217 lines
20 KiB
XML
217 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.443.7880" ID-GBIF-Dataset="33ef48a3-7145-4103-9159-e11be2ba9adc" ID-PMC="PMC4205504" ID-Pensoft-Pub="1313-2970-443-61" ID-PubMed="25349497" ID-ZBK="E33841396A6E426C840D85BC32A12E78" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-443-61" ModsDocOrigin="ZooKeys 443" ModsDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" checkinTime="1451245273153" checkinUser="pensoft" docAuthor="Sokolov, Igor M. & Kavanaugh, David H." docDate="2014" docId="D731F77B785ED6564010A82BC8A0E89E" docLanguage="en" docName="ZooKeys 443: 61-118" docOrigin="ZooKeys 443" docSource="http://dx.doi.org/10.3897/zookeys.443.7880" docTitle="Geocharidius comayaguanus Sokolov & Kavanaugh, 2014, sp. n." docType="treatment" docUuid="CC2857E4-7428-4FC0-A2BC-CFA5A571E8C3" docUuidSource="ZooBank" docVersion="4" lastPageNumber="82" masterDocId="3546FFA8CF78F064801706227F76A615" masterDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" masterLastPageNumber="118" masterPageNumber="61" pageNumber="82" updateTime="1668159453511" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>443</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>61</mods:start>
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<mods:end>118</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.443.7880</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.443.7880</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-443-61</mods:identifier>
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<mods:identifier type="ZBK">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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<mods:identifier type="ZooBank">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152055688" LSID="urn:lsid:zoobank.org:act:CC2857E4-7428-4FC0-A2BC-CFA5A571E8C3" httpUri="http://treatment.plazi.org/id/D731F77B785ED6564010A82BC8A0E89E" lastPageNumber="82" pageId="21" pageNumber="82">
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<subSubSection pageId="21" pageNumber="82" type="multiple">
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<paragraph pageId="21" pageNumber="82">
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<pageBreakToken pageId="21" pageNumber="82" start="start">Taxon</pageBreakToken>
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classification Animalia Coleoptera Carabidae
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="nomenclature">
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<paragraph pageId="21" pageNumber="82">
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<taxonomicName LSID="http://zoobank.org/CC2857E4-7428-4FC0-A2BC-CFA5A571E8C3" class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius comayaguanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="comayaguanus">Geocharidius comayaguanus</taxonomicName>
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<taxonomicNameLabel pageId="21" pageNumber="82">sp. n.</taxonomicNameLabel>
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Figs 2C, 3C, 3F, 6H, 7D, H, 18C, 19
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<normalizedToken originalValue="K–P">K-P</normalizedToken>
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, 20D, 21D, 22, 23
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="type material">
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<paragraph pageId="21" pageNumber="82">Type material.</paragraph>
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<paragraph pageId="21" pageNumber="82">
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HOLOTYPE, a male, in CMNC, point-mounted, dissected, labeled: \ HONDURAS: Comayagua, 18km ENE Comayagua, 1950m, 20.VIII.1994, S. & J. Peck, wet oak-pine forest litter, S&JPeck 1994-52 \ CMNC \ HOLOTYPE
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius comayaguanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="comayaguanus">Geocharidius comayaguanus</taxonomicName>
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Sokolov and Kavanaugh 2014 [red label] \. PARATYPES: A total of 23 specimens (6 males and 2 females were dissected), deposited in CAS, CMNC and KUNHM; 4 specimens labeled same as holotype; 5 specimens labeled: \ HONDURAS: Comayagua, Comayagua (18km E.N.E.), 1950m, 20.VIII.1994, S. Peck wet oak-pine forest litter, SBP 94-52 \ CMNC \; 8 specimens labeled: HONDURAS: La Paz Dept. Tutule, Res. Biol. Guajiquiro,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="14.166667">14°10'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-87.833336">87°50'W</geoCoordinate>
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, 2130m, 7-V-2002, R.Anderson, cloud forest litter, RSA2002-010 \ SM0
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<normalizedToken originalValue="…">...</normalizedToken>
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KUNHM-ENT \; 1 specimen labeled: HONDURAS: LA PAZ: Tutule, Res. Biol. Guajiquiro,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="14.166667">14°10'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-87.833336">87°50'W</geoCoordinate>
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, 2130m, 7.V.2002, R.Anderson, cloud forest litter, 2002-010H \ CMNC \; 1 specimen labeled: \ HONDURAS: LA PAZ: Tutule, Res. Biol. Guajiquiro,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="14.166667">14°10'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-87.833336">87°50'W</geoCoordinate>
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, 2130m, 7.V.2002, R.Anderson, cloud forest litter, 2002-010D \ CMNC \; 1 specimen labeled: \ HONDURAS: LA PAZ: Tutule, Res. Biol. Guajiquiro,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="14.166667">14°10'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-87.833336">87°50'W</geoCoordinate>
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, 2130m, 7.V.2002, R.Anderson, cloud forest litter, 2002-010E \ CMNC \; 2 specimens labeled: \ HONDURAS: LA PAZ: Tutule, Res. Biol. Guajiquiro, 14°10', N
