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<document id="7C1E2A9E4B961E38FEBD7AD5B7C01921" ID-CLB-Dataset="55344" ID-DOI="10.5852/ejt.2023.880.2173" ID-GBIF-Dataset="339bce42-a7e7-418b-b9ac-4588e4219b15" ID-ISSN="2118-9773" ID-Zenodo-Dep="8155472" ID-ZooBank="3F806FD6-2EB3-456A-AFD7-780A0FBEB2DA" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1689603386695" checkinUser="felipe" docAuthor="Huber, Bernhard A., Meng, Guanliang, Valdez-Mondragón, Alejandro, Král, Jiří, Ávila Herrera, Ivalú M. &amp; Carvalho, Leonardo S." docDate="2023" docId="03F087F1AC5AFFFA096E652EE75DFC70" docLanguage="en" docName="ejt-880_1-89.pdf" docOrigin="European Journal of Taxonomy 880 (1)" docSource="http://zoobank.org/3f806fd6-2eb3-456a-afd7-780a0fbeb2da" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pholcophora Banks 1896" docType="treatment" docVersion="6" lastPageNumber="45" masterDocId="FFC9FF89AC76FFD60B146426E305FFD0" masterDocTitle="Short-legged daddy-long-leg spiders in North America: the genera Pholcophora and Tolteca (Araneae, Pholcidae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="45" updateTime="1699540719498" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A08D469D2C69A7AE02D65A2F60AA3F96">Short-legged daddy-long-leg spiders in North America: the genera Pholcophora and Tolteca (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="CA68AC9CAA9FBB897746B02B3FAC1BE6">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="A8A395BEDC9ADED09A60D972270EBD0D">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany. Laboratory of Arachnology (LATLAX), Institute of Biology, Universidad Nacional Autónoma de México-Tlaxcala, San Miguel Contla, Santa Cruz Tlaxcala, Tlaxcala, México. Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic. Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, Floriano, Piauí, Brazil.</mods:affiliation>
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<mods:namePart id="595F596F28858305A83994F4E382E653">Meng, Guanliang</mods:namePart>
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<mods:affiliation id="421697FA5BE2E164FE64A89B7E31A694">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany. Laboratory of Arachnology (LATLAX), Institute of Biology, Universidad Nacional Autónoma de México-Tlaxcala, San Miguel Contla, Santa Cruz Tlaxcala, Tlaxcala, México. Department of Genetics and Microbiology, Faculty of Science, Charles University, Prague, Czech Republic. Campus Amílcar Ferreira Sobral, Universidade Federal do Piauí, Floriano, Piauí, Brazil.</mods:affiliation>
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<mods:namePart id="6AC88451D156EDDCF1D506B9BB6ED0D1">Valdez-Mondragón, Alejandro</mods:namePart>
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<mods:namePart id="3D19703B3DF067D574826474660F9EE6">Král, Jiří</mods:namePart>
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<mods:namePart id="8D31A33446B6DB8A0D66604210D2E6FF">Ávila Herrera, Ivalú M.</mods:namePart>
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<taxonomicName id="4C594D64AC5AFFFA096E652EE013FEF3" ID-CoL="6NWL" ID-ENA="393828" authorityName="Banks" authorityYear="1896" box="[634,790,264,291]" class="Arachnida" family="Pholcidae" genus="Pholcophora" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF5AC5AFFFA096E652EE013FEF3" bold="true" box="[634,790,264,291]" italics="true" pageId="44" pageNumber="45">Pholcophora</emphasis>
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? “Cu12-325”
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<figureCitation id="13622A62AC5AFFFA09DA650AE022FE96" box="[718,807,300,326]" captionStart="Fig" captionStartId="42.[189,232,1892,1918]" captionTargetBox="[200,1388,524,1848]" captionTargetId="figure-78@42.[200,1388,524,1848]" captionTargetPageId="42" captionText="Fig. 32. Pholcophora? “Cu12-325”, female from Cuba, Santiago de Cuba, Siboney (ZFMK Cu12-325). AB. Habitus, dorsal and lateral views. CD. Abdomen, ventral and lateral views. E. Cleared female genitalia, ventral view. F. Detail of median internal structures, dorsal view. Scale bars: AB = 1 mm; CD = 0.3 mm; EF = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8155568" httpUri="https://zenodo.org/record/8155568/files/figure.png" pageId="44" pageNumber="45">
Figs
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,
<figureCitation id="13622A62AC5AFFFA0827650AE063FE96" box="[819,870,300,326]" captionStart="Fig" captionStartId="43.[189,232,1681,1707]" captionTargetBox="[246,1335,267,1633]" captionTargetId="figure-12@43.[246,1342,265,1633]" captionTargetPageId="43" captionText="Fig. 33. Pholcophora internal female genitalia, median sections of main transversal internal sclerite. AB. P. americana Banks, 1896, females from USA, California, Inyo Nat. Forest (ZFMK G089) and Colorado, near Golden (ZFMK USA16). CD. P. mazatlan Huber sp. nov., females from Mexico, Guerrero, N of Mazatlán (ZFMK Ar 23943). EF. P. papanoa Huber sp. nov., females from Mexico, Guerrero, S of Papanoa (ZFMK Ar 23945). GH. P. “Mex354”, females from Mexico, Puebla, SE of Tehuacan (ZFMK Mex354). IJ. P. texana Gertsch, 1935, females from Mexico, Hidalgo, SW of Jacala (ZFMK Ar 23952). KL. P. tehuacan Huber sp. nov., females from Mexico, Puebla, SE of Tehuacan (ZFMK Ar 23949). M. Pholcophora? “Car544”, female from Puerto Rico, Isla Mono (USNM ENT 783464). N. Pholcophora? “Cu12-325”, female from Cuba, Santiago de Cuba, Siboney (ZFMK Cu12- 325). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8155572" httpUri="https://zenodo.org/record/8155572/files/figure.png" pageId="44" pageNumber="45">33N</figureCitation>
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S323 Gen.
<emphasis id="B92DEAF5AC5AFFFA0A546548E2B0FE58" box="[320,437,366,392]" italics="true" pageId="44" pageNumber="45">Cu12-325</emphasis>
Cu12-325
<bibRefCitation id="EFC84B16AC5AFFFA095B6548E025FE58" author="Eberle J. &amp; Dimitrov D. &amp; Valdez-Mondragon A. &amp; Huber B. A." box="[591,800,366,392]" pageId="44" pageNumber="45" pagination="141" refId="ref33384" refString="Eberle J., Dimitrov D., Valdez-Mondragon A. &amp; Huber B. A. 2018. Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology 18: 141. https: // doi. org / 10.1186 / s 12862 - 018 - 1244 - 8" type="journal article" year="2018">
Eberle
<emphasis id="B92DEAF5AC5AFFFA09B76548E1DBFE58" box="[675,734,366,392]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2018
</bibRefCitation>
(molecular data). —
<bibRefCitation id="EFC84B16AC5AFFFA0F086548E61CFE58" author="Huber B. A. &amp; Eberle J. &amp; Dimitrov D." box="[1052,1305,365,392]" pageId="44" pageNumber="45" pagination="51 - 101" refId="ref34395" refString="Huber B. A., Eberle J. &amp; Dimitrov D. 2018. The phylogeny of pholcid spiders: a critical evaluation of relationships suggested by molecular data (Araneae, Pholcidae). ZooKeys 789: 51 - 101. https: // doi. org / 10.3897 / zookeys. 789.22781" type="journal article" year="2018">
Huber
<emphasis id="B92DEAF5AC5AFFFA0F786548E7A2FE58" box="[1132,1191,366,392]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2018: 55
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.
</paragraph>
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<emphasis id="B92DEAF5AC5AFFFA0BA96589E22AFE19" bold="true" box="[189,303,431,457]" pageId="44" pageNumber="45">Remarks</emphasis>
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The
<specimenCount id="9D5FFD6EAC5AFFFA0BEC65F8E269FE27" box="[248,364,477,504]" pageId="44" pageNumber="45" type="adult">two adult</specimenCount>
females available of this species resemble other large representatives of the genus (
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); body length 2.4; tibia 1 length 1.28, 1.36. Epigynum (
<figureCitation id="13622A62AC5AFFFA08D06627E74BFDCB" box="[964,1102,513,539]" captionStart="Fig" captionStartId="42.[189,232,1892,1918]" captionTargetBox="[200,1388,524,1848]" captionTargetId="figure-78@42.[200,1388,524,1848]" captionTargetPageId="42" captionText="Fig. 32. Pholcophora? “Cu12-325”, female from Cuba, Santiago de Cuba, Siboney (ZFMK Cu12-325). AB. Habitus, dorsal and lateral views. CD. Abdomen, ventral and lateral views. E. Cleared female genitalia, ventral view. F. Detail of median internal structures, dorsal view. Scale bars: AB = 1 mm; CD = 0.3 mm; EF = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8155568" httpUri="https://zenodo.org/record/8155568/files/figure.png" pageId="44" pageNumber="45">Fig. 32CD</figureCitation>
) consisting of trapezoidal anterior plate and short but wide posterior plate; internal genitalia (
<figureCitation id="13622A62AC5AFFFA08DF6602E757FDEE" box="[971,1106,548,574]" captionStart="Fig" captionStartId="42.[189,232,1892,1918]" captionTargetBox="[200,1388,524,1848]" captionTargetId="figure-78@42.[200,1388,524,1848]" captionTargetPageId="42" captionText="Fig. 32. Pholcophora? “Cu12-325”, female from Cuba, Santiago de Cuba, Siboney (ZFMK Cu12-325). AB. Habitus, dorsal and lateral views. CD. Abdomen, ventral and lateral views. E. Cleared female genitalia, ventral view. F. Detail of median internal structures, dorsal view. Scale bars: AB = 1 mm; CD = 0.3 mm; EF = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8155568" httpUri="https://zenodo.org/record/8155568/files/figure.png" pageId="44" pageNumber="45">Fig. 32EF</figureCitation>
) with distinctive median tube, curled up and ~600 µm long (reminding of the putatively distantly related
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<emphasis id="B92DEAF5AC5AFFFA0F676661E672FDB1" box="[1139,1399,583,609]" italics="true" pageId="44" pageNumber="45">Gertschiola neuquena</emphasis>
<bibRefCitation id="EFC84B16AC5AFFFA0BA9664CE257FD54" author="Huber B. A." box="[189,338,618,644]" pageId="44" pageNumber="45" pagination="1 - 348" refId="ref34125" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / brh 26 h" type="journal article" year="2000">Huber, 2000</bibRefCitation>
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; see
<bibRefCitation id="EFC84B16AC5AFFFA0A9F664CE11FFD54" author="Huber B. A." box="[395,538,618,644]" pageId="44" pageNumber="45" pagination="1 - 348" refId="ref34125" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / brh 26 h" type="journal article" year="2000">Huber 2000</bibRefCitation>
: fig. 354).
</paragraph>
<paragraph id="8BE636E7AC5AFFFA0BA9668AE297FCDD" blockId="44.[189,1399,684,781]" pageId="44" pageNumber="45">
Our molecular data suggest that the present species is more closely related with other Caribbean taxa (
<taxonomicName id="4C594D64AC5AFFFA0BD366E9E2C3FD39" authorityName="Gertsch" authorityYear="1982" box="[199,454,719,745]" class="Arachnida" family="Pholcidae" genus="Pholcophora" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="bahama">
<emphasis id="B92DEAF5AC5AFFFA0BD366E9E2C3FD39" box="[199,454,719,745]" italics="true" pageId="44" pageNumber="45">Pholcophora bahama</emphasis>
</taxonomicName>
, the undescribed
<taxonomicName id="4C594D64AC5AFFFA098166E9E02BFD39" authorityName="Banks" authorityYear="1896" box="[661,814,719,745]" class="Arachnida" family="Pholcidae" genus="Pholcophora" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF5AC5AFFFA098166E9E02BFD39" box="[661,814,719,745]" italics="true" pageId="44" pageNumber="45">Pholcophora</emphasis>
</taxonomicName>
? “
<taxonomicName id="4C594D64AC5AFFFA085966E9E07EFD3A" box="[845,891,719,746]" class="Insecta" family="Caridae" genus="Car" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="40" phylum="Arthropoda" rank="genus">Car</taxonomicName>
544”, and the genus
<taxonomicName id="4C594D64AC5AFFFA0F7E66F6E7FDFD39" authorityName="Huber" authorityYear="2000" box="[1130,1272,720,745]" class="Arachnida" family="Pholcidae" genus="Papiamenta" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF5AC5AFFFA0F7E66F6E7FDFD39" box="[1130,1272,720,745]" italics="true" pageId="44" pageNumber="45">Papiamenta</emphasis>
</taxonomicName>
) than with true
<taxonomicName id="4C594D64AC5AFFFA0BE666D4E28EFCDC" authorityName="Banks" authorityYear="1896" box="[242,395,754,780]" class="Arachnida" family="Pholcidae" genus="Pholcophora" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF5AC5AFFFA0BE666D4E28EFCDC" box="[242,395,754,780]" italics="true" pageId="44" pageNumber="45">Pholcophora</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C343656CAC5AFFFA0BA96715E75DFC70" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BE636E7AC5AFFFA0BA96715E2AFFC9D" blockId="44.[189,426,819,845]" box="[189,426,819,845]" pageId="44" pageNumber="45">
<heading id="D0AE818BAC5AFFFA0BA96715E2AFFC9D" bold="true" box="[189,426,819,845]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B92DEAF5AC5AFFFA0BA96715E2AFFC9D" bold="true" box="[189,426,819,845]" pageId="44" pageNumber="45">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE636E7AC5AFFFA0BA96744E75DFC70" blockId="44.[189,1399,866,928]" pageId="44" pageNumber="45">
<materialsCitation id="3B313CBAAC5AFFFA0BA96744E750FC70" ID-GBIF-Occurrence="4161533335" collectingDate="2012-04" collectionCode="ZFMK" collectorName="F. Cala Riquelme" country="Cuba" county="Siboney" elevation="90" latitude="19.9623" location="Cueva de los Majases" longLