BE7B5672FFBAFF77FB465C90FA83" blockId="7.[151,1437,618,2011]" pageId="7" pageNumber="122">
<emphasis id="F8FA62695672FFBAFF77FB46590DFAA7" box="[199,266,1268,1291]" italics="true" pageId="7" pageNumber="122">Wings</emphasis>
. Membrane smoky Colored; venation and pterostigma blaCk; Px
<quantity id="0D76139E5672FFBAFC5EFB415C2FFAA7" box="[1006,1064,1266,1291]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="7" pageNumber="122" unit="in" value="39.0">39 in</quantity>
FW,
<quantity id="0D76139E5672FFBAFBD8FB415CCBFAA7" box="[1128,1228,1266,1291]" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="7.874" metricValueMin="7.62" pageId="7" pageNumber="122" unit="in" value="30.5" valueMax="31.0" valueMin="30.0">30–31 in</quantity>
HW; pt overlying six Cells; FW nodus Closer to the pterositgma (13.0mm) than to the base of wing (
<quantity id="0D76139E5672FFBAFB98FAA45C8CFA83" box="[1064,1163,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="7" pageNumber="122" unit="mm" value="14.5">14.5mm</quantity>
).
</paragraph>
<paragraph id="CA31BE7B5672FFBAFF77FA885CC2FA4F" blockId="7.[151,1437,618,2011]" pageId="7" pageNumber="122">
<emphasis id="F8FA62695672FFBAFF77FA885936FAFF" box="[199,305,1338,1363]" italics="true" pageId="7" pageNumber="122">Abdomen</emphasis>
. BlaCk exCept dorsum of S1 and anterior 2/3 of S2 dorsum whiCh are blue (
<figureCitation id="52B5A2FE5672FFBAFB14FA885D13FAFF" box="[1188,1300,1338,1363]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4a, e</figureCitation>
). S10 has a middorsal horn on apex, pointed in dorsoposterior direCtion (
<figureCitation id="52B5A2FE5672FFBAFCEAFAEC5BD6FADB" box="[858,977,1374,1399]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig 4 C–d</figureCitation>
). Genital ligula with filamentous lateral lobes, small midlateral proCesses, and wide lateroapiCal projeCtions (
<figureCitation id="52B5A2FE5672FFBAFC2BFA305BE8FA37" box="[923,1007,1410,1435]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4b</figureCitation>
). CerCus tip Curving inwards in dorsal view (
<figureCitation id="52B5A2FE5672FFBAFF53FA14593CFA13" box="[227,315,1446,1471]" captionStart="FIGURE 1" captionStartId="1.[151,250,1326,1348]" captionTargetBox="[160,1430,520,1258]" captionTargetId="figure@1.[151,1436,511,1270]" captionTargetPageId="1" captionText="FIGURE 1. Colombian National System of Protected Areas (SINAP) including the register Civil Society Natural Reserves reported to the Ministry of Environment until 2015." httpUri="https://zenodo.org/record/1441440/files/figure.png" pageId="7" pageNumber="122">Fig. 1C</figureCitation>
), in lateral view almost straight with a midventral tooth (
<figureCitation id="52B5A2FE5672FFBAFC49FA145C54FA13" box="[1017,1107,1446,1471]" captionStart="FIGURE 4" captionStartId="10.[151,250,1931,1953]" captionTargetBox="[206,1381,193,1921]" captionTargetId="figure@10.[206,1381,193,1921]" captionTargetPageId="10" captionText="FIGURE 4. Cora verapax sp. nov. holotype: a. head and thorax in dorsal and lateral views respectively; b. genital ligula in ventral view; c. S10 and cercus in dorsal view; d. S9–10 and cercus in lateral view; e. habitus in lateral view. a-l. p.: apicolateral processes, f. l. l.: filamentous lateral lobes, m-l. p.: midlateral processes. Photos: a, c–e: CABS; b: Camilo Flórez." httpUri="https://zenodo.org/record/1441446/files/figure.png" pageId="7" pageNumber="122">Fig. 4d</figureCitation>
). ParaproCts not developed. Measurements (mm). Total length 69.5, length of abdomen 57.1, FW 31.6 and HW 29.6 length.
</paragraph>
</subSubSection>
<subSubSection id="8294EDF05672FFBAFF77FA5C594FF877" pageId="7" pageNumber="122" type="diagnosis">
<paragraph id="CA31BE7B5672FFBAFF77FA5C594FF877" blockId="7.[151,1437,618,2011]" pageId="7" pageNumber="122">
<emphasis id="F8FA62695672FFBAFF77FA5C5A92F9AB" bold="true" box="[199,661,1518,1543]" pageId="7" pageNumber="122">
Notes on the
<emphasis id="F8FA62695672FFBAFED1FA5C599CF9AA" bold="true" box="[353,411,1518,1542]" italics="true" pageId="7" pageNumber="122">Cora</emphasis>
genus and diagnosis.
</emphasis>
The last phylogenetiC analysis (SánChez-Herrera
<emphasis id="F8FA62695672FFBAFB70FA5D5CFFF9AB" box="[1216,1272,1518,1543]" italics="true" pageId="7" pageNumber="122">et al.</emphasis>
2018) showed the genus
<emphasis id="F8FA62695672FFBAFEBDF9A15940F986" box="[269,327,1555,1578]" italics="true" pageId="7" pageNumber="122">Cora</emphasis>
as traditionally grouped is polyphyletiC in the light of the genetiC evidenCe. The
<taxonomicName id="0D8EC5F85672FFBAFB5FF9A65D37F987" baseAuthorityName="Selys" baseAuthorityYear="1853" box="[1263,1328,1556,1579]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="cyane">
<emphasis id="F8FA62695672FFBAFB5FF9A65D37F987" box="[1263,1328,1556,1579]" italics="true" pageId="7" pageNumber="122">cyane</emphasis>
</taxonomicName>
and
<taxonomicName id="0D8EC5F85672FFBAFADCF9A65D9BF987" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[1388,1436,1556,1579]" class="Insecta" family="Coenagrionidae" genus="Aeolagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="F8FA62695672FFBAFADCF9A65D9BF987" box="[1388,1436,1556,1579]" italics="true" pageId="7" pageNumber="122">inca</emphasis>
</taxonomicName>
groups proposed by
<bibRefCitation id="AE1FC38A5672FFBAFE32F9845A6FF9E3" box="[386,616,1590,1615]" pageId="7" pageNumber="122" refId="ref4441" refString="Bick, G. H. &amp; Bick, J. C. (1991) Two new damselflies: Cora dorada spec. nov. from Ecuador and C. parda spec. nov. from Peru (Zygoptera: Polythoridae). Odonatologica, 20 (4), 453 - 458." type="journal article">BiCk &amp; BiCk (1991)</bibRefCitation>
share several CharaCters but were separated beCause of the lateroapiCal projeCtions of the genital ligula observed in
<taxonomicName id="0D8EC5F85672FFBAFD31F9E95B8BF9DF" authority="(Selys, 1873)" authorityName="Selys" authorityYear="1873" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[641,908,1626,1651]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="F8FA62695672FFBAFD31F9E95AF6F9DF" box="[641,753,1627,1651]" italics="true" pageId="7" pageNumber="122">Cora inca</emphasis>
(Selys, 1873)
</taxonomicName>
. SánChez-Herrera
<emphasis id="F8FA62695672FFBAFBECF9EE5C93F9DF" box="[1116,1172,1626,1651]" italics="true" pageId="7" pageNumber="122">et al.</emphasis>
(2018) support them as part of a Clade Called
<emphasis id="F8FA62695672FFBAFE38F9CD59C5F93A" box="[392,450,1663,1686]" italics="true" pageId="7" pageNumber="122">Cora</emphasis>
s.s., grouping
<taxonomicName id="0D8EC5F85672FFBAFDD6F9CD5AF7F93A" authorityName="McLachlan" authorityYear="1881" box="[614,752,1663,1686]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="jocosa">
<emphasis id="F8FA62695672FFBAFDD6F9CD5AF7F93A" box="[614,752,1663,1686]" italics="true" pageId="7" pageNumber="122">Cora jocosa</emphasis>
</taxonomicName>
(MCLaChlan, 1881),
<taxonomicName id="0D8EC5F85672FFBAFC6DF9CD5CFBF93B" authority="(Selys, 1853)" authorityName="Selys" authorityYear="1853" baseAuthorityName="Selys" baseAuthorityYear="1853" box="[989,1276,1662,1687]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="cyane">
<emphasis id="F8FA62695672FFBAFC6DF9CD5C58F93A" box="[989,1119,1663,1686]" italics="true" pageId="7" pageNumber="122">Cora cyane</emphasis>
(Selys, 1853)
</taxonomicName>
,
<taxonomicName id="0D8EC5F85672FFBAFABAF9CD591EF917" authority="Selys, 1868" authorityName="Selys" authorityYear="1868" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="marina">
<emphasis id="F8FA62695672FFBAFABAF9CD5D9BF93A" box="[1290,1436,1663,1686]" italics="true" pageId="7" pageNumber="122">Cora marina</emphasis>
Selys, 1868
</taxonomicName>
,
<taxonomicName id="0D8EC5F85672FFBAFE97F9115A15F917" authority="Ris, 1918" authorityName="Ris" authorityYear="1918" box="[295,530,1698,1723]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="irene">
<emphasis id="F8FA62695672FFBAFE97F91159A7F917" box="[295,416,1699,1723]" italics="true" pageId="7" pageNumber="122">Cora irene</emphasis>
Ris, 1918
</taxonomicName>
,
<taxonomicName id="0D8EC5F85672FFBAFD90F9115B66F917" authority="Ris, 1918" authorityName="Ris" authorityYear="1918" box="[544,865,1698,1723]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="xanthostoma">
<emphasis id="F8FA62695672FFBAFD90F9115AE8F917" box="[544,751,1698,1723]" italics="true" pageId="7" pageNumber="122">Cora xanthostoma</emphasis>
Ris, 1918
</taxonomicName>
and
<taxonomicName id="0D8EC5F85672FFBAFC27F9115BEFF917" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[919,1000,1699,1723]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="F8FA62695672FFBAFC27F9115BEFF917" box="[919,1000,1699,1723]" italics="true" pageId="7" pageNumber="122">C. inca</emphasis>
</taxonomicName>
, a Clade morphologiCally supported by the plaCement of the FW nodus Closer to the pt than to the base of the wing, and the blue Coloration on dorsum of S2.
