297 lines
35 KiB
XML
297 lines
35 KiB
XML
<document id="374829286B2E840563D39DBB4C60C82E" ID="10.11646/zootaxa.4139.1.4" ID-DOI="10.11646/zootaxa.4139.1.4" ID-GBIF-Dataset="3cf8edb7-4be5-4ec3-803e-ca1f224d703b" ID-ISSN="1175-5326" ID-Zenodo-Dep="271999" ID-ZooBank="2744EC46-70A1-4267-AAB8-2131F5DC21DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468909435473" checkinUser="plazi" docAuthor="Nunes, Rafael V. & Vaz-De-Mello, Fernando Z." docDate="2016" docId="03C187AE4532FF8A0EC8F88DFD16FAAF" docLanguage="en" docName="zootaxa.4139.1.4.pdf" docOrigin="Zootaxa 4139 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichotomius (Selenocopris) paresi Nunes & Vaz-De-Mello, 2016, new species" docType="treatment" docVersion="7" masterDocId="FFF8FFD6453BFF810E5FFF9FFF9CFFFA" masterDocTitle="New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus" updateTime="1698365276952" updateUser="plazi">
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<mods:mods id="BCABE5E7E445E3E0849799772E70CE7A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="92BC4A9490B094958A51D03B0EBB20AB">
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<mods:title id="FA99189F13E1F4EA1E0577C2E5BD8373">New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus</mods:title>
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</mods:titleInfo>
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<mods:name id="E1DFFD19E080CB17A580AF624A532451" type="personal">
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<mods:role id="95D3E92FED7E3770B3FB4727E3961EDD">
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<mods:roleTerm id="AA09D6CC66B564DCD7BE29B5A377A645">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="18CA124A66F058BADEF69412D1B455EE">Nunes, Rafael V.</mods:namePart>
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</mods:name>
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<mods:name id="5A75B6E19C0C1428AA9AB21ECA98B952" type="personal">
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<mods:role id="DE474FCD4555B6EEE76B7ECB98213D82">
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<mods:roleTerm id="64541C52335A5A9F2E82D76FF77FE8EA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="35A349184FC1EA9A1FEC89B9F0196839">Vaz-De-Mello, Fernando Z.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1CA98D94BAF354020339C421DC7A3995">text</mods:typeOfResource>
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<mods:relatedItem id="4D9A0064BC9706E35B26B8E3A8D70174" type="host">
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<mods:titleInfo id="234EA1DB9B3AE80BA29973D916105EC3">
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<mods:title id="B2B11029EE9A81561AB4DBBD035FE63A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BBFA14D6276F74732AB197B61C1B46FE">
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<mods:date id="6E273F7ED6B1F25F53C11A31CA33FCFB">2016</mods:date>
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<mods:detail id="C6678DFBB5AB401B2432625FF95C5F86" type="volume">
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<mods:number id="128736818F611F63E6831DF10A034D41">4139</mods:number>
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</mods:detail>
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<mods:detail id="42840447F31648590235B4D34E87304D" type="issue">
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<mods:number id="42C65BC306F9FF7D2585710B904E983A">1</mods:number>
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</mods:detail>
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||
</mods:part>
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</mods:relatedItem>
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<mods:classification id="8DC2E796E9483962584E650165B63156">journal volume</mods:classification>
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<mods:identifier id="80F523B7CD6FABE409CAEAC394F4732F" type="DOI">10.11646/zootaxa.4139.1.4</mods:identifier>
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<mods:identifier id="C672786D69EAA2E803F897235B60FA7F" type="GBIF-Dataset">3cf8edb7-4be5-4ec3-803e-ca1f224d703b</mods:identifier>
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<mods:identifier id="518B5B275AFA6CE9F0C443AD33C724D7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0235F753D57CB9C7C65562DDA1B92EA6" type="Zenodo-Dep">271999</mods:identifier>
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<mods:identifier id="004135AA645654AA01152A109AFEC694" type="ZooBank">2744EC46-70A1-4267-AAB8-2131F5DC21DC</mods:identifier>
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</mods:mods>
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<treatment id="03C187AE4532FF8A0EC8F88DFD16FAAF" ID-DOI="http://doi.org/10.5281/zenodo.6090775" ID-GBIF-Taxon="123198331" ID-Zenodo-Dep="6090775" LSID="urn:lsid:plazi:treatment:03C187AE4532FF8A0EC8F88DFD16FAAF" httpUri="http://treatment.plazi.org/id/03C187AE4532FF8A0EC8F88DFD16FAAF" lastPageId="11" pageId="9">
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<subSubSection id="C37265334532FF880EC8F88DFF77F8B7" pageId="9" type="nomenclature">
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||
<paragraph id="8BD736B84532FF880EC8F88DFD53F8D6" blockId="9.[151,719,1810,1869]" box="[151,719,1810,1836]" pageId="9">
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||
<heading id="D09F81D44532FF880EC8F88DFD53F8D6" bold="true" box="[151,719,1810,1836]" fontSize="11" level="1" pageId="9" reason="1">
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||
<emphasis id="B91CEAAA4532FF880EC8F88DFD53F8D6" bold="true" box="[151,719,1810,1836]" pageId="9">
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||
<taxonomicName id="4C684D3B4532FF880EC8F88DFDA4F8D6" box="[151,568,1810,1836]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="paresi" status="sp. nov." subGenus="Selenocopris">
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<emphasis id="B91CEAAA4532FF880EC8F88DFDA4F8D6" bold="true" box="[151,568,1810,1836]" italics="true" pageId="9">Dichotomius (Selenocopris) paresi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F57D14532FF880C60F88DFD53F8D6" box="[575,719,1810,1836]" pageId="9" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BD736B84532FF880EC8F8ABFF77F8B7" blockId="9.[151,719,1810,1869]" box="[151,235,1844,1869]" pageId="9">
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||
(
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<figureCitation id="13532A3D4532FF880EC0F8ABFF7FF8B7" box="[159,227,1844,1869]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="9">Fig. 6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265334532FF8B0EC8F8E3FBDDFF02" lastPageId="10" pageId="9" type="materials_examined">
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<paragraph id="8BD736B84532FF880EC8F8E3FA00F824" blockId="9.[151,1437,1916,2014]" pageId="9">
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<emphasis id="B91CEAAA4532FF880EC8F8E3FE6FF86F" bold="true" box="[151,499,1916,1941]" pageId="9">
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Material studied.
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<typeStatus id="54D3881A4532FF880F22F8E3FE72F86F" box="[381,494,1916,1941]" pageId="9" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Ƌ
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<collectingCountry id="F37F76284532FF880C77F8E2FD0CF86F" box="[552,656,1917,1941]" name="Brazil" pageId="9">BRASIL</collectingCountry>
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:
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<emphasis id="B91CEAAA4532FF880CFBF8E3FCCCF86F" bold="true" box="[676,848,1916,1941]" pageId="9">
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<collectingRegion id="49ACF85A4532FF880CFBF8E3FCD0F86F" box="[676,844,1916,1941]" country="Brazil" name="Mato Grosso" pageId="9">Mato Grosso</collectingRegion>
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:
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</emphasis>
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Nova Marilândia. Fazenda Aparecida da Serra.
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<geoCoordinate id="EE5C507F4532FF880EC8F83EFE82F843" box="[151,286,1953,1977]" direction="south" orientation="latitude" pageId="9" precision="15" value="-14.353889">14°21’14”S</geoCoordinate>
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;
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<geoCoordinate id="EE5C507F4532FF880F75F83EFE26F840" box="[298,442,1953,1978]" direction="west" orientation="longitude" pageId="9" precision="15" value="-57.731667">57°43’54”W</geoCoordinate>
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. Human feces.
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<date id="FFD610784532FF880C39F83EFD45F843" box="[614,729,1953,1977]" pageId="9" value="2010-01-17">17-I-2010</date>
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. RJSilva (leg.) /
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<taxonomicName id="4C684D3B4532FF880DCBF83EFBF3F843" box="[916,1135,1953,1977]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="paresi" status="sp. nov.">
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<emphasis id="B91CEAAA4532FF880DCBF83EFBF3F843" box="[916,1135,1953,1977]" italics="true" pageId="9">Dichotomius paresi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F57D14532FF880A29F83DFB2DF843" box="[1142,1201,1954,1977]" pageId="9" rank="species">n. sp.</taxonomicNameLabel>
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,
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<typeStatus id="54D3881A4532FF880A9FF83EFABAF840" box="[1216,1318,1953,1978]" pageId="9" type="holotype">holotype</typeStatus>
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. RVNunes & FZVaz-de-Mello, 2011 [
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<collectionCode id="ED79AE7D4532FF880F9CF859FD85F824" box="[451,537,1990,2014]" pageId="9">CEMT</collectionCode>
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].
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<emphasis id="B91CEAAA4532FF880C74F85AFD30F824" bold="true" box="[555,684,1989,2014]" pageId="9">
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<typeStatus id="54D3881A4532FF880C74F85AFD3BF824" box="[555,679,1989,2014]" pageId="9" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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same data of
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<typeStatus id="54D3881A4532FF880D14F85AFC31F824" box="[843,941,1989,2014]" pageId="9" type="holotype">holotype</typeStatus>
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[2 Ƌ,
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<specimenCount id="9D6EFD314532FF880DA8F859FB87F824" box="[1015,1051,1990,2014]" pageId="9" type="female">2♀</specimenCount>
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,
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<collectionCode id="ED79AE7D4532FF880A76F859FBE1F824" box="[1065,1149,1990,2014]" pageId="9">CEMT</collectionCode>
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]; Nova Marilândia, 2008.
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</paragraph>
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<paragraph id="8BD736B84531FF8B0EC8FF08FBDDFF02" blockId="10.[151,1437,151,392]" pageId="10">
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Cerrado
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<emphasis id="B91CEAAA4531FF8B0EA0FF06FE06FF55" box="[255,410,152,176]" italics="true" pageId="10">sensu stricto.</emphasis>
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Pitfall s/ isca. Tombo 13 [7 Ƌ,
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<specimenCount id="9D6EFD314531FF8B0D6EFF07FCC8FF4A" box="[817,852,152,176]" pageId="10" type="female">2♀</specimenCount>
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<collectionCode id="ED79AE7D4531FF8B0D3DFF07FC2AFF4A" box="[866,950,152,176]" pageId="10">CEMT</collectionCode>
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]; Same but Fazenda Aparados da Serra.
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<geoCoordinate id="EE5C507F4531FF8B0EC8FF23FE83FF2E" box="[151,287,188,212]" direction="south" orientation="latitude" pageId="10" precision="15" value="-14.366944">14°22’01”S</geoCoordinate>
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;
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<geoCoordinate id="EE5C507F4531FF8B0F74FF23FE24FF2F" box="[299,440,188,213]" direction="east" orientation="longitude" pageId="10" precision="15" value="57.725002">57°43’30”O</geoCoordinate>
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. 575mosl. Cerrado
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<emphasis id="B91CEAAA4531FF8B0CCBFF22FD4DFF2E" box="[660,721,189,212]" italics="true" pageId="10">s. str.</emphasis>
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Pitfall s. isca. 8-
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<date id="FFD610784531FF8B0DD1FF23FC43FF2F" box="[910,991,188,213]" pageId="10" value="1916-02-02">16-II-2</date>
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0 0 8 [2 Ƌ
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<specimenCount id="9D6EFD314531FF8B0A12FF23FBE4FF2E" box="[1101,1144,188,212]" pageId="10" type="female">1 ♀</specimenCount>
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<collectionCode id="ED79AE7D4531FF8B0ADFFF23FB48FF2E" box="[1152,1236,188,212]" pageId="10">CEMT</collectionCode>
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]; same but
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<quantity id="4C909B5D4531FF8B0B0BFF23FA00FF2F" box="[1364,1436,188,213]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" unit="km" value="30.0">30 km</quantity>
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NW Tangará da Serra.
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<date id="FFD610784531FF8B0FC7FF7FFDAEFF02" box="[408,562,224,248]" pageId="10" value="2012-12-03" valueMax="2012-12-09" valueMin="2012-12-03">3–9.XII.2012</date>
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. Vaz-de-Mello, Pokorný & Beneč [5 Ƌ
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<collectionCode id="ED79AE7D4531FF8B0DB0FF7FFBABFF02" box="[1007,1079,224,248]" pageId="10">SPPC</collectionCode>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265334531FF8B0E98FE9BFC95FE72" pageId="10" type="diagnosis">
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<paragraph id="8BD736B84531FF8B0E98FE9BFC95FE72" blockId="10.[151,1437,151,392]" pageId="10">
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<emphasis id="B91CEAAA4531FF8B0E98FE9BFEDDFEE7" bold="true" box="[199,321,260,285]" pageId="10">Diagnosis.</emphasis>
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In the group,
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<taxonomicName id="4C684D3B4531FF8B0C5BFE9AFDE8FEE6" box="[516,628,261,284]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="paresi" status="sp. nov.">
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<emphasis id="B91CEAAA4531FF8B0C5BFE9AFDE8FEE6" box="[516,628,261,284]" italics="true" pageId="10">D. paresi</emphasis>
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</taxonomicName>
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<emphasis id="B91CEAAA4531FF8B0CDBFE9BFD50FEE7" bold="true" box="[644,716,260,285]" pageId="10">
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<taxonomicNameLabel id="A22F57D14531FF8B0CDBFE9BFD50FEE7" box="[644,716,260,285]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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is separated of other species in the group (including the brachypterous) by the following combined characters: posterior portion of the pronotal disc bearing ocellate punctures (
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<figureCitation id="13532A3D4531FF8B0F4BFED3FEFEFE9F" box="[276,354,332,357]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="10">Figs. 6</figureCitation>
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A–B); elytral striae widened near the basis and lacking punctures (
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<figureCitation id="13532A3D4531FF8B0A09FED3FB05FE9F" box="[1110,1177,332,357]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="10">Fig. 6</figureCitation>
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E) and anterior portion of pronotal border flattened (dorsal view) (
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<figureCitation id="13532A3D4531FF8B0C29FEF0FD58FE72" box="[630,708,367,392]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="10">Figs. 6</figureCitation>
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A–B).
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</paragraph>
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</subSubSection>
|
||
<caption id="DF1766304531FF8B0EC8FACDFF4EFA3E" httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="10" targetBox="[283,1306,458,1329]" targetPageId="10">
|
||
<paragraph id="8BD736B84531FF8B0EC8FACDFF4EFA3E" blockId="10.[151,1436,1361,1476]" pageId="10">
|
||
<emphasis id="B91CEAAA4531FF8B0EC8FACDFE8FFA9D" bold="true" box="[151,275,1362,1384]" pageId="10">FIGURE 6.</emphasis>
|
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Paratypes of
|
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<taxonomicName id="4C684D3B4531FF8B0FFEFACDFC9EFA92" box="[417,770,1362,1384]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="paresi" status="sp. nov." subGenus="Selenocopris">
|
||
<emphasis id="B91CEAAA4531FF8B0FFEFACDFC9EFA92" box="[417,770,1362,1384]" italics="true" pageId="10">Dichotomius (Selenocopris) paresi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CEAAA4531FF8B0D55FACDFCD8FA9D" bold="true" box="[778,836,1362,1383]" pageId="10">
|
||
<taxonomicNameLabel id="A22F57D14531FF8B0D55FACDFCD8FA9D" box="[778,836,1362,1383]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B91CEAAA4531FF8B0D0DFACEFCEAFA92" bold="true" box="[850,886,1361,1384]" pageId="10">(A)</emphasis>
|
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male dorsal habitus;
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||
<emphasis id="B91CEAAA4531FF8B0A0DFACEFBE8FA92" bold="true" box="[1106,1140,1361,1384]" pageId="10">(B)</emphasis>
|
||
female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc);
|
||
<emphasis id="B91CEAAA4531FF8B0A5AFAEFFBB4FA7D" bold="true" box="[1029,1064,1392,1415]" pageId="10">(C)</emphasis>
|
||
dorsal view of male paramera;
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||
<emphasis id="B91CEAAA4531FF8B0B26FAEFFA00FA7D" bold="true" box="[1401,1436,1392,1415]" pageId="10">(D)</emphasis>
|
||
lateral view of male paramera;
|
||
<emphasis id="B91CEAAA4531FF8B0F8BFA11FE6AFA5F" bold="true" box="[468,502,1422,1445]" pageId="10">(E)</emphasis>
|
||
detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37265334531FF8A0E98FA74FB98FC5E" lastPageId="11" pageId="10" type="materials_examined">
|
||
<paragraph id="8BD736B84531FF8A0E98FA74FE0AFD52" blockId="10.[151,1437,1515,2008]" lastBlockId="11.[151,1437,151,1365]" lastPageId="11" pageId="10">
|
||
<emphasis id="B91CEAAA4531FF8B0E98FA74FE17F9FE" bold="true" box="[199,395,1515,1540]" pageId="10">
|
||
<typeStatus id="54D3881A4531FF8B0E98FA74FEAAF9FE" box="[199,310,1515,1540]" pageId="10" type="holotype">Holotype</typeStatus>
|
||
. male:
|
||
</emphasis>
|
||
length: 17.0 mm. Maximum width (pronotum):
|
||
<quantity id="4C909B5D4531FF8B0D8DFA74FBA3F9F9" box="[978,1087,1515,1540]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="10" unit="mm" value="10.5">10.5 mm</quantity>
|
||
. Black, shinny.
|
||
<emphasis id="B91CEAAA4531FF8B0B5EFA74FAD7F9FE" bold="true" box="[1281,1355,1515,1540]" pageId="10">Head:</emphasis>
|
||
dorsal surface smooth, bearing few occellate puncturess close to the internal border of the eyes. Clypeogenal junction obtuse. Margins of clypeal teeth bearing a single row of short scattered setae. Ventral clypeal process strongly bifurcated. Cephalic process smooth, consisting of a single conical horn. Clypeo-genal suture distinct, producing a posterior knob. Antennal club with dark brown tumescence.
|
||
<emphasis id="B91CEAAA4531FF8B0D63F9E4FC22F96E" bold="true" box="[828,958,1659,1684]" pageId="10">Pronotum:</emphasis>
|
||
wider than long and sub equal to elytra on width. Anterior excavation bearing ocellate punctures very close to each other. A bilobed projection delimits the pronotal disc from the anterior declivity. Pronotal disc shining: smooth anteriorly and bearing ocellate punctures posteriorly – the diameter of the punctures increase toward the posterior margin of pronotum. Ocellate punctures also present on pronotal margins and at the anterior angles. Anterior angles acute. Anterior portion of pronotal sides flattened (dorsal view).
|
||
<emphasis id="B91CEAAA4531FF8B0FA6F8B0FD0DF8B2" bold="true" box="[505,657,1839,1864]" pageId="10">Hypomeron:</emphasis>
|
||
anterior portion with a lateral band of ocellate setose punctures. Posterior portion almost glabrous with few sparse ocellate setose punctures.
|
||
<emphasis id="B91CEAAA4531FF8B0A49F8CCFB32F896" bold="true" box="[1046,1198,1875,1900]" pageId="10">Prosternum:</emphasis>
|
||
with ocellate setose punctures: setae do not reach one quarter the length of hypomeron setae.
|
||
<emphasis id="B91CEAAA4531FF8B0A55F8E8FB2BF86A" bold="true" box="[1034,1207,1911,1936]" pageId="10">Mesosternum:</emphasis>
|
||
strongly narrowed, glabrous and chagrinated medially.
|
||
<emphasis id="B91CEAAA4531FF8B0C73F804FD72F84E" bold="true" box="[556,750,1947,1972]" pageId="10">Mesepisternum:</emphasis>
|
||
having ocellate setose punctures. Posterior portion of upper margin ending in an acute angle and reaching pseudoepipleuron (
|
||
<figureCitation id="13532A3D4531FF8B0DD2F820FC45F822" box="[909,985,1983,2008]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="10">Figs. 1</figureCitation>
|
||
B–C).
|
||
<emphasis id="B91CEAAA4531FF8B0A74F820FB4AF822" bold="true" box="[1067,1238,1983,2008]" pageId="10">Metasternum:</emphasis>
|
||
mesometasternal suture indistinct. Anterior lobe strong narrowed medially. Ocellate setose puncture present along the margins of the metasternal lobe – these points became denser around the anterior margin. Mestasternal disc bearing a longitudinal sulcus that ends in a feeble posterior concavity.
|
||
<emphasis id="B91CEAAA4530FF8A0CEEFF40FC9BFF02" bold="true" box="[689,775,223,248]" pageId="11">Elytra:</emphasis>
|
||
black, shinny, lacking humeral callus. Striae widened near the elytral basis, lacking punctures, at most with some indication of points near the elytra apex (
|
||
<figureCitation id="13532A3D4530FF8A0ABDFE9BFABBFEE7" box="[1250,1319,260,285]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="11">Fig. 6</figureCitation>
|
||
E). Elytral interstriae feebly convex.
|
||
<emphasis id="B91CEAAA4530FF8A0F91FEB8FDFAFEBA" bold="true" box="[462,614,295,320]" pageId="11">Hind wings:</emphasis>
|
||
length:
|
||
<quantity id="4C909B5D4530FF8A0C8FFEB7FC86FEBA" box="[720,794,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="11" unit="mm" value="11.0">11mm</quantity>
|
||
. Having folding articulation. Basal sclerites glabrous. Subcostal vein intersecting costal axis at its basal quarter. Medial vein extending to the folding articulation. Distinct setae present on the basal third of costal axis and on folding articulation. Cubital vein present, anal vein absent. Small developed membranous portion apical to folding articulation with incomplete or weakly-marked venations.
|
||
<emphasis id="B91CEAAA4530FF8A0F4AFE28FECAFE2A" bold="true" box="[277,342,439,464]" pageId="11">Legs:</emphasis>
|
||
anterior tibiae spur curved inward at apical third. Anterior tibiae bearing four teeth. Ventral surface of the anterior femur glabrous. Length of 5th metatarsomere twice the length of 4th metatarsomere.
|
||
<emphasis id="B91CEAAA4530FF8A0EC8FE60FE89FDE2" bold="true" box="[151,277,511,536]" pageId="11">Abdomen:</emphasis>
|
||
sternites 1 to 6 glabrous and having ocellate punctures on the upper margin.
|
||
<emphasis id="B91CEAAA4530FF8A0A30FE60FB70FDE2" bold="true" box="[1135,1260,511,536]" pageId="11">Pygidium:</emphasis>
|
||
shinny, lacking punctures, wider than long.
|
||
<emphasis id="B91CEAAA4530FF8A0F80FDBBFDE8FDC7" bold="true" box="[479,628,548,573]" pageId="11">Paramera: (</emphasis>
|
||
as in
|
||
<figureCitation id="13532A3D4530FF8A0CE8FDBBFC9BFDC7" box="[695,775,548,573]" captionStart="FIGURE 6" captionStartId="10.[151,250,1362,1384]" captionTargetBox="[283,1306,458,1329]" captionTargetId="figure@10.[267,1320,445,1340]" captionTargetPageId="10" captionText="FIGURE 6. Paratypes of Dichotomius (Selenocopris) paresi n. sp.: (A) male dorsal habitus; (B) female dorsal habitus (black arrows indicate the ocellate punctures at the posterior portion of the pronotal disc); (C) dorsal view of male paramera; (D) lateral view of male paramera; (E) detail of elytra, white arrow indicating the elytral striae widened near the basis. Scale bar = 5 mm." httpUri="https://zenodo.org/record/272005/files/figure.png" pageId="11">Figs. 6</figureCitation>
|
||
C–D) having rounded/circular excavations near the basis (dorsal view) and longitudinal excavation toward the apex (lateral and dorsal view). Anterior border of circular/ rounded excavation carinates. Longitudinal excavation taking 1/3 the length of paramera. Apex divergent, curved outward (dorsal view).
|
||
</paragraph>
|
||
<paragraph id="8BD736B84530FF8A0E98FD2BFB98FC5E" blockId="11.[151,1437,151,1365]" pageId="11">
|
||
<emphasis id="B91CEAAA4530FF8A0E98FD2BFE68FD37" bold="true" box="[199,500,692,717]" pageId="11">Morphological variation.</emphasis>
|
||
length varies from 17.0–
|
||
<quantity id="4C909B5D4530FF8A0D45FD2BFCE3FD36" box="[794,895,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="11" unit="mm" value="17.5">17.5 mm</quantity>
|
||
on males and from 17.5–18.0 mm on females. Pronotum width varies from
|
||
<quantity id="4C909B5D4530FF8A0F84FD48FDE1FD15" box="[475,637,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.15" metricValueMin="1.05" pageId="11" unit="mm" value="11.0" valueMax="11.5" valueMin="10.5">10.5–11.5 mm</quantity>
|
||
on males and from 10.5–12.0 mm on females. Both males and females show variations in following characters:
|
||
<emphasis id="B91CEAAA4530FF8A0C35FD63FD31FCEF" bold="true" box="[618,685,764,789]" pageId="11">head:</emphasis>
|
||
height of the clypeo-frontal carina.
|
||
<emphasis id="B91CEAAA4530FF8A0A11FD63FB4DFCEF" bold="true" box="[1102,1233,764,789]" pageId="11">Pronotum:</emphasis>
|
||
anterior declivity may vary in size: on most of the females, the declivity is lower. Also on females, ocellate punctures at anterior lobe are more sparse and smaller (3/4 the diameter of the male biggest occellus).
|
||
<emphasis id="B91CEAAA4530FF8A0DA7FCDBFBD1FCA7" bold="true" box="[1016,1101,836,861]" pageId="11">Elytra:</emphasis>
|
||
both males and females have variations on the color of the elytra base that may exhibit brown reflections.
|
||
<emphasis id="B91CEAAA4530FF8A0DABFCF8FBEFFC7A" bold="true" box="[1012,1139,871,896]" pageId="11">Abdomen:</emphasis>
|
||
only on females, the sixth abdominal segment has a pair of central tubercles vertically oriented (
|
||
<figureCitation id="13532A3D4530FF8A0DFBFC13FC7EFC5E" box="[932,994,908,933]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="11">Fig. 1</figureCitation>
|
||
A)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37265334530FF8A0E98FC30FD03FC16" pageId="11" type="distribution">
|
||
<paragraph id="8BD736B84530FF8A0E98FC30FD03FC16" blockId="11.[151,1437,151,1365]" pageId="11">
|
||
<emphasis id="B91CEAAA4530FF8A0E98FC30FEFCFC32" bold="true" box="[199,352,943,968]" pageId="11">Distribution.</emphasis>
|
||
Known from the region of Chapada dos Parecis, municipalities of Nova Marilândia and Tangará da Serra, state of
|
||
<collectingRegion id="49ACF85A4530FF8A0F05FC4AFE72FC16" box="[346,494,980,1004]" country="Brazil" name="Mato Grosso" pageId="11">Mato Grosso</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37F76284530FF8A0FA6FC4BFDA1FC17" box="[505,573,980,1005]" name="Brazil" pageId="11">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="13532A3D4530FF8A0C12FC4BFD0DFC16" box="[589,657,980,1005]" captionStart="FIGURE 8" captionStartId="13.[151,250,1913,1935]" captionTargetBox="[235,1353,676,1876]" captionTargetId="figure@13.[222,1365,661,1892]" captionTargetPageId="13" captionText="FIGURE 8. Distribution of the brachypterous species of Selenocopris." httpUri="https://zenodo.org/record/272007/files/figure.png" pageId="11">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37265334530FF8A0E98FC68FC7AFBA2" pageId="11" type="etymology">
|
||
<paragraph id="8BD736B84530FF8A0E98FC68FC7AFBA2" blockId="11.[151,1437,151,1365]" pageId="11">
|
||
<emphasis id="B91CEAAA4530FF8A0E98FC68FED2FBEA" bold="true" box="[199,334,1015,1040]" pageId="11">Etymology.</emphasis>
|
||
The species is named after the region that covers the type-locality known as Chapada dos Parecis (Parecis plateau, usually spelled with “c”) at the brazilian states of
|
||
<collectingRegion id="49ACF85A4530FF8A0D20FB82FB8CFBCE" box="[895,1040,1052,1076]" country="Brazil" name="Mato Grosso" pageId="11">Mato Grosso</collectingRegion>
|
||
and
|
||
<collectingRegion id="49ACF85A4530FF8A0A19FB83FB2BFBCF" box="[1094,1207,1052,1077]" country="Brazil" name="Rondonia" pageId="11">Rondônia</collectingRegion>
|
||
.
|
||
<taxonomicName id="4C684D3B4530FF8A0A9FFB83FA98FBCF" box="[1216,1284,1052,1077]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="paresi">Paresi</taxonomicName>
|
||
(spelled with “s”) is also the denomination of the ethnic groups living at the same region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37265334530FF8A0E98FBFBFD16FAAF" pageId="11" type="discussion">
|
||
<paragraph id="8BD736B84530FF8A0E98FBFBFD16FAAF" blockId="11.[151,1437,151,1365]" pageId="11">
|
||
<emphasis id="B91CEAAA4530FF8A0E98FBFBFEA7FB87" bold="true" box="[199,315,1124,1149]" pageId="11">Remarks.</emphasis>
|
||
this is the only species of
|
||
<taxonomicName id="4C684D3B4530FF8A0C2CFBFBFC98FB86" box="[627,772,1124,1148]" class="Insecta" family="Scarabaeidae" genus="Selenocopris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91CEAAA4530FF8A0C2CFBFBFC98FB86" box="[627,772,1124,1148]" italics="true" pageId="11">Selenocopris</emphasis>
|
||
</taxonomicName>
|
||
that bears ocellate punctures on the pronotal disc and the unique in the
|
||
<taxonomicName id="4C684D3B4530FF8A0F6EFB17FE5FFB5A" box="[305,451,1160,1184]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="quadraticeps">
|
||
<emphasis id="B91CEAAA4530FF8A0F6EFB17FE5FFB5A" box="[305,451,1160,1184]" italics="true" pageId="11">quadraticeps</emphasis>
|
||
</taxonomicName>
|
||
group that lacks punctures in the elytral striae. Among the brachypterous species in the group, the male paramera is most similar to
|
||
<taxonomicName id="4C684D3B4530FF8A0CC9FB32FC9DFB3E" box="[662,769,1197,1220]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="manni">
|
||
<emphasis id="B91CEAAA4530FF8A0CC9FB32FC9DFB3E" box="[662,769,1197,1220]" italics="true" pageId="11">D. manni</emphasis>
|
||
</taxonomicName>
|
||
while the males head and pronotum characters are shared with
|
||
<taxonomicName id="4C684D3B4530FF8A0E8BFB4EFED8FB1D" box="[212,324,1232,1256]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="ingens">
|
||
<emphasis id="B91CEAAA4530FF8A0E8BFB4EFED8FB1D" box="[212,324,1232,1256]" italics="true" pageId="11">D. ingens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91CEAAA4530FF8A0FDCFB4EFD67FB1D" box="[387,763,1232,1256]" italics="true" pageId="11">
|
||
<taxonomicName id="4C684D3B4530FF8A0FDCFB4EFD90FB12" box="[387,524,1232,1256]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="fissiceps">D. fissiceps</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C684D3B4530FF8A0C43FB4FFD67FB1D" box="[540,763,1232,1256]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="paresi">Dichotomius paresi</taxonomicName>
|
||
</emphasis>
|
||
has been collected at open habitat (cerrado
|
||
<emphasis id="B91CEAAA4530FF8A0B54FB4EFA00FB1D" box="[1291,1436,1232,1256]" italics="true" pageId="11">sensu stricto</emphasis>
|
||
formation) at
|
||
<taxonomicName id="4C684D3B4530FF8A0F6FFB6BFEE8FAF7" box="[304,372,1268,1293]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="paresi">Paresi</taxonomicName>
|
||
Plateau, which takes part of the states of
|
||
<collectingRegion id="49ACF85A4530FF8A0D1DFB6AFC4EFAF6" box="[834,978,1268,1292]" country="Brazil" name="Mato Grosso" pageId="11">Mato Grosso</collectingRegion>
|
||
and
|
||
<collectingRegion id="49ACF85A4530FF8A0A57FB6BFBE5FAF7" box="[1032,1145,1268,1293]" country="Brazil" name="Rondonia" pageId="11">Rondônia</collectingRegion>
|
||
. Ecologically, this species is similar to
|
||
<emphasis id="B91CEAAA4530FF8A0F75FA86FE02FAD5" box="[298,414,1304,1328]" italics="true" pageId="11">
|
||
<taxonomicName id="4C684D3B4530FF8A0F75FA86FE06FAD5" box="[298,410,1304,1328]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="ingens">D. ingens</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
both were collected at open habitats, at the hottest hours of the day (between 11:00 and 13:00) and being mostly attracted to carrion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |