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<document id="6088EA6632953BC149AEAE10BF531507" ID-DOI="10.5281/zenodo.281994" ID-GBIF-Dataset="9ed4f1e5-ffdd-4db6-b83f-bff541492839" ID-ISSN="1175-5326" ID-Zenodo-Dep="281994" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460507760580" checkinUser="plazi" docAuthor="T, Duniesky Rios &amp; Goloboff, Pablo A." docDate="2012" docId="03C43846FFCD22736AC2FAA27D84FC53" docLanguage="en" docName="zt03422p051.pdf" docOrigin="Zootaxa 3422" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mediothele Raven &amp; Platnick 1978" docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="FFFD403EFFC4227A6A55FA7D7E30FFB8" masterDocTitle="New species of Chilean Hexathelidae (Araneae, Mygalomorphae)" masterLastPageNumber="51" masterPageNumber="32" pageNumber="41" updateTime="1698312978760" updateUser="plazi">
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<mods:title id="56025F39644A7C15A7C26E1DB39ED4E5">New species of Chilean Hexathelidae (Araneae, Mygalomorphae)</mods:title>
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<mods:namePart id="8825636D5594C579387700C9A2B100FE">T, Duniesky Rios</mods:namePart>
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<mods:namePart id="16B84BCA0691219A35DAFCB4B51930FF">Goloboff, Pablo A.</mods:namePart>
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<mods:date id="65CC87B2AE6D013737DE292C8DEB369D">2012</mods:date>
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<mods:identifier id="11824B69696529F1E708E876C7477BC4" type="DOI">10.5281/zenodo.281994</mods:identifier>
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<mods:identifier id="9082B545CB13AAE3892C4C76956AED2B" type="Zenodo-Dep">281994</mods:identifier>
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<subSubSection id="C377DADBFFCD22736AC2FAA27EF5FDAE" pageId="9" pageNumber="41" type="nomenclature">
<paragraph id="8BD28950FFCD22736AC2FAA27CB4FF41" blockId="9.[151,644,223,249]" box="[151,644,223,249]" pageId="9" pageNumber="41">
<heading id="D09A3E3CFFCD22736AC2FAA27CB4FF41" bold="true" box="[151,644,223,249]" fontSize="11" level="1" pageId="9" pageNumber="41" reason="1">
<treatmentCitationGroup id="AB7DAE7EFFCD22736AC2FAA27CB4FF41" box="[151,644,223,249]" pageId="9" pageNumber="41">
<emphasis id="B9195542FFCD22736AC2FAA27CB4FF41" bold="true" box="[151,644,223,249]" pageId="9" pageNumber="41">
<taxonomicName id="4C6DF2D3FFCD22736AC2FAA27C4DFF41" ID-CoL="62ZJQ" ID-ENA="1956467" authority="Raven &amp; Platnick, 1978: 74" authorityName="Raven &amp; Platnick" authorityPageNumber="74" authorityYear="1978" box="[151,637,223,249]" class="Arachnida" family="Hexathelidae" genus="Mediothele" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9195542FFCD22736AC2FAA27F2EFF41" bold="true" box="[151,286,223,249]" italics="true" pageId="9" pageNumber="41">Mediothele</emphasis>
<treatmentCitation id="0ACCAF41FFCD22736B70FAA27C4DFF41" author="Raven" box="[293,637,223,249]" page="74" pageId="9" pageNumber="41" year="1978">
<bibRefCitation id="EFFCF4A1FFCD22736B70FAA27C62FF41" author="Raven" box="[293,594,223,249]" pageId="9" pageNumber="41" refString="Raven, R. J. &amp; N. I. Platnick (1978) A new genus of the spider family Dipluridae from Chile (Araneae, Mygalomorphae). Journal of Arachnology, 6, 73 - 77." type="journal article" year="1978">Raven &amp; Platnick, 1978</bibRefCitation>
: 74
</treatmentCitation>
</taxonomicName>
.
</emphasis>
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</heading>
</paragraph>
<paragraph id="8BD28950FFCD22736AC2FB5B7A7BFE3F" blockId="9.[151,1436,294,1003]" pageId="9" pageNumber="41">
Until now, the genus
<taxonomicName id="4C6DF2D3FFCD22736BD2FB5B7C35FE86" box="[391,517,294,318]" class="Arachnida" family="Hexathelidae" genus="Mediothele" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9195542FFCD22736BD2FB5B7C35FE86" box="[391,517,294,318]" italics="true" pageId="9" pageNumber="41">Mediothele</emphasis>
</taxonomicName>
included a single species,
<taxonomicName id="4C6DF2D3FFCD22736964FB557D89FE86" box="[817,953,294,319]" class="Arachnida" family="Hexathelidae" genus="Mediothele" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B9195542FFCD22736964FB557D89FE86" box="[817,953,294,319]" italics="true" pageId="9" pageNumber="41">M. australis</emphasis>
</taxonomicName>
, described from a single male from Southern
<collectingCountry id="F37AC9C0FFCD22736A91FB377F34FEDA" box="[196,260,330,354]" name="Chile" pageId="9" pageNumber="41">Chile</collectingCountry>
; no further males are known of the genus. The genus was originally included in
<taxonomicName id="4C6DF2D3FFCD22736EE6FB377B1EFEDA" box="[1203,1326,330,354]" class="Arachnida" family="Dipluridae" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="family">Dipluridae</taxonomicName>
, but later transferred (
<bibRefCitation id="EFFCF4A1FFCD22736B4AFB137F9FFE3E" author="Raven" box="[287,431,366,390]" pageId="9" pageNumber="41" refString="Raven, R. J. (1985) The spider infraorder Mygalomorphae (Araneae): Cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven, 1985</bibRefCitation>
) to
<taxonomicName id="4C6DF2D3FFCD22736B8EFB137C42FE3E" box="[475,626,366,390]" class="Arachnida" family="Hexathelidae" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="family">Hexathelidae</taxonomicName>
, proposing
<taxonomicName id="4C6DF2D3FFCD227368A1FB127D4AFE3E" box="[756,890,367,390]" class="Arachnida" family="Hexathelidae" genus="Scotinoecus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9195542FFCD227368A1FB127D4AFE3E" box="[756,890,367,390]" italics="true" pageId="9" pageNumber="41">Scotinoecus</emphasis>
</taxonomicName>
as its sister group.
</paragraph>
<paragraph id="8BD28950FFCD22736A9CFBEE7EF5FDAE" blockId="9.[151,1436,294,1003]" pageId="9" pageNumber="41">
In this genus, morphological differences between species are more marked than in
<taxonomicName id="4C6DF2D3FFCD22736E2CFBEE7ACFFE12" box="[1145,1279,403,426]" class="Arachnida" family="Hexathelidae" genus="Scotinoecus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9195542FFCD22736E2CFBEE7ACFFE12" box="[1145,1279,403,426]" italics="true" pageId="9" pageNumber="41">Scotinoecus</emphasis>
</taxonomicName>
, especially in leg, cephalothorax and sternum shape, and chaetotaxy. The much simpler spermathecae have only minor differences between species (often within the observed range of individual variation among specimens of a single locality).
</paragraph>
</subSubSection>
<subSubSection id="C377DADBFFCD22736A92F85E7D54FD82" box="[199,868,546,571]" pageId="9" pageNumber="41" type="materials_examined">
<paragraph id="8BD28950FFCD22736A92F85E7D54FD82" blockId="9.[151,1436,294,1003]" box="[199,868,546,571]" pageId="9" pageNumber="41">
<emphasis id="B9195542FFCD22736A92F85E7F52FD83" bold="true" box="[199,354,546,571]" pageId="9" pageNumber="41">
<typeStatus id="54D637F2FFCD22736A92F85E7F32FD83" box="[199,258,547,571]" pageId="9" pageNumber="41">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6DF2D3FFCD22736B3CF85E7D54FD82" authority="Raven &amp; Platnick, 1978" authorityName="Raven &amp; Platnick" authorityYear="1978" box="[361,868,546,571]" class="Arachnida" family="Hexathelidae" genus="Mediothele" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B9195542FFCD22736B3CF85E7C7FFD83" box="[361,591,547,571]" italics="true" pageId="9" pageNumber="41">Mediothele australis</emphasis>
<bibRefCitation id="EFFCF4A1FFCD22736802F85E7D54FD82" author="Raven" box="[599,868,546,570]" pageId="9" pageNumber="41" refString="Raven, R. J. &amp; N. I. Platnick (1978) A new genus of the spider family Dipluridae from Chile (Araneae, Mygalomorphae). Journal of Arachnology, 6, 73 - 77." type="journal article" year="1978">Raven &amp; Platnick, 1978</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C377DADBFFCD22736A92F83B7D84FC53" pageId="9" pageNumber="41" type="diagnosis">
<paragraph id="8BD28950FFCD22736A92F83B7F3DFD56" blockId="9.[151,1436,294,1003]" pageId="9" pageNumber="41">
<emphasis id="B9195542FFCD22736A92F83B7F71FDE7" bold="true" box="[199,321,582,607]" pageId="9" pageNumber="41">Diagnosis.</emphasis>
Differs from
<taxonomicName id="4C6DF2D3FFCD22736BB4F83A7C57FDE6" box="[481,615,583,606]" class="Arachnida" family="Hexathelidae" genus="Scotinoecus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9195542FFCD22736BB4F83A7C57FDE6" box="[481,615,583,606]" italics="true" pageId="9" pageNumber="41">Scotinoecus</emphasis>
</taxonomicName>
in the labium and maxillae having few (female) or no (male) cuspules; fangs with basal outer tooth. PLS spinnerets short, ALS two-segmented much shorter than PMS; thickened femora III; patellae III with strong prolateral spines; patellae III and IV with many prolateral and dorsal anterior strong setae. Male tibia I with one large and one small spur, and associated thorn on metatarsus; bulb pyriform, without conductor.
</paragraph>
<paragraph id="8BD28950FFCD22736A92F8867DA3FC8E" blockId="9.[151,1436,294,1003]" pageId="9" pageNumber="41">Many of these somatic characters (fang tooth, shorter spinnerets, thickened femora, and patellae with strong spines and setae) are probably associated with a burrowing lifestyle.</paragraph>
<paragraph id="8BD28950FFCD22736A92F93F7AFEFCE2" blockId="9.[151,1436,294,1003]" box="[199,1230,834,859]" pageId="9" pageNumber="41">
<emphasis id="B9195542FFCD22736A92F93F7F2DFCE3" bold="true" box="[199,285,834,859]" pageId="9" pageNumber="41">Habits.</emphasis>
Spiders of all known species live in small silk-lined burrows with an open entrance.
</paragraph>
<paragraph id="8BD28950FFCD22736A92F91A7D84FC53" blockId="9.[151,1436,294,1003]" pageId="9" pageNumber="41">
<emphasis id="B9195542FFCD22736A92F91A7F65FCC7" bold="true" box="[199,341,871,895]" pageId="9" pageNumber="41">Monophyly.</emphasis>
By out-group comparison, essentially all of the characters in which it differs from
<emphasis id="B9195542FFCD22736EAEF91A7B43FCC6" box="[1275,1395,871,894]" italics="true" pageId="9" pageNumber="41">Scotinecus</emphasis>
are synapomorphies of
<taxonomicName id="4C6DF2D3FFCD22736B22F9F67FC5FC1B" box="[375,501,907,931]" class="Arachnida" family="Hexathelidae" genus="Mediothele" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9195542FFCD22736B22F9F67FC5FC1B" box="[375,501,907,931]" italics="true" pageId="9" pageNumber="41">Mediothele</emphasis>
</taxonomicName>
. The monophyly of the genus is thus well established. The low numbers of labial (and maxillary) cuspules are best interpreted as a reversal, as (according to Ravens 1978, 1985 hypotheses of hexathelid relationships) they occur in several successive sister groups.
</paragraph>
</subSubSection>
</treatment>
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