<taxonomicName id="0D8EC5F85672FFBAFF74F9595967F8AF" authorityName="Bota-Sierra &amp; Sánchez-Herrera &amp; Palacino-Rodríguez" authorityYear="2018" box="[196,352,1771,1795]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="verapax">
<emphasis id="F8FA62695672FFBAFF74F9595967F8AF" box="[196,352,1771,1795]" italics="true" pageId="7" pageNumber="122">Cora verapax</emphasis>
</taxonomicName>
shares the plaCement of its nodus and the blue Coloration on S2, with this Clade, whiCh Can be easily separated from the other speCies by the seCond segment of genital ligula, whiCh resembles
<emphasis id="F8FA62695672FFBAFB5CF8BD5D42F88A" box="[1260,1349,1807,1830]" italics="true" pageId="7" pageNumber="122">
<taxonomicName id="0D8EC5F85672FFBAFB5CF8BD5D46F88A" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[1260,1345,1807,1830]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="inca">C. inca</taxonomicName>
,
</emphasis>
but the apiColateral projeCtions are wider, by its middorsal horn on apex of S10 pointing in dorsoposterior direCtion, unique among the
<emphasis id="F8FA62695672FFBAFEA4F8E55949F8C2" box="[276,334,1879,1902]" italics="true" pageId="7" pageNumber="122">Cora</emphasis>
s.s. group.
<taxonomicName id="0D8EC5F85672FFBAFE61F8E55A6AF8C2" authorityName="Bota-Sierra &amp; Sánchez-Herrera &amp; Palacino-Rodríguez" authorityYear="2018" box="[465,621,1879,1902]" class="Insecta" family="Polythoridae" genus="Cora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="verapax">
<emphasis id="F8FA62695672FFBAFE61F8E55A6AF8C2" box="[465,621,1879,1902]" italics="true" pageId="7" pageNumber="122">Cora verapax</emphasis>
</taxonomicName>
also has two unique CharaCteristiCs in the family
<taxonomicName id="0D8EC5F85672FFBAFB2CF8E45D29F8C3" authorityName="Munz" authorityYear="1919" box="[1180,1326,1878,1903]" class="Insecta" family="Polythoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="family">Polythoridae</taxonomicName>
, a ventral thoraCiC tuberCle, and the abdomen almost two times longer than wings, whiCh is unusual for the family, resembling speCies of other families as
<taxonomicName id="0D8EC5F85672FFBAFDC4F82C5AF0F81B" box="[628,759,1950,1975]" class="Insecta" family="Perilestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="family">Perilestidae</taxonomicName>
or some genera in
<taxonomicName id="0D8EC5F85672FFBAFC5AF82C5C9CF81B" authorityName="Kirby" authorityYear="1890" box="[1002,1179,1950,1975]" class="Insecta" family="Coenagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
(e.g.,
<taxonomicName id="0D8EC5F85672FFBAFB5EF82C593CF877" authority="Sjostedt, 1918" authorityName="Sjostedt" authorityYear="1918" class="Insecta" family="Coenagrionidae" genus="Mesoleptobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="F8FA62695672FFBAFB5EF82C5D9BF81B" box="[1262,1436,1950,1975]" italics="true" pageId="7" pageNumber="122">Mesoleptobasis</emphasis>
Sjöstedt, 1918
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="9EF1EEF3567DFFB5FF27F83F5B01F84D" httpUri="https://zenodo.org/record/1441442/files/figure.png" pageId="8" pageNumber="123" startId="8.[151,250,1933,1955]" targetBox="[304,1261,205,1865]" targetPageId="8">
<paragraph id="CA31BE7B567DFFB5FF27F83F5B01F84D" blockId="8.[151,1435,1933,2017]" pageId="8" pageNumber="123">
<emphasis id="F8FA6269567DFFB5FF27F83F5913F80F" bold="true" box="[151,276,1933,1955]" pageId="8" pageNumber="123">FIGURE 2.</emphasis>
Protected areas reporting odonates from 2001 to 2017 in Colombia. a) Geographical distribution of the sampled areas. Polygons for SFF, RNSC, RFPN and No Register Reserves are too small to be noticed at the map scale. b) Number of sampled areas under the categories recognized by the SINAP.
</paragraph>
</caption>
<caption id="9EF1EEF3567CFFB4FF27FCBE5B4EFCF3" httpUri="https://zenodo.org/record/1441444/files/figure.png" pageId="9" pageNumber="124" startId="9.[151,250,780,802]" targetBox="[181,1410,197,764]" targetPageId="9">
<paragraph id="CA31BE7B567CFFB4FF27FCBE5B4EFCF3" blockId="9.[151,1437,780,863]" pageId="9" pageNumber="124">
<emphasis id="F8FA6269567CFFB4FF27FCBE590BFC8E" bold="true" box="[151,268,780,802]" pageId="9" pageNumber="124">FIGURE 3</emphasis>
. Taxonomical distribution of odonates species reported in Colombian protected areas a) literature compilation from 2001 to 2017 and new records reported in this work b) new records based on six protected areas surveyed in this study. Numbers at the end of the bars correspond to the number of species.
</paragraph>
</caption>
<subSubSection id="8294EDF0567CFFB4FF77FC3E5BC0FC64" pageId="9" pageNumber="124" type="distribution">
<paragraph id="CA31BE7B567CFFB4FF77FC3E5BC0FC64" blockId="9.[151,1437,908,1112]" pageId="9" pageNumber="124">
<emphasis id="F8FA6269567CFFB4FF77FC3E5967FC09" bold="true" box="[199,352,908,933]" pageId="9" pageNumber="124">Distribution.</emphasis>
The speCies is only known from its
<typeStatus id="153500D9567CFFB4FD4DFC3F5B2AFC09" box="[765,813,909,933]" pageId="9" pageNumber="124">type</typeStatus>
loCality at National Natural Park Tatamá in the PaCifiC OCean slope of the Andean Cordillera OCCidental (
<collectingRegion id="084A7099567CFFB4FD7EFC1D5B3BFC64" box="[718,828,943,968]" country="Colombia" name="Risaralda" pageId="9" pageNumber="124">Risaralda</collectingRegion>
,
<collectingCountry id="B299FEEB567CFFB4FCF7FC1D5BBCFC64" box="[839,955,943,968]" name="Colombia" pageId="9" pageNumber="124">Colombia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="8294EDF0567CFFB4FF77FC665A30FBF4" pageId="9" pageNumber="124" type="biology_ecology">
<paragraph id="CA31BE7B567CFFB4FF77FC665A30FBF4" blockId="9.[151,1437,908,1112]" pageId="9" pageNumber="124">
<emphasis id="F8FA6269567CFFB4FF77FC665920FC41" bold="true" box="[199,295,980,1005]" pageId="9" pageNumber="124">Biology.</emphasis>
The biology of this speCies remains a mystery, 13 months of sampling were aCCumulated from
<quantity id="0D76139E567CFFB4FAD4FC66590FFBBC" metricMagnitude="1" metricUnit="m" metricValue="5.11937" metricValueMax="5.12318" metricValueMin="5.11556" pageId="9" pageNumber="124" unit="in" value="2015.5" valueMax="2017.0" valueMin="2014.0">
<date id="BE3098BB567CFFB4FAD4FC6658ECFBBC" pageId="9" pageNumber="124" value="2014" valueMax="2017">2014 to 2017</date>
in
</quantity>
the
<typeStatus id="153500D9567CFFB4FE87FC4A5960FBBC" box="[311,359,1016,1040]" pageId="9" pageNumber="124">type</typeStatus>
loCality and only one speCimen was observed and fortunately Caught, showing this is a very rare speCies. The only known speCimen was at a stream in a well-preserved Cloud mountain forest where this speCies probably inhabits in the high Canopy.
</paragraph>
</subSubSection>
</treatment>
</document>
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