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<document id="92953DAC8618834D3C128E8C61B22E58" ID-DOI="10.5852/ejt.2023.913.2375" ID-ISSN="2118-9773" ID-Zenodo-Dep="10405782" ID-ZooBank="F9BD2A46-57EC-4B9D-A52E-E6A45EC67B56" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1702989613919" checkinUser="plazi" docAuthor="Fernández, Omar Rafael Regalado, Stöhr, Henrik, Kästle, Benedikt &amp; Werneburg, Ingmar" docDate="2023" docId="03C14140FF8543339E483214FEBEFBC4" docLanguage="en" docName="EJT.2023.913.1-88.pdf" docOrigin="European Journal of Taxonomy 913 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2375/10423" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Teratosaurus suevicus Meyer 1861" docType="treatment" docVersion="1" lastPageNumber="62" masterDocId="FFF83938FFB8430E9C0D3041FFE8FF98" masterDocTitle="Diversity and taxonomy of the Late Triassic sauropodomorphs (Saurischia, Sauropodomorpha) stored in the Palaeontological Collection of Tübingen, Germany, historically referred to Plateosaurus" masterLastPageNumber="88" masterPageNumber="1" pageNumber="62" updateTime="1703256379235" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9A835B90017091F9844DE10224331A99">Diversity and taxonomy of the Late Triassic sauropodomorphs (Saurischia, Sauropodomorpha) stored in the Palaeontological Collection of Tübingen, Germany, historically referred to Plateosaurus</mods:title>
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<mods:namePart id="202C3B03CD976FAA7A463ACE80B2B3B8">Fernández, Omar Rafael Regalado</mods:namePart>
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<mods:affiliation id="208AEF6B84F527AE698EE1C4E26684DC">Fachbereich Geowissenschaften an der Universität Tübingen, Hölderlinstrasse 12, 72074 Tübingen, Germany. &amp; Senckenberg Centre for Human Evolution and Palaeoenvironment an der Universität Tübingen, Sigwartstrasse 10, 72076 Tübingen, Germany.</mods:affiliation>
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<mods:namePart id="77358D555C46CECB1CA51828AE91C9F7">Stöhr, Henrik</mods:namePart>
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<mods:affiliation id="1825DCF54FFC1E7C3BA724AB77820056">Fachbereich Geowissenschaften an der Universität Tübingen, Hölderlinstrasse 12, 72074 Tübingen, Germany.</mods:affiliation>
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<mods:namePart id="78349C6AA096E9855F5E0F3A04712548">Kästle, Benedikt</mods:namePart>
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<mods:affiliation id="E71B989027FAE5975C4E3A216D955C2E">Fachbereich Geowissenschaften an der Universität Tübingen, Hölderlinstrasse 12, 72074 Tübingen, Germany.</mods:affiliation>
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<mods:namePart id="3ECA43338413BF0CFE63F34FA5AEDE9E">Werneburg, Ingmar</mods:namePart>
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<mods:affiliation id="A2D0904CBAE06B23BFF288BFC4AA3570">Fachbereich Geowissenschaften an der Universität Tübingen, Hölderlinstrasse 12, 72074 Tübingen, Germany. &amp; Senckenberg Centre for Human Evolution and Palaeoenvironment an der Universität Tübingen, Sigwartstrasse 10, 72076 Tübingen, Germany.</mods:affiliation>
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<mods:date id="FFF7E03CDB63A94B0A8126209CC9FDE6">2023</mods:date>
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<taxonomicName id="4C688BD5FF8543339E483214FC06FDF7" authority="Meyer, 1861" authorityName="Meyer" authorityYear="1861" box="[581,1006,597,623]" class="Reptilia" family="Megalosauridae" genus="Teratosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B91C2C44FF8543339E483214FCBAFDF7" bold="true" box="[581,850,597,623]" italics="true" pageId="61" pageNumber="62">Teratosaurus suevicus</emphasis>
<bibRefCitation id="EFF98DA7FF8543339F543214FC06FDF7" author="Meyer" box="[857,1006,597,623]" pageId="61" pageNumber="62" pagination="253 - 346" refId="ref39578" refString="von Meyer. H. 1861. Reptilien aus dem Stubensandstein des oberen Keupers. Palaeontographica 7: 253 - 346." type="journal article" year="1861">Meyer, 1861</bibRefCitation>
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<taxonomicName id="4C688BD5FF8543339CB032D7FE4DFD37" baseAuthorityName="Meyer" baseAuthorityYear="1861" box="[189,421,662,688]" genus="Zanclodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B91C2C44FF8543339CB032D7FE4DFD37" box="[189,421,662,688]" italics="true" pageId="61" pageNumber="62">Zanclodon suevicus</emphasis>
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<treatmentCitation id="0AC9D647FF8543339DCF32D7FD9CFD28" author="Fraas E." box="[450,628,661,688]" page="18" pageId="61" pageNumber="62" year="1896">
<bibRefCitation id="EFF98DA7FF8543339DCF32D7FD9CFD28" author="Fraas E." box="[450,628,661,688]" pageId="61" pageNumber="62" refId="ref35426" refString="Fraas E. 1896. Die Schwabischen Trias-Saurier nach dem Material der Kgl. Naturalien-Sammlung in Stuttgart zusammengestellt. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart." type="book" year="1896">Fraas 1896: 18</bibRefCitation>
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.
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<heading id="D09F473AFF8543339CB03296FE84FD68" bold="true" box="[189,364,726,753]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="B91C2C44FF8543339CB03296FE84FD68" bold="true" box="[189,364,726,753]" pageId="61" pageNumber="62">Type material</emphasis>
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<emphasis id="B91C2C44FF8543339CE73345FEB4FC86" bold="true" box="[234,348,772,798]" pageId="61" pageNumber="62">
<typeStatus id="54D34EF4FF8543339CE73345FEB4FC86" box="[234,348,772,798]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
</emphasis>
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<paragraph id="8BD7F056FF8543339CB03366FE09FCFB" blockId="61.[189,1398,772,868]" pageId="61" pageNumber="62">
<materialsCitation id="3B00FA0BFF8543339CB03366FE09FCFB" collectionCode="NHMUK" country="Germany" location="Mittlerer Stubensandstein" municipality="Mittelkeuper" pageId="61" pageNumber="62" specimenCode="38646" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F37FB0C6FF8543339CB03366FEBBFCD9" box="[189,339,807,833]" name="Germany" pageId="61" pageNumber="62">GERMANY</collectingCountry>
• right maxilla;
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,
<collectingMunicipality id="6BB36A2CFF8543339F523366FC11FCD9" box="[863,1017,807,833]" pageId="61" pageNumber="62">Mittelkeuper</collectingMunicipality>
(middle Norian, Late Triassic);
<collectionCode id="ED796893FF8543339CB0330BFEC4FCFC" box="[189,300,842,868]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="61" pageNumber="62" type="Museum">NHMUK</collectionCode>
PV OR
<specimenCode id="DBCE582DFF8543339D9D3308FE35FCFB" box="[400,477,841,867]" pageId="61" pageNumber="62">38646</specimenCode>
.
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</paragraph>
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<heading id="D09F473AFF8543339CB033CBFEC7FC3C" bold="true" box="[189,303,906,932]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="B91C2C44FF8543339CB033CBFEC7FC3C" bold="true" box="[189,303,906,932]" pageId="61" pageNumber="62">Remarks</emphasis>
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<paragraph id="8BD7F056FF8543339CB033F9FEBEFBC4" blockId="61.[188,1399,952,1116]" pageId="61" pageNumber="62">
<bibRefCitation id="EFF98DA7FF8543339CB033F9FEB8FC4A" author="Fraas E." box="[189,336,952,978]" pageId="61" pageNumber="62" refId="ref35426" refString="Fraas E. 1896. Die Schwabischen Trias-Saurier nach dem Material der Kgl. Naturalien-Sammlung in Stuttgart zusammengestellt. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart." type="book" year="1896">Fraas (1896)</bibRefCitation>
referred to this maxilla as
<taxonomicName id="4C688BD5FF8543339E8D33F9FC8FFC4A" baseAuthorityName="Meyer" baseAuthorityYear="1861" box="[640,871,952,978]" genus="Zanclodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="suevicus">
<emphasis id="B91C2C44FF8543339E8D33F9FC8FFC4A" box="[640,871,952,978]" italics="true" pageId="61" pageNumber="62">Zanclodon suevicus</emphasis>
</taxonomicName>
(species
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<figureCitation id="1353ECD3FF854333981C33F9FBBDFC4A" box="[1041,1109,952,978]" captionStart="Fig" captionStartId="6.[186,229,1363,1392]" captionTargetBox="[204,1924,549,1324]" captionTargetId="figure-18@6.[186,1940,538,1324]" captionTargetPageId="6" captionText="Fig. 3. Timeline showing changes in the taxonomic placement of the specimens of early-diverging sauropodomorphs considered to be Plateosaurus Meyer, 1837 from 1900 to 2004. This timeline considers only papers discussing taxonomy and papers describing and naming new species. Lines indicate continuity or change in specimen composition of species across revisions. Arrows outside the timeline indicate that the taxa were named before 1900 and/or are still valid after 2004. When more lines depart from one node, part of the specimens included in that species were referred to as different species. Lines converging in one node indicate that the specimens were lumped together into the species in the node. Stars at the end of a line indicate that in the taxonomic revision, the name became either a nomen dubium or a nomen nudum. Lines continuing after this point indicate that the specimens were referred to the new node. This timeline does not include specimens removed from the genus Plateosaurus, namely Efraasia (von Huene, 19071908), Ruehleia Galton, 2001, Camelotia Galton, 1985 and Tuebingosaurus Regalado Fernández &amp; Werneburg, 2022. The green lines correspond to species that have been synonymized withPl. trossingensisFraas, 1913, and the blue lines correspond to those that have been synonymized with Pl. gracilis(von Huene, 1905). The orange-red lines refer to species removed from Sauropodomorpha Huene, 1932 altogether. Changes after 2004 are discussed in the text." figureDoi="http://doi.org/10.5281/zenodo.10405789" httpUri="https://zenodo.org/record/10405789/files/figure.png" pageId="61" pageNumber="62">Fig. 3</figureCitation>
). The morphology of the maxilla has been identified as an early-branching rauisuchian, a group of carnivore archosaurs from the Triassic (
<bibRefCitation id="EFF98DA7FF8543339D2433BCFDF2FB8F" author="Brusatte S. L. &amp; Butler R. J. &amp; Sulej T. &amp; Niedzwiedzki G." box="[297,538,1021,1047]" pageId="61" pageNumber="62" pagination="221 - 230" refId="ref34903" refString="Brusatte S. L., Butler R. J., Sulej T. &amp; Niedzwiedzki G. 2009. The taxonomy and anatomy of rauisuchian archosaurs from the Late Triassic of Germany and Poland. Acta Palaeontologica Polonica 54 (2): 221 - 230. https: // doi. org / 10. 4202 / app. 2008. 0065" type="journal article" year="2009">
Brusatte
<emphasis id="B91C2C44FF8543339D9A33BFFE3AFB8F" box="[407,466,1021,1047]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
2009
</bibRefCitation>
). The genus
<taxonomicName id="4C688BD5FF8543339EBE33BFFCA5FB8F" authorityName="Meyer" authorityYear="1861" box="[691,845,1022,1047]" class="Reptilia" family="Megalosauridae" genus="Teratosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="61" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B91C2C44FF8543339EBE33BFFCA5FB8F" box="[691,845,1022,1047]" italics="true" pageId="61" pageNumber="62">Teratosaurus</emphasis>
</taxonomicName>
is therefore excluded from
<taxonomicName id="4C688BD5FF854333989B33BCFA9BFB8F" authorityName="Huene" authorityYear="1932" box="[1174,1395,1021,1047]" class="Reptilia" kingdom="Animalia" order="Saurischia" pageId="61" pageNumber="66" phylum="Chordata" rank="subOrder" subOrder="Sauropodomorpha">Sauropodomorpha</taxonomicName>
. The species of
<taxonomicName id="4C688BD5FF8543339D663461FDEDFBA1" authorityName="Meyer" authorityYear="1861" box="[363,517,1056,1081]" class="Reptilia" family="Megalosauridae" genus="Teratosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="61" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B91C2C44FF8543339D663461FDEDFBA1" box="[363,517,1056,1081]" italics="true" pageId="61" pageNumber="62">Teratosaurus</emphasis>
</taxonomicName>
have been traditionally considered as part of
<taxonomicName id="4C688BD5FF854333981B345EFAF9FBA1" baseAuthorityName="von Huene" baseAuthorityYear="1905" box="[1046,1297,1055,1081]" class="Reptilia" family="Plateosauridae" genus="Plateosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B91C2C44FF854333981B345EFAF9FBA1" box="[1046,1297,1055,1081]" italics="true" pageId="61" pageNumber="62">Plateosaurus gracilis</emphasis>
</taxonomicName>
(species
<quantity id="4C905DB3FF8543339CB03403FF11FBC4" box="[189,249,1090,1116]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="61" pageNumber="62" unit="in" value="20.0">20 in</quantity>
<figureCitation id="1353ECD3FF8543339D0D3403FEAEFBC4" box="[256,326,1090,1116]" captionStart="Fig" captionStartId="6.[186,229,1363,1392]" captionTargetBox="[204,1924,549,1324]" captionTargetId="figure-18@6.[186,1940,538,1324]" captionTargetPageId="6" captionText="Fig. 3. Timeline showing changes in the taxonomic placement of the specimens of early-diverging sauropodomorphs considered to be Plateosaurus Meyer, 1837 from 1900 to 2004. This timeline considers only papers discussing taxonomy and papers describing and naming new species. Lines indicate continuity or change in specimen composition of species across revisions. Arrows outside the timeline indicate that the taxa were named before 1900 and/or are still valid after 2004. When more lines depart from one node, part of the specimens included in that species were referred to as different species. Lines converging in one node indicate that the specimens were lumped together into the species in the node. Stars at the end of a line indicate that in the taxonomic revision, the name became either a nomen dubium or a nomen nudum. Lines continuing after this point indicate that the specimens were referred to the new node. This timeline does not include specimens removed from the genus Plateosaurus, namely Efraasia (von Huene, 19071908), Ruehleia Galton, 2001, Camelotia Galton, 1985 and Tuebingosaurus Regalado Fernández &amp; Werneburg, 2022. The green lines correspond to species that have been synonymized withPl. trossingensisFraas, 1913, and the blue lines correspond to those that have been synonymized with Pl. gracilis(von Huene, 1905). The orange-red lines refer to species removed from Sauropodomorpha Huene, 1932 altogether. Changes after 2004 are discussed in the text." figureDoi="http://doi.org/10.5281/zenodo.10405789" httpUri="https://zenodo.org/record/10405789/files/figure.png" pageId="61" pageNumber="62">Fig. 3</figureCitation>
).
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</subSubSection>
</treatment>
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