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<mods:title id="17CB3E958B98422E2006E33BE6AA5B46">A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<taxonomicName id="7DFCF58EFF9EFFA4FF3E75962D47D962" authorityName="Brenske" authorityYear="1897" box="[151,304,498,524]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="8888521FFF9EFFA4FF3E75962D47D962" bold="true" box="[151,304,498,524]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
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(sbg.) 
<taxonomicName id="7DFCF58EFF9EFFA4FE2D75962E78D963" authority="Ahrens &amp; Liu &amp; Lukic &amp; Bai, 2023" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" box="[388,527,498,525]" kingdom="Animalia" pageId="8" pageNumber="9" rank="subGenus" status="subgen. nov." subGenus="Helioserica">
<emphasis id="8888521FFF9EFFA4FE2D75962E78D963" bold="true" box="[388,527,498,525]" italics="true" pageId="8" pageNumber="9">Helioserica</emphasis>
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Ahrens, Lukic &amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9EFFA4FC8275962FAED962" box="[811,985,498,524]" pageId="8" pageNumber="9" rank="subGenus">new subgenus</taxonomicNameLabel>
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<paragraph id="BA438E0DFF9EFFA4FF3E765E28B2D93A" blockId="8.[151,1221,570,597]" box="[151,1221,570,597]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FF3E765E2E5BD93A" bold="true" box="[151,556,570,596]" pageId="8" pageNumber="9">
<typeStatus id="654730AFFF9EFFA4FF3E765E2CA5D93A" box="[151,210,570,596]" pageId="8" pageNumber="9">Type</typeStatus>
species of the new subgenus:
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<taxonomicName id="7DFCF58EFF9EFFA4FD9A765F2FDED93B" authority="Moser, 1915" authorityName="Moser" authorityYear="1915" box="[563,937,570,597]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="8888521FFF9EFFA4FD9A765F2F61D93A" box="[563,790,571,596]" italics="true" pageId="8" pageNumber="9">Microserica varians</emphasis>
<bibRefCitation id="DE6DF3FCFF9EFFA4FCB4765F2FDED93B" author="Moser, J." box="[797,937,570,597]" pageId="8" pageNumber="9" pagination="337 - 393" refId="ref97123" refString="Moser, J. (1915) Neue Serica - Arten. Deutsche Entomologische Zeitschrift, 1915, 337 - 393. https: // doi. org / 10.1002 / mmnd. 48019150402" type="journal article" year="1915">Moser, 1915</bibRefCitation>
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(by current designation).
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<paragraph id="BA438E0DFF9EFFA4FF3E76E62FD1D866" blockId="8.[151,1437,642,1101]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FF3E76E62D65D9F2" bold="true" box="[151,274,642,668]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
The new subgenus differs from its sister clade (
<figureCitation id="22C79288FF9EFFA4FC8F76E62F10D9F2" box="[806,871,642,668]" captionStart="FIGURE 1" captionStartId="84.[152,255,1943,1967]" captionTargetBox="[236,1352,186,1917]" captionTargetId="figure-17@84.[225,1362,181,1919]" captionTargetPageId="84" captionText="FIGURE 1. Phylogenetic placement of lineages so far assigned to Microserica (and some of its closest relatives) according to the phylogeny of Eberle et al. (2017)." figureDoi="http://doi.org/10.5281/zenodo.7639738" httpUri="https://zenodo.org/record/7639738/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
), 
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<emphasis id="8888521FFF9EFFA4FCD676E72867D9F3" box="[895,1040,643,669]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
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, by the well-produced and distinctly angled anterior angles of pronotum, as well as by the distinctly emarginated anterior margin of labroclypeus. Furthermore, the apical abdomen is not as robustly developed as in most 
<taxonomicName id="7DFCF58EFF9EFFA4FBDF76AF2970D98B" authorityName="Brenske" authorityYear="1897" box="[1142,1287,715,741]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8888521FFF9EFFA4FBDF76AF2970D98B" box="[1142,1287,715,741]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
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species, with the pygidium barely protruding under the apex of elytra.
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<emphasis id="8888521FFF9EFFA4FF6E77762D4CD842" bold="true" box="[199,315,786,812]" pageId="8" pageNumber="9">Remarks.</emphasis>
Given, of being the sister lineage of 
<taxonomicName id="7DFCF58EFF9EFFA4FD5877772FF5D843" authorityName="Brenske" authorityYear="1897" box="[753,898,787,813]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="8888521FFF9EFFA4FD5877772FF5D843" box="[753,898,787,813]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
</taxonomicName>
(
<figureCitation id="22C79288FF9EFFA4FC3C77762FACD842" box="[917,987,786,812]" captionStart="FIGURE 1" captionStartId="84.[152,255,1943,1967]" captionTargetBox="[236,1352,186,1917]" captionTargetId="figure-17@84.[225,1362,181,1919]" captionTargetPageId="84" captionText="FIGURE 1. Phylogenetic placement of lineages so far assigned to Microserica (and some of its closest relatives) according to the phylogeny of Eberle et al. (2017)." figureDoi="http://doi.org/10.5281/zenodo.7639738" httpUri="https://zenodo.org/record/7639738/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
; see also 
<bibRefCitation id="DE6DF3FCFF9EFFA4FBF877762952D842" author="Eberle, J. &amp; Fabrizi, S. &amp; Lago P. &amp; Ahrens, D." box="[1105,1317,786,812]" pageId="8" pageNumber="9" pagination="183 - 197" refId="ref95839" refString="Eberle, J., Fabrizi, S., Lago P. &amp; Ahrens, D. (2017) A historical biogeography of megadiverse Sericini - another story &quot; out of Africa &quot;? Cladistics, 33, 183 - 197. https: // doi. org / 10.1111 / cla. 12162" type="journal article" year="2017">
Eberle 
<emphasis id="8888521FFF9EFFA4FB0D777728A8D842" box="[1188,1247,786,812]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2017
</bibRefCitation>
), and thus representing a monophyletic group with the species of 
<taxonomicName id="7DFCF58EFF9EFFA4FD5677532FE7D83F" authorityName="Brenske" authorityYear="1897" box="[767,912,823,849]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8888521FFF9EFFA4FD5677532FE7D83F" box="[767,912,823,849]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93BBEF64FF9EFFA4FC3E77532851D83E" box="[919,1062,822,849]" pageId="8" pageNumber="9" sensu="stricto">sensu stricto</taxonomicNameLabel>
, we nevertheless decided to erect a new and separate subgenus for these species, since key diagnostic features of 
<taxonomicName id="7DFCF58EFF9EFFA4FB89773F28C6D81B" authorityName="Brenske" authorityYear="1897" box="[1056,1201,859,885]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8888521FFF9EFFA4FB89773F28C6D81B" box="[1056,1201,859,885]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93BBEF64FF9EFFA4FB10773F2931D81A" box="[1209,1350,858,885]" pageId="8" pageNumber="9" sensu="stricto">sensu stricto</taxonomicNameLabel>
are not present among these species, namely the obsolete anterior angles of pronotum and the straight, non-emarginate anterior margin of labroclypeus.
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF6E77A22DAEDF6A" blockId="8.[151,1437,642,1101]" pageId="8" pageNumber="9">
All species so far classified as 
<taxonomicName id="7DFCF58EFF9EFFA4FDBE77A32FB2D88E" authority="(Ahrens &amp; Bezdek 2016)" baseAuthorityName="Ahrens &amp; Bezdek" baseAuthorityYear="2016" box="[535,965,966,993]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="8888521FFF9EFFA4FDBE77A32EDFD88F" box="[535,680,967,993]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
(
<bibRefCitation id="DE6DF3FCFF9EFFA4FD1F77A22FCAD88E" author="Ahrens, D. &amp; Bezdek, A." box="[694,957,966,992]" pageId="8" pageNumber="9" refId="ref94531" refString="Ahrens, D. &amp; Bezdek, A. (2016) Sericini. In: Lobl, I. &amp; Lobl, D (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. Brill, Leiden, pp. 281 - 317." type="book" year="2016">Ahrens &amp; Bezděk 2016</bibRefCitation>
)
</taxonomicName>
are subsequently assigned to the subgenus 
<taxonomicName id="7DFCF58EFF9EFFA4FF3E778F2DA3DF6A" authority="Brenske, 1897" authorityName="Brenske" authorityYear="1897" box="[151,468,1002,1029]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="8888521FFF9EFFA4FF3E778F2D5FDF6B" box="[151,296,1003,1029]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
<bibRefCitation id="DE6DF3FCFF9EFFA4FE86778E2DA3DF6A" author="Brenske, E." box="[303,468,1002,1028]" pageId="8" pageNumber="9" pagination="345 - 468" refId="ref95715" refString="Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310" type="journal article" year="1897">Brenske, 1897</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F2E6DD86FF9EFFA4FF6E706A292CDF22" pageId="8" pageNumber="9" type="etymology">
<paragraph id="BA438E0DFF9EFFA4FF6E706A292CDF22" blockId="8.[151,1437,642,1101]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FF6E706A2D39DF46" bold="true" box="[199,334,1038,1064]" pageId="8" pageNumber="9">Etymology.</emphasis>
The name of the new subgenus is composed of the Greek word “ 
<emphasis id="8888521FFF9EFFA4FB90706A280BDF46" box="[1081,1148,1038,1064]" italics="true" pageId="8" pageNumber="9">helios</emphasis>
” (sun) and the word root 
<taxonomicName id="7DFCF58EFF9EFFA4FF3E70572CA8DF23" authorityName="MacLeay" authorityYear="1819" box="[151,223,1075,1101]" class="Insecta" family="Melolonthidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="8888521FFF9EFFA4FF3E70572CA8DF23" box="[151,223,1075,1101]" italics="true" pageId="8" pageNumber="9">Serica</emphasis>
</taxonomicName>
(
<typeStatus id="654730AFFF9EFFA4FF4470562D68DF22" box="[237,287,1074,1100]" pageId="8" pageNumber="9">type</typeStatus>
genus of the tribe 
<taxonomicName id="7DFCF58EFF9EFFA4FE5970562E3BDF22" authorityName="Kirby" authorityYear="1837" box="[496,588,1074,1100]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
), with reference to the diurnal activity of the species of this subgenus.
</paragraph>
</subSubSection>
<subSubSection id="F2E6DD86FF9EFFA5FF3E70FA29EAD996" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="key">
<paragraph id="BA438E0DFF9EFFA4FF3E70FA2878DFD6" blockId="8.[151,1039,1179,1209]" box="[151,1039,1179,1209]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FF3E70FA2878DFD6" bold="true" box="[151,1039,1179,1209]" pageId="8" pageNumber="9">
Key to the species of the 
<taxonomicName id="7DFCF58EFF9EFFA4FE6270FA2E13DFD6" authorityName="Brenske" authorityYear="1897" box="[459,612,1182,1208]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="8888521FFF9EFFA4FE6270FA2E13DFD6" bold="true" box="[459,612,1182,1208]" italics="true" pageId="8" pageNumber="9">Gastroserica</emphasis>
</taxonomicName>
(
<taxonomicName id="7DFCF58EFF9EFFA4FDDC70FA2F77DFD7" authority="Ahrens &amp; Liu &amp; Lukic &amp; Bai, 2023" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" box="[629,768,1182,1209]" kingdom="Animalia" pageId="8" pageNumber="9" rank="subGenus" status="subgen. nov." subGenus="Helioserica">
<emphasis id="8888521FFF9EFFA4FDDC70FA2F77DFD7" bold="true" box="[629,768,1182,1209]" italics="true" pageId="8" pageNumber="9">Helioserica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93BBEF64FF9EFFA4FCAE70FA2FC0DFD6" box="[775,951,1182,1208]" pageId="8" pageNumber="9" rank="subGenus">new subgenus</taxonomicNameLabel>
) (♁♁):
</emphasis>
</paragraph>
<key id="CE6DD93CFF9EFFA5FF3E70B929EAD996" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<keyStep id="010896A8FF9EFFA4FF3E70B929EBDE61" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E70B929EBDF9C" blockId="8.[151,1437,1244,2029]" box="[151,1436,1244,1266]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E70B929EBDF9C" box="[151,1436,1244,1266]" pageId="8" pageNumber="9">1 Dorsal surface mostly dull or iridescent.................................................................... 2</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E709E29EBDE61" blockId="8.[151,1437,1244,2029]" box="[151,1436,1274,1296]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E709E29EBDE61" box="[151,1436,1274,1296]" pageId="8" pageNumber="9">- Dorsal surface completely simply shiny................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E717329EBDE24" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E717329EADE43" blockId="8.[151,1437,1244,2029]" box="[151,1437,1303,1325]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E717329EADE43" box="[151,1437,1303,1325]" pageId="8" pageNumber="9">
2 Elytra with multiple dark spots and darker striae.......................................... 
<taxonomicName id="7DFCF58EFF9EFFA4FB19717329EADE43" authority="(Frey, 1972)" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1200,1437,1303,1325]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cognata">
<emphasis id="8888521FFF9EFFA4FB19717329EADE43" bold="true" box="[1200,1437,1303,1325]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FB197173296BDE42" bold="true" box="[1200,1308,1303,1324]" italics="true" pageId="8" pageNumber="9">G. cognata</emphasis>
(
<bibRefCitation id="DE6DF3FCFF9EFFA4FA80717329E2DE43" author="Frey, G." box="[1321,1429,1303,1325]" pageId="8" pageNumber="9" pagination="186 - 216" refId="ref96076" refString="Frey, G. (1972) Neue Sericinen aus Indien und Indochina, sowie Abbildungen von Parameren bekannter Arten (Col., Scar., Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey, 23, 186 - 216." type="journal article" year="1972">Frey, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E715129EBDE24" blockId="8.[151,1437,1244,2029]" box="[151,1436,1332,1354]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E715129EBDE24" box="[151,1436,1332,1354]" pageId="8" pageNumber="9">- Elytra without multiple dark spots and striae of the same colour as intervals....................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E713629EADEEB" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E713629EBDE09" blockId="8.[151,1437,1244,2029]" box="[151,1436,1362,1384]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E713629EBDE09" box="[151,1436,1362,1384]" pageId="8" pageNumber="9">3 Left paramere with a distinct basal lobe.................................................................... 4</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E711429EADEEB" blockId="8.[151,1437,1244,2029]" box="[151,1437,1391,1413]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E711429EADEEB" box="[151,1437,1391,1413]" pageId="8" pageNumber="9">
- Left paramere without a distinct basal lobe............................................. 
<taxonomicName id="7DFCF58EFF9EFFA4FB3A710B29EADEEB" authority="(Frey, 1972)" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1171,1437,1391,1413]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="quateorum">
<emphasis id="8888521FFF9EFFA4FB3A710B29EADEEB" bold="true" box="[1171,1437,1391,1413]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FB3A710B296BDEEA" bold="true" box="[1171,1308,1391,1412]" italics="true" pageId="8" pageNumber="9">G. quateorum</emphasis>
(
<bibRefCitation id="DE6DF3FCFF9EFFA4FA80710B29E2DEEB" author="Frey, G." box="[1321,1429,1391,1413]" pageId="8" pageNumber="9" pagination="186 - 216" refId="ref96076" refString="Frey, G. (1972) Neue Sericinen aus Indien und Indochina, sowie Abbildungen von Parameren bekannter Arten (Col., Scar., Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey, 23, 186 - 216." type="journal article" year="1972">Frey, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E71E929EBDEAE" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E71E929EADECC" blockId="8.[151,1437,1244,2029]" box="[151,1437,1420,1442]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E71E929EADECC" box="[151,1437,1420,1442]" pageId="8" pageNumber="9">
4 Left distal portion of phallobase strongly widened (lateral view)............................ 
<taxonomicName id="7DFCF58EFF9EFFA4FB3271E829EADECC" authority="(Brenske, 1897)" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[1179,1437,1420,1442]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hiulca">
<emphasis id="8888521FFF9EFFA4FB3271E829EADECC" bold="true" box="[1179,1437,1420,1442]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FB3271E82880DECC" bold="true" box="[1179,1271,1420,1442]" italics="true" pageId="8" pageNumber="9">G. hiulca</emphasis>
(
<bibRefCitation id="DE6DF3FCFF9EFFA4FAAA71E829E2DECC" author="Brenske, E." box="[1283,1429,1420,1442]" pageId="8" pageNumber="9" pagination="345 - 468" refId="ref95715" refString="Brenske, E. (1897) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 42, 345 - 468. https: // doi. org / 10.1002 / mmnd. 18970420310" type="journal article" year="1897">Brenske, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E71CE29EBDEAE" blockId="8.[151,1437,1244,2029]" box="[151,1436,1450,1472]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E71CE29EBDEAE" box="[151,1436,1450,1472]" pageId="8" pageNumber="9">- Distal portion of phallobase at both sides narrowed towards apex (lateral view)..................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E71A329EBDE94" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E71A329EBDEB3" blockId="8.[151,1437,1244,2029]" box="[151,1436,1479,1501]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E71A329EBDEB3" box="[151,1436,1479,1501]" pageId="8" pageNumber="9">5 Left paramere bilobed in distal part....................................................................... 6</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E718129EBDE94" blockId="8.[151,1437,1244,2029]" box="[151,1436,1508,1530]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E718129EBDE94" box="[151,1436,1508,1530]" pageId="8" pageNumber="9">- Left paramere simple in distal part (with just one lobe or branch)............................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E726629EADD5B" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E726629EBDD79" blockId="8.[151,1437,1244,2029]" box="[151,1436,1538,1560]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E726629EBDD79" box="[151,1436,1538,1560]" pageId="8" pageNumber="9">6 Phallobase with a ventral lamellose extension on the left side.................................................. 7</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E724429EADD5B" blockId="8.[151,1437,1244,2029]" box="[151,1437,1567,1589]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E724429EADD5B" box="[151,1437,1567,1589]" pageId="8" pageNumber="9">
- Phallobase without a ventral lamellose extension on the left side......................... 
<taxonomicName id="7DFCF58EFF9EFFA4FBD4727B29EADD5B" authority="(Nomura, 1974)" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" baseAuthorityName="Nomura" baseAuthorityYear="1974" box="[1149,1437,1567,1589]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bisignata">
<emphasis id="8888521FFF9EFFA4FBD4727B29EADD5B" bold="true" box="[1149,1437,1567,1589]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FBD4727B2882DD5B" bold="true" box="[1149,1269,1567,1589]" italics="true" pageId="8" pageNumber="9">G. bisignata</emphasis>
(
<bibRefCitation id="DE6DF3FCFF9EFFA4FAA8727B29E2DD5B" author="Nomura, S." box="[1281,1429,1567,1589]" pageId="8" pageNumber="9" pagination="81 - 115" refId="ref97274" refString="Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115." type="journal article" year="1974">Nomura, 1974</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E725929EADDC4" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E725929EADD01" blockId="8.[151,1437,1244,2029]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E725929EADD01" pageId="8" pageNumber="9">
7 Left paramere robust, as wide as apical phallobase. Both distal lobes of left paramere subequal in length to the right paramere.................................................................... 
<taxonomicName id="7DFCF58EFF9EFFA4FBDC723E29EADD01" authority="(Nomura, 1974)" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" baseAuthorityName="Nomura" baseAuthorityYear="1974" box="[1141,1437,1625,1647]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nitidipyga">
<emphasis id="8888521FFF9EFFA4FBDC723E29EADD01" bold="true" box="[1141,1437,1625,1647]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FBDC723E2882DD01" bold="true" box="[1141,1269,1625,1647]" italics="true" pageId="8" pageNumber="9">G. nitidipyga</emphasis>
(
<bibRefCitation id="DE6DF3FCFF9EFFA4FAA8723D29E2DD01" author="Nomura, S." box="[1281,1429,1625,1647]" pageId="8" pageNumber="9" pagination="81 - 115" refId="ref97274" refString="Nomura, S. (1974) On the Sericini of Taiwan. Toho-Gakuho, 24, 81 - 115." type="journal article" year="1974">Nomura, 1974</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E721C29EADDC4" blockId="8.[151,1437,1244,2029]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E721C29EADDC4" pageId="8" pageNumber="9">
- Left paramere narrower, distinctly more narrow than apical phallobase. External distal lobes of left paramere half of length to the right paramere............................................. 
<emphasis id="8888521FFF9EFFA4FC1C72F029EADDC4" bold="true" box="[949,1437,1684,1706]" pageId="8" pageNumber="9">
<taxonomicName id="7DFCF58EFF9EFFA4FC1C72F028A1DDC4" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[949,1238,1684,1706]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yuebaensis">
<emphasis id="8888521FFF9EFFA4FC1C72F02849DDC4" bold="true" box="[949,1086,1684,1706]" italics="true" pageId="8" pageNumber="9">G. yuebaensis</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9EFFA4FA8F72F129EADDC4" box="[1318,1437,1684,1706]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E72D629EBDD8A" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E72D629EBDDA9" blockId="8.[151,1437,1244,2029]" box="[151,1436,1714,1736]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E72D629EBDDA9" box="[151,1436,1714,1736]" pageId="8" pageNumber="9">8 Right side of apical phallobase with a small granulate raster area................................................ 9</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E72B429EBDD8A" blockId="8.[151,1437,1244,2029]" box="[151,1436,1743,1765]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E72B429EBDD8A" box="[151,1436,1743,1765]" pageId="8" pageNumber="9">- Right side of apical phallobase without a granulate raster area................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E728929EADC34" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E728929EADC4E" blockId="8.[151,1437,1244,2029]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E728929EADC4E" pageId="8" pageNumber="9">
9 Left paramere basally wider (lateral view). Right paramere before apex almost straight............................................................................................. 
<emphasis id="8888521FFF9EFFA4FC78736E29EADC4E" bold="true" box="[977,1437,1801,1824]" pageId="8" pageNumber="9">
<taxonomicName id="7DFCF58EFF9EFFA4FC78736E28A1DC4E" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[977,1238,1801,1824]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fumaria">
<emphasis id="8888521FFF9EFFA4FC78736E2849DC71" bold="true" box="[977,1086,1801,1823]" italics="true" pageId="8" pageNumber="9">G. fumaria</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9EFFA4FA8F736E29EADC4E" box="[1318,1437,1802,1824]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E734C29EADC34" blockId="8.[151,1437,1244,2029]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E734C29EADC34" pageId="8" pageNumber="9">
- Left paramere basally more narrow (lateral view). Right paramere before apex strongly bent externally............................................................................ 
<emphasis id="8888521FFF9EFFA4FC0E732029EADC34" bold="true" box="[935,1437,1860,1882]" pageId="8" pageNumber="9">
<taxonomicName id="7DFCF58EFF9EFFA4FC0E732028A1DC34" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[935,1238,1860,1882]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="phukradung">
<emphasis id="8888521FFF9EFFA4FC0E73202837DC34" bold="true" box="[935,1088,1860,1882]" italics="true" pageId="8" pageNumber="9">G. phukradung</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9EFFA4FA8F732129EADC34" box="[1318,1437,1860,1882]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E730629EADCFB" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E730629EBDC19" blockId="8.[151,1437,1244,2029]" box="[151,1436,1890,1912]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E730629EBDC19" box="[151,1436,1890,1912]" pageId="8" pageNumber="9">10 Right paramere in basal two thirds strongly widened........................................................ 11</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E73E429EADCFB" blockId="8.[151,1437,1244,2029]" box="[151,1437,1919,1941]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E73E429EADCFB" box="[151,1437,1919,1941]" pageId="8" pageNumber="9">
- Right paramere in basal half not particularly widened..................................... 
<taxonomicName id="7DFCF58EFF9EFFA4FB0D731B29EADCFB" authority="(Moser, 1915)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1188,1437,1919,1941]" class="Insecta" family="Melolonthidae" genus="Microserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="8888521FFF9EFFA4FB0D731B29EADCFB" bold="true" box="[1188,1437,1919,1941]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FB0D731B297CDCFA" bold="true" box="[1188,1291,1919,1940]" italics="true" pageId="8" pageNumber="9">G. varians</emphasis>
(
<bibRefCitation id="DE6DF3FCFF9EFFA4FAB1731B29E2DCFB" author="Moser, J." box="[1304,1429,1919,1941]" pageId="8" pageNumber="9" pagination="337 - 393" refId="ref97123" refString="Moser, J. (1915) Neue Serica - Arten. Deutsche Entomologische Zeitschrift, 1915, 337 - 393. https: // doi. org / 10.1002 / mmnd. 48019150402" type="journal article" year="1915">Moser, 1915</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA4FF3E73F929EADCA1" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E73F929EBDCDC" blockId="8.[151,1437,1244,2029]" box="[151,1436,1948,1970]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E73F929EBDCDC" box="[151,1436,1948,1970]" pageId="8" pageNumber="9">11 Left paramere ventrally strongly excavated and bent......................................................... 12</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9EFFA4FF3E73DE29EADCA1" blockId="8.[151,1437,1244,2029]" box="[151,1437,1977,2000]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E73DE29EADCA1" box="[151,1437,1977,2000]" pageId="8" pageNumber="9">
- Left paramere ventrally not excavated and straight........................... 
<taxonomicName id="7DFCF58EFF9EFFA4FBBA73DE29EADCA1" authority="(Fabrizi &amp; Ahrens, 2016)" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" baseAuthorityName="Fabrizi &amp; Ahrens" baseAuthorityYear="2016" box="[1043,1437,1977,2000]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="roingensis">
<emphasis id="8888521FFF9EFFA4FBBA73DE29EADCA1" bold="true" box="[1043,1437,1977,2000]" pageId="8" pageNumber="9">
<emphasis id="8888521FFF9EFFA4FBBA73DE28E0DCA1" bold="true" box="[1043,1175,1978,1999]" italics="true" pageId="8" pageNumber="9">G. roingensis</emphasis>
(Fabrizi &amp; Ahrens, 2016)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9EFFA5FF3E73B329EADBC3" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<paragraph id="BA438E0DFF9EFFA4FF3E73B329EBDC82" blockId="8.[151,1437,1244,2029]" box="[151,1436,2007,2029]" pageId="8" pageNumber="9">
<keyLead id="010D2D38FF9EFFA4FF3E73B329EBDC82" box="[151,1436,2007,2029]" pageId="8" pageNumber="9">12 Right paramere at apex strongly bent externally (dorsal view).................................................. 13</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E74FC29EADBC3" blockId="9.[151,1437,151,760]" box="[151,1437,151,173]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E74FC29EADBC3" box="[151,1437,151,173]" pageId="9" pageNumber="10">
- Right paramere at apex weakly bent externally (dorsal view)..................... 
<taxonomicName id="7DFCF58EFF9FFFA5FB8E74F329EADBC3" authority="(Ahrens &amp; Fabrizi, 2009)" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" baseAuthorityName="Ahrens &amp; Fabrizi" baseAuthorityYear="2009" box="[1063,1437,151,173]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dohertyi">
<emphasis id="8888521FFF9FFFA5FB8E74F329EADBC3" bold="true" box="[1063,1437,151,173]" pageId="9" pageNumber="10">
<emphasis id="8888521FFF9FFFA5FB8E74F328E2DBC3" bold="true" box="[1063,1173,151,173]" italics="true" pageId="9" pageNumber="10">G. dohertyi</emphasis>
(Ahrens &amp; Fabrizi, 2009)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9FFFA5FF3E74D129EADA6B" pageId="9" pageNumber="10">
<paragraph id="BA438E0DFF9FFFA5FF3E74D129EBDBA4" blockId="9.[151,1437,151,760]" box="[151,1436,180,203]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E74D129EBDBA4" box="[151,1436,180,203]" pageId="9" pageNumber="10">
13 Elytra colour yellow to dark brown. Basal lobe of left paramere short.......................... 
<taxonomicName id="7DFCF58EFF9FFFA5FB1C74D129EBDBA4" authority="Frey, 1972" authorityName="Frey" authorityYear="1972" box="[1205,1436,180,202]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="8888521FFF9FFFA5FB1C74D129EBDBA4" bold="true" box="[1205,1436,180,202]" pageId="9" pageNumber="10">
<emphasis id="8888521FFF9FFFA5FB1C74D1295BDBA4" bold="true" box="[1205,1324,180,202]" italics="true" pageId="9" pageNumber="10">G. fukiensis</emphasis>
<bibRefCitation id="DE6DF3FCFF9FFFA5FA9B74D129EBDBA4" author="Frey, G." box="[1330,1436,181,202]" pageId="9" pageNumber="10" pagination="186 - 216" refId="ref96076" refString="Frey, G. (1972) Neue Sericinen aus Indien und Indochina, sowie Abbildungen von Parameren bekannter Arten (Col., Scar., Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey, 23, 186 - 216." type="journal article" year="1972">Frey, 1972</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E74B729EADA6B" blockId="9.[151,1437,151,760]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E74B729EADA6B" pageId="9" pageNumber="10">
- Elytra colour blackish, pronotum reddish. Basal lobe of left paramere long..................................................................................................... 
<emphasis id="8888521FFF9FFFA5FC1F748B29EADA6B" bold="true" box="[950,1437,239,261]" pageId="9" pageNumber="10">
<taxonomicName id="7DFCF58EFF9FFFA5FC1F748B28A1DA6B" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[950,1238,239,261]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="simaoensis">
<emphasis id="8888521FFF9FFFA5FC1F748B2848DA6A" bold="true" box="[950,1087,239,260]" italics="true" pageId="9" pageNumber="10">G. simaoensis</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9FFFA5FA8F749429EADA6B" box="[1318,1437,239,261]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9FFFA5FF3E756929EBDA51" pageId="9" pageNumber="10">
<paragraph id="BA438E0DFF9FFFA5FF3E756929EBDA4D" blockId="9.[151,1437,151,760]" box="[151,1436,269,291]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E756929EBDA4D" box="[151,1436,269,291]" pageId="9" pageNumber="10">14 Phallobase ventrally with prominent lateral elevations....................................................... 15</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E754F29EBDA51" blockId="9.[151,1437,151,760]" box="[151,1436,298,320]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E754F29EBDA51" box="[151,1436,298,320]" pageId="9" pageNumber="10">- Phallobase ventrally without prominent lateral elevations..................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9FFFA5FF3E752329EBDA14" pageId="9" pageNumber="10">
<paragraph id="BA438E0DFF9FFFA5FF3E752329EBDA33" blockId="9.[151,1437,151,760]" box="[151,1436,327,349]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E752329EBDA33" box="[151,1436,327,349]" pageId="9" pageNumber="10">15 Prominent lateral elevations of phallobase at apex.......................................................... 16</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E750129EBDA14" blockId="9.[151,1437,151,760]" box="[151,1436,357,379]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E750129EBDA14" box="[151,1436,357,379]" pageId="9" pageNumber="10">- Prominent lateral elevations of phallobase at middle......................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9FFFA5FF3E75E629EADA9E" pageId="9" pageNumber="10">
<paragraph id="BA438E0DFF9FFFA5FF3E75E629EADADB" blockId="9.[151,1437,151,760]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E75E629EADADB" pageId="9" pageNumber="10">
16 Right paramere more than half as long as phallobase. Eyes moderately large, ratio diameter/interocular width: 0.64................................................................. 
<emphasis id="8888521FFF9FFFA5FC3E75FB29EADADB" bold="true" box="[919,1437,415,437]" pageId="9" pageNumber="10">
<taxonomicName id="7DFCF58EFF9FFFA5FC3E75FB28A1DADB" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[919,1238,415,437]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="piceocoerulea">
<emphasis id="8888521FFF9FFFA5FC3E75FB2849DADB" bold="true" box="[919,1086,415,437]" italics="true" pageId="9" pageNumber="10">G. piceocoerulea</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9FFFA5FA8F75C429EADADB" box="[1318,1437,415,437]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E75D929EADA9E" blockId="9.[151,1437,151,760]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E75D929EADA9E" pageId="9" pageNumber="10">
- Right paramere less than half as long as phallobase. Eyes very large, ratio diameter/interocular width: 0.93.................................................................................. 
<emphasis id="8888521FFF9FFFA5FC5775BE29EADA9E" bold="true" box="[1022,1437,474,496]" pageId="9" pageNumber="10">
<taxonomicName id="7DFCF58EFF9FFFA5FC5775BE28A1DA9E" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[1022,1238,474,496]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="loei">
<emphasis id="8888521FFF9FFFA5FC5775BE2848DA9E" bold="true" box="[1022,1087,474,496]" italics="true" pageId="9" pageNumber="10">G. loei</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9FFFA5FA8F75BE29EADA9E" box="[1318,1437,474,496]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9FFFA5FF3E759C29EAD926" pageId="9" pageNumber="10">
<paragraph id="BA438E0DFF9FFFA5FF3E759C29EBD963" blockId="9.[151,1437,151,760]" box="[151,1436,503,525]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E759C29EBD963" box="[151,1436,503,525]" pageId="9" pageNumber="10">17 Left paramere with a distinct and subequal ventral and dorsal/basal lobe......................................... 18</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E767129EAD926" blockId="9.[151,1437,151,760]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E767129EAD926" pageId="9" pageNumber="10">
- Left paramere without a ventral and dorsal/basal lobe, the latter basally reduced in length and developed short and medially........................................................................ 
<taxonomicName id="7DFCF58EFF9FFFA5FB91765629EAD926" authority="(Ahrens &amp; Fabrizi, 2009)" authorityName="Ahrens &amp; Liu &amp; Lukic &amp; Bai" authorityYear="2023" baseAuthorityName="Ahrens &amp; Fabrizi" baseAuthorityYear="2009" box="[1080,1437,562,584]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="8888521FFF9FFFA5FB91765629EAD926" bold="true" box="[1080,1437,562,584]" pageId="9" pageNumber="10">
<emphasis id="8888521FFF9FFFA5FB91765628E2D926" bold="true" box="[1080,1173,562,584]" italics="true" pageId="9" pageNumber="10">G. lucens</emphasis>
(Ahrens &amp; Fabrizi, 2009)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9FFFA5FF3E763429EAD9CE" pageId="9" pageNumber="10">
<paragraph id="BA438E0DFF9FFFA5FF3E763429EAD90B" blockId="9.[151,1437,151,760]" box="[151,1437,591,613]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E763429EAD90B" box="[151,1437,591,613]" pageId="9" pageNumber="10">
18 Dorsal/basal lobe shorter than the ventral one, sharply pointed, simple......... 
<emphasis id="8888521FFF9FFFA5FC54762B29EAD90B" bold="true" box="[1021,1437,591,613]" pageId="9" pageNumber="10">
<taxonomicName id="7DFCF58EFF9FFFA5FC54762B28A1D90B" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[1021,1238,591,613]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ivoi">
<emphasis id="8888521FFF9FFFA5FC54762B2849D90A" bold="true" box="[1021,1086,591,612]" italics="true" pageId="9" pageNumber="10">G. ivoi</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9FFFA5FA8F763429EAD90B" box="[1318,1437,591,613]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E760929EAD9CE" blockId="9.[151,1437,151,760]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E760929EAD9CE" pageId="9" pageNumber="10">
- Dorsal/basal lobe as long as the ventral one, rounded at apex, tubuliform with a membranous opening at median apex.............................................................. 
<emphasis id="8888521FFF9FFFA5FCD376EE29EAD9CE" bold="true" box="[890,1437,650,672]" pageId="9" pageNumber="10">
<taxonomicName id="7DFCF58EFF9FFFA5FCD376EE28A1D9CE" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[890,1238,650,672]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lucidomarginalis">
<emphasis id="8888521FFF9FFFA5FCD376EE2848D9CE" bold="true" box="[890,1087,650,672]" italics="true" pageId="9" pageNumber="10">G. lucidomarginalis</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9FFFA5FA8F76EE29EAD9CE" box="[1318,1437,650,672]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="010896A8FF9FFFA5FF3E76CC29EAD996" pageId="9" pageNumber="10">
<paragraph id="BA438E0DFF9FFFA5FF3E76CC29EAD9D3" blockId="9.[151,1437,151,760]" box="[151,1437,679,701]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E76CC29EAD9D3" box="[151,1437,679,701]" pageId="9" pageNumber="10">
19 Left paramere with a distinct and subequal ventral and dorsal lobe....... 
<emphasis id="8888521FFF9FFFA5FC1F76C329EAD9D3" bold="true" box="[950,1437,679,701]" pageId="9" pageNumber="10">
<taxonomicName id="7DFCF58EFF9FFFA5FC1F76C328A1D9D3" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[950,1238,679,701]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rubropicea">
<emphasis id="8888521FFF9FFFA5FC1F76C32848D9D3" bold="true" box="[950,1087,679,701]" italics="true" pageId="9" pageNumber="10">G. rubropicea</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9FFFA5FA8F76CC29EAD9D3" box="[1318,1437,679,701]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="BA438E0DFF9FFFA5FF3E76A129EAD996" blockId="9.[151,1437,151,760]" pageId="9" pageNumber="10">
<keyLead id="010D2D38FF9FFFA5FF3E76A129EAD996" pageId="9" pageNumber="10">
- Left paramere without a ventral and dorsal lobe, the latter basally reduced in length and developed short and medially.................................................................... 
<emphasis id="8888521FFF9FFFA5FC7F768629EAD996" bold="true" box="[982,1437,738,760]" pageId="9" pageNumber="10">
<taxonomicName id="7DFCF58EFF9FFFA5FC7F768628A1D996" authority="Ahrens, Lukic" authorityName="Ahrens, Lukic" box="[982,1238,738,760]" class="Insecta" family="Melolonthidae" genus="Gastroserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bannok">
<emphasis id="8888521FFF9FFFA5FC7F76862849D996" bold="true" box="[982,1086,738,760]" italics="true" pageId="9" pageNumber="10">G. bannok</emphasis>
Ahrens, Lukic
</taxonomicName>
&amp; Liu, 
<taxonomicNameLabel id="93BBEF64FF9FFFA5FA8F768629EAD996" box="[1318,1437,738,760]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>