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-87.833336">87°50'W</geoCoordinate>
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, 2130m, 7.V.2002, R.Anderson, cloud forest litter, 2002-010I \ CMNC \; 1 specimen labeled: \ HONDURAS: Yoro Dept., P.N. Pico Pijol, 1300m, 15°09.4', N
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-87.62666">87°37.6'W</geoCoordinate>
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, 11.V.2002, R. Anderson, upper montane forest litter, 2002-017A\ CMNC.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="type locality">
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<paragraph pageId="21" pageNumber="82">Type locality.</paragraph>
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<paragraph pageId="21" pageNumber="82">Honduras, Comayagua Department, 18 km ENE of Comayagua.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="etymology">
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<paragraph pageId="21" pageNumber="82">Etymology.</paragraph>
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<paragraph pageId="21" pageNumber="82">The specific epithet is a Latinized adjective in the masculine form based on the name of the city of Comayagua, from the vicinity of which the new species is described.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="recognition">
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<paragraph pageId="21" pageNumber="82">Recognition.</paragraph>
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<paragraph pageId="21" pageNumber="82">
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Adults of this species are practically indistinguishable externally from those of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius disjunctus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="disjunctus">Geocharidius disjunctus</taxonomicName>
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, described below, and are distinguished from the latter, as from those of the other members of the integripennis species group, by the structure of the male median lobe and the shape of spermatheca in females.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="description">
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<paragraph pageId="21" pageNumber="82">Description.</paragraph>
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<paragraph pageId="21" pageNumber="82">
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Size. Small to medium for genus (SBL range 1.19-1.34 mm, mean 1.28
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.072mm, n=20).
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</paragraph>
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<paragraph pageId="21" pageNumber="82">
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Habitus. Body form (Fig. 18C) moderately convex, ovoid, general proportions (WE/SBL 0.40
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.011), proportions of head (WH/WPm 0.74
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.017) and pronotum (WPm/WE 0.78
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.018) moderately wide.
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</paragraph>
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<paragraph pageId="21" pageNumber="82">Color. Body brunneorufous, appendages testaceous.</paragraph>
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<paragraph pageId="21" pageNumber="82">Microsculpture. Mesh pattern of irregularly isodiametric sculpticells present over all dorsal surfaces of head and elytra. Pronotum and proepisternum smooth (without evident microsculpture).</paragraph>
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<paragraph pageId="21" pageNumber="82">
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Prothorax. Pronotum moderately transverse (WPm/LP 1.29
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.024), with lateral margins markedly constricted posteriorly (WPm/WPp 1.35
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.027). Posterior angles obtuse (110-120°). Width between posterior angles equal to the width between anterior angles (WPa/WPp 1.02
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.026). Ventral aspect (Fig. 3C).
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</paragraph>
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<paragraph pageId="21" pageNumber="82">Pterothorax (Fig. 3F).</paragraph>
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<paragraph pageId="21" pageNumber="82">
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Elytra (Fig. 2C). Moderately convex, slightly depressed along suture, moderately wide (WE/LE 0.68
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.022), without traces of striae. Humeri rounded, in outline forming right angle with longitudinal axis of body. Lateral margins convex, evenly divergent at basal third, evenly rounded to apex in apical third.
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</paragraph>
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<paragraph pageId="21" pageNumber="82">Legs. Protibia (Fig. 6H). Mesotibia (Fig. 7D). Metatibia (Fig. 7H).</paragraph>
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<paragraph pageId="21" pageNumber="82">
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Male genitalia. Median lobe (Fig. 19K) with shaft long and dorsally convex, apex small and narrowly rounded. Ventral margin straight. Dorsal sclerites of internal sac small, in form of a short hook-like fig, slightly varied among different populations (Fig. 19
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<normalizedToken originalValue="K–L">K-L</normalizedToken>
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,
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<normalizedToken originalValue="O–P">O-P</normalizedToken>
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). Right paramere with long apical constriction (Fig. 19N). Left paramere with long and narrow apical constriction (Fig. 19M). Ring sclerite with handle triangular, pointed at apex (Fig. 20D).
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</paragraph>
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<paragraph pageId="21" pageNumber="82">Female internal genitalia. Spermatheca sclerotized, fusiform, slightly tapered apically, straight, with cornu and nodulus of equal length (Fig. 21D). Length of spermathecal gland less than length of spermatheca. Spermathecal duct not coiled.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="geographical distribution">
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<paragraph pageId="21" pageNumber="82">Geographical distribution.</paragraph>
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<paragraph pageId="21" pageNumber="82">This species is known from La Paz, Comayagua and Yoro Departments, thus having a range that crosses nearly the entire Honduran Interior Highlands from the Pacific to the Antlantic slope (Fig. 22, yellow circles).</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="way of life">
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<paragraph pageId="21" pageNumber="82">Way of life.</paragraph>
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<paragraph pageId="21" pageNumber="82">Specimens were collected in litter samples from cloud, upper montane and wet oak-pine forests at middle and high elevations of 1300 and 2130 m, respectively.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="82" type="relationships">
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<paragraph pageId="21" pageNumber="82">Relationships.</paragraph>
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<paragraph pageId="21" pageNumber="82">
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This species is unique within the integripennis species group in the shape of dorsal sclerites of the internal sac (Fig. 19K) of males and of the spermatheca (Fig. 21D) of females. Hence,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius comayaguanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="comayaguanus">Geocharidius comayaguanus</taxonomicName>
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appears to be only remotely related to the other members of the species group.
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</paragraph>
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<caption pageId="21" pageNumber="82">
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<paragraph pageId="21" pageNumber="82">
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Figures 19. Line drawings of aedeagus of Guatemalan and Honduran
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="genus">Geocharidius</taxonomicName>
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species.
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius antigua" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="antigua">Geocharidius antigua</taxonomicName>
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(GUATEMALA,
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<normalizedToken originalValue="Sacatepéquez">Sacatepequez</normalizedToken>
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, Antigua), holotype: A median lobe with internal sac and dorsal sclerites, right lateral aspect B dorsal sclerite of median lobe, dorsal aspect C left paramere, left lateral aspect D right paramere, right lateral aspect
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<normalizedToken originalValue="E–G">E-G</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius celaquensis" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="celaquensis">Geocharidius celaquensis</taxonomicName>
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(HONDURAS, Lempira, Celaque National Park), holotype: E median lobe with internal sac and dorsal sclerites, right lateral aspect F left paramere, left lateral aspect G right paramere, right lateral aspect
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<normalizedToken originalValue="H–J">H-J</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius lencanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="lencanus">Geocharidius lencanus</taxonomicName>
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(HONDURAS, Lempira, Celaque National Park), paratype: H median lobe with internal sac and dorsal sclerites, right lateral aspect I left paramere, left lateral aspect J right paramere, right lateral aspect
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<normalizedToken originalValue="K–N">K-N</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius comayaguanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="comayaguanus">Geocharidius comayaguanus</taxonomicName>
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(HONDURAS, Comayagua, Comayagua), paratype: K median lobe with internal sac and dorsal sclerites, right lateral aspect L variation in a shape of dorsal sclerite of internal sac, right lateral aspect M left paramere, left lateral aspect N right paramere, right lateral aspect
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<normalizedToken originalValue="O–P">O-P</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius comayaguanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="comayaguanus">Geocharidius comayaguanus</taxonomicName>
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(HONDURAS, La Paz, Guajicuiro): variations in a shape of dorsal sclerite of internal sac, right lateral aspect
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<normalizedToken originalValue="Q–S">Q-S</normalizedToken>
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius disjunctus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="disjunctus">Geocharidius disjunctus</taxonomicName>
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(HONDURAS, Francisco
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<normalizedToken originalValue="Morazán">Morazan</normalizedToken>
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, La Tigra National Park), holotype: Q median lobe with internal sac and dorsal sclerites, right lateral aspect R left paramere, left lateral aspect S right paramere, right lateral aspect. T
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius disjunctus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="disjunctus">Geocharidius disjunctus</taxonomicName>
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(HONDURAS, Yoro, Pico Pijol National Park): shape of dorsal sclerite of median lobe, right lateral aspect. Scale = 0.05mm.
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</paragraph>
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</caption>
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<caption pageId="21" pageNumber="82">
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<paragraph pageId="21" pageNumber="82">
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Figures 20. Line drawings of ring sclerite of Guatemalan and Honduran
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="genus">Geocharidius</taxonomicName>
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species, male genitalia, dorsal aspect. A
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius antigua" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="antigua">Geocharidius antigua</taxonomicName>
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(GUATEMALA,
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<normalizedToken originalValue="Sacatepéquez">Sacatepequez</normalizedToken>
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, Antigua), holotype B
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius celaquensis" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="celaquensis">Geocharidius celaquensis</taxonomicName>
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(HONDURAS, Lempira, Celaque National Park), holotype C
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius lencanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="lencanus">Geocharidius lencanus</taxonomicName>
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(HONDURAS, Lempira, Celaque National Park), paratype D
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius comayaguanus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="comayaguanus">Geocharidius comayaguanus</taxonomicName>
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(HONDURAS, Comayagua, Comayagua), paratype E
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius disjunctus" order="Coleoptera" pageId="21" pageNumber="82" phylum="Arthropoda" rank="species" species="disjunctus">Geocharidius disjunctus</taxonomicName>
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(HONDURAS, Francisco
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<normalizedToken originalValue="Morazán">Morazan</normalizedToken>
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, La Tigra National Park), holotype. Scale = 0.1mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |