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<document id="77D4DDAC93A222FAAA95854F6882D81D" ID-CLB-Dataset="37404" ID-DOI="10.11646/zootaxa.4205.4.1" ID-GBIF-Dataset="eb5dfbad-e024-4318-b58e-6d9b9e19eac3" ID-ISSN="1175-5326" ID-Zenodo-Dep="208241" ID-ZooBank="BAC8F107-21D6-49FE-BAC7-BF4EE6C3E6A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1482105213028" checkinUser="plazi" docAuthor="Kolesik, Peter &amp; Gagné, Raymond J." docDate="2016" docId="D759878B0E66EE4A5BF6FA012EFDFAD0" docLanguage="en" docName="Zootaxa.4205.4.301-338" docOrigin="Zootaxa 4205 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphenolasioptera vastatrix Skuse 1888, new combination" docType="treatment" docVersion="12" lastPageNumber="327" masterDocId="2B60FFF30E7FEE505B61FFDA2A35FFDF" masterDocTitle="Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae)" masterLastPageNumber="338" masterPageNumber="301" pageNumber="326" updateTime="1698434883341" updateUser="ExternalLinkService">
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<mods:title id="51DBE8EDEA97225475662F449A6AD654">Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae)</mods:title>
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<mods:namePart id="51BE36CAE996F831AB0CCFEC3589E3B7">Kolesik, Peter</mods:namePart>
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<mods:namePart id="EF44F003485F0B60F22C331138624B70">Gagné, Raymond J.</mods:namePart>
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<mods:title id="F6569D7A113B701C8B8C9AE4DE1F2938">Zootaxa</mods:title>
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<treatment id="D759878B0E66EE4A5BF6FA012EFDFAD0" ID-DOI="http://doi.org/10.5281/zenodo.4752574" ID-GBIF-Taxon="125140099" ID-Zenodo-Dep="4752574" LSID="urn:lsid:plazi:treatment:D759878B0E66EE4A5BF6FA012EFDFAD0" httpUri="http://treatment.plazi.org/id/D759878B0E66EE4A5BF6FA012EFDFAD0" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
<subSubSection id="17EA65160E66EE495BF6FA012ACDF9C8" pageId="25" pageNumber="326" type="nomenclature">
<paragraph id="5F4F369D0E66EE495BF6FA01295FFA29" blockId="25.[151,874,1499,1559]" box="[151,874,1499,1526]" pageId="25" pageNumber="326">
<heading id="040781F10E66EE495BF6FA01295FFA29" bold="true" box="[151,874,1499,1526]" fontSize="11" level="1" pageId="25" pageNumber="326" reason="1">
<emphasis id="6D84EA8F0E66EE495BF6FA01295FFA29" bold="true" box="[151,874,1499,1526]" pageId="25" pageNumber="326">
<taxonomicName id="98F04D1E0E66EE495BF6FA0128B3FA29" ID-CoL="8F482" authority="Skuse, 1888" authorityName="Skuse" authorityYear="1888" box="[151,646,1499,1526]" class="Insecta" family="Cecidomyiidae" genus="Sphenolasioptera" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="vastatrix" status="comb. nov.">
<emphasis id="6D84EA8F0E66EE495BF6FA012BEDFA2A" bold="true" box="[151,472,1499,1525]" italics="true" pageId="25" pageNumber="326">Sphenolasioptera vastatrix</emphasis>
(
<bibRefCitation id="3B614B6C0E66EE495A86FA06284BFA29" author="Skuse" box="[487,638,1500,1526]" pageId="25" pageNumber="326" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse, 1888</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="76B757F40E66EE4959F5FA06295FFA29" box="[660,874,1500,1526]" pageId="25" pageNumber="326">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5F4F369D0E66EE495BF6FA242ACDF9C8" blockId="25.[151,874,1499,1559]" box="[151,248,1534,1559]" pageId="25" pageNumber="326">[Fig. 14]</paragraph>
</subSubSection>
<subSubSection id="17EA65160E66EE495BF6F99D282AF983" box="[151,543,1606,1628]" pageId="25" pageNumber="326" type="reference_group">
<paragraph id="5F4F369D0E66EE495BF6F99D282AF983" blockId="25.[151,543,1606,1628]" box="[151,543,1606,1628]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="7FE011B30E66EE495BF6F99D282AF983" box="[151,543,1606,1628]" pageId="25" pageNumber="326">
<taxonomicName id="98F04D1E0E66EE495BF6F99D2829F983" ID-CoL="895N5" authority="Skuse, 1888: 128" authorityName="Skuse" authorityPageNumber="128" authorityYear="1888" box="[151,540,1606,1628]" class="Insecta" family="Cecidomyiidae" genus="Lasioptera" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="vastatrix">
<emphasis id="6D84EA8F0E66EE495BF6F99D2B56F983" box="[151,355,1607,1628]" italics="true" pageId="25" pageNumber="326">Lasioptera vastatrix</emphasis>
<treatmentCitation id="DE51108C0E66EE495A08F99C2829F983" author="Skuse" box="[361,540,1606,1628]" page="128" pageId="25" pageNumber="326" year="1888">
<bibRefCitation id="3B614B6C0E66EE495A08F99C2BDCF983" author="Skuse" box="[361,489,1606,1628]" pageId="25" pageNumber="326" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse, 1888</bibRefCitation>
: 128
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E66EE4A5BF6F9532AC9FEBA" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326" type="materials_examined">
<paragraph id="5F4F369D0E66EE495BF6F9532BD4F91A" blockId="25.[151,1437,1673,2022]" pageId="25" pageNumber="326">
<materialsCitation id="EF983CC00E66EE495BF6F9532E74F97E" ID-GBIF-Occurrence="1416805638" box="[151,1089,1673,1698]" collectionCode="NSW" location="Parkes district" pageId="25" pageNumber="326" specimenCount="1" typeStatus="syntype">
<emphasis id="6D84EA8F0E66EE495BF6F9532B6BF97D" bold="true" box="[151,350,1673,1698]" pageId="25" pageNumber="326">Material studied</emphasis>
.
<typeStatus id="804B883F0E66EE495A0FF9532BECF97E" box="[366,473,1673,1697]" pageId="25" pageNumber="326" type="syntype">Syntypes</typeStatus>
: males, females, pupa, larvae, “
<location id="5A2F60460E66EE495821F95329D1F97D" LSID="urn:lsid:plazi:treatment:D759878B0E66EE4A5BF6FA012EFDFAD0:5A2F60460E66EE495821F95329D1F97D" box="[832,996,1673,1698]" name="Parkes district" pageId="25" pageNumber="326">Parkes district</location>
[
<collectionCode id="39E1AE580E66EE495892F9532E0FF97E" box="[1011,1082,1673,1697]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="25" pageNumber="326">NSW</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="EF983CC00E66EE495F2DF9532BE8F91A" ID-GBIF-Occurrence="1416805629" collectingDate="5 th December" collectionCode="ANIC" location="Imagines" pageId="25" pageNumber="326" specimenCount="1" typeStatus="syntype">
<location id="5A2F60460E66EE495F2DF9532E86F97D" LSID="urn:lsid:plazi:treatment:D759878B0E66EE4A5BF6FA012EFDFAD0:5A2F60460E66EE495F2DF9532E86F97D" box="[1100,1203,1673,1698]" name="Imagines" pageId="25" pageNumber="326">Imagines</location>
began to emerge on the
<date id="2B4E105D0E66EE495BA1F9762B56F91A" box="[192,355,1708,1733]" pageId="25" pageNumber="326">
<collectingDate id="3B0AE9B50E66EE495BA1F9762B56F91A" box="[192,355,1708,1733]" pageId="25" pageNumber="326">5th December</collectingDate>
</date>
”, in
<collectionCode id="39E1AE580E66EE495AF4F9772BE8F91A" box="[405,477,1709,1733]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="25" pageNumber="326">ANIC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="5F4F369D0E66EE495BA6F90B2BF6F842" blockId="25.[151,1437,1673,2022]" pageId="25" pageNumber="326">
<emphasis id="6D84EA8F0E66EE495BA6F90B2B4BF935" bold="true" box="[199,382,1745,1770]" pageId="25" pageNumber="326">Associated gall</emphasis>
. The
<typeStatus id="804B883F0E66EE495AA6F9082837F935" box="[455,514,1746,1770]" pageId="25" pageNumber="326">types</typeStatus>
were reared from “saffron-yellow” larvae gregariously living within indistinct swellings on stems of an unidentified grass: ”Inhabiting grass-stems, generally that portion underneath the spathe. The deformation caused is a scarcely perceptible swelling, extending from an inch to an inch and a-half in length, and containing from ten to a dozen larvae, lying somewhat obliquely, enveloped in delicate white filmy cocoons. These larvae have as yet only been found in a species of grass in the Parkes district, where they prove very destructive to the pasture.”
</paragraph>
<paragraph id="5F4F369D0E66EE4A5BA6F8702AC9FEBA" blockId="25.[151,1437,1673,2022]" lastBlockId="26.[151,1437,151,1296]" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
We mounted one male, designated here as
<emphasis id="6D84EA8F0E66EE4959CCF873292DF81D" bold="true" box="[685,792,1961,1986]" pageId="25" pageNumber="326">lectotype</emphasis>
(ANIC 29-38509), three females (
<emphasis id="6D84EA8F0E66EE495FC2F8732F7DF81D" bold="true" box="[1187,1352,1961,1986]" pageId="25" pageNumber="326">paralectoypes</emphasis>
, ANIC 29-38508,10,11) and four larvae (
<emphasis id="6D84EA8F0E66EE495973F8172882F839" bold="true" box="[530,695,1997,2022]" pageId="25" pageNumber="326">paralectoypes</emphasis>
, ANIC 29-385125) while one pin bearing a card with two wings and five legs was left intact for future DNA study. No pupa was found in ANIC. Surviving body parts of the mounted types: lectotype male: head without flagellomeres, end of abdomen with terminalia, thorax with one leg; female 1: wings, head with antennae, thorax with legs, abdomen with terminalia; female 2: wings, head with antennae, thorax with two legs, abdomen with terminalia; female 3: wings, damaged but present head with antennae, thorax with three legs, abdomen with terminalia; four whole larvae, except one missing the terminal segment.
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E65EE4A5BA6FEB52EFDFAD0" pageId="26" pageNumber="327" type="description">
<paragraph id="5F4F369D0E65EE4A5BA6FEB52BF2FC80" blockId="26.[151,1437,151,1296]" pageId="26" pageNumber="327">
<emphasis id="6D84EA8F0E65EE4A5BA6FEB52B6CFE57" bold="true" box="[199,345,367,392]" pageId="26" pageNumber="327">Description.</emphasis>
<emphasis id="6D84EA8F0E65EE4A5A05FEAA2B95FE57" box="[356,416,368,392]" italics="true" pageId="26" pageNumber="327">Adult</emphasis>
. Colour (extracted from
<bibRefCitation id="3B614B6C0E65EE4A59A9FEB5295BFE57" author="Skuse" box="[712,878,367,392]" pageId="26" pageNumber="327" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse (1888))</bibRefCitation>
: Antennae black to dark brown with short pale pubescence; palpus yellowish brown; thorax deep brown covered with golden-yellow scales and pubescence; halteres with pale brown base, white scales on stalk, club brown, darker in female than male, with some scattered dark brown or black scales; abdomen covered anteriorly on each tergite with deep brown scales, bordered posteriorly with band of white scales in male and yellowish in female; pale brown between segments, sternites pale brown, covered with white scales; male terminalia densely covered with white scales and long hairs; female terminalia pale brownish; legs densely covered with scales appearing deep brown with pale reflexions when viewed at angle; wings yellowish brown at base, hyaline with weak cupreous reflexion, deeply ciliated on posterior border, costal vein deep brown, thickly scaled with yellowish white spot at junction with R4+5 vein, R1 and R4+5 veins pale brown. Wing in both sexes
<quantity id="98089B780E65EE4A5924FD6D289CFD10" box="[581,681,695,719]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="26" pageNumber="327" unit="mm" value="2.27">2.27 mm</quantity>
long,
<quantity id="98089B780E65EE4A5990FD6D2960FD10" box="[753,853,695,719]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="26" pageNumber="327" unit="mm" value="0.88">0.88 mm</quantity>
wide (
<bibRefCitation id="3B614B6C0E65EE4A58C3FD6C2E1FFD10" author="Skuse" box="[930,1066,694,719]" pageId="26" pageNumber="327" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse 1888</bibRefCitation>
).
<bibRefCitation id="3B614B6C0E65EE4A5F5FFD6C2EE6FD10" author="Skuse" box="[1086,1235,694,719]" pageId="26" pageNumber="327" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse (1888)</bibRefCitation>
gives the number of flagellomeres as
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male and
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female. While the male
<typeStatus id="804B883F0E65EE4A5815FD0029EEFD2C" box="[884,987,730,755]" pageId="26" pageNumber="327" type="lectotype">lectotype</typeStatus>
lacks antennae, all
<specimenCount id="49F6FD140E65EE4A5FDBFD012F13FD2C" box="[1210,1318,730,755]" pageId="26" pageNumber="327" type="female">3 females</specimenCount>
have both antennae with 15 flagellomeres, agreeing with
<bibRefCitation id="3B614B6C0E65EE4A59C8FD242975FCC8" author="Skuse" box="[681,832,766,791]" pageId="26" pageNumber="327" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse (1888)</bibRefCitation>
, so we consider the number of flagellomeres as
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both sexes. First palpal segment wider and shorter than second and third, scape 1.5x wider and longer than pedicel, eye bridge five facets long.
</paragraph>
<paragraph id="5F4F369D0E65EE4A5BA6FCB02FBFFBCC" blockId="26.[151,1437,151,1296]" pageId="26" pageNumber="327">Male terminalia: gonocoxites cylindrical beyond base; gonocoxal apodemes joined basally; mediobasal lobe subdivided, dorsal part of lobe short, hemispherical, long-microtrichose, ventral part long-tapered, closely sheathing one side of aedeagus, nearly as long as aedeagus, mostly microtrichose but glabrous apically, tipped with short setae; aedeagus as long as gonocoxites, tapered apically; cerci triangular, tapering; hypoproct bilobed, lobes round, incision between lobes shallow; gonostylus with basal third rounded, microtrichose, narrow and carinate beyond.</paragraph>
<paragraph id="5F4F369D0E65EE4A5BA6FBC42F77FB84" blockId="26.[151,1437,151,1296]" pageId="26" pageNumber="327">Ovipositor: fused, bilaterally flattened cerci with scooped setae along dorsal ridge, 2025 of them along most of ridge, followed by 1520 additional, smaller ones subapically, with sparse short simple setae elsewhere.</paragraph>
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<emphasis id="6D84EA8F0E65EE4A5BA6FBB22B37FBA0" box="[199,258,1128,1151]" italics="true" pageId="26" pageNumber="327">Pupa</emphasis>
.
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in length, brownish-yellow in colour except thorax which is paler (
<bibRefCitation id="3B614B6C0E65EE4A5F07FBBC2ED8FBA0" author="Skuse" box="[1126,1261,1126,1151]" pageId="26" pageNumber="327" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse 1888</bibRefCitation>
).
</paragraph>
<paragraph id="5F4F369D0E65EE4A5BA6FB512EFDFAD0" blockId="26.[151,1437,151,1296]" pageId="26" pageNumber="327">
<emphasis id="6D84EA8F0E65EE4A5BA6FB512B27FB7D" box="[199,274,1163,1186]" italics="true" pageId="26" pageNumber="327">Larva.</emphasis>
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in length, saffron-yellow (
<bibRefCitation id="3B614B6C0E65EE4A59D1FB50290CFB7C" author="Skuse" box="[688,825,1162,1187]" pageId="26" pageNumber="327" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse 1888</bibRefCitation>
); head capsule hemispherical to slightly triangular, as long as posterolateral apodemes, antennae slightly longer than wide at base; spatula with 4 pointed lobes, central two slightly longer than lateral; 4 lateral papillae on each side of spatula, 2 with tiny setae, 2 without; ventral papilla without seta, closely adjacent to lateral papillae. Terminal segment with 6 short papillae.
</paragraph>
</subSubSection>
</treatment>
</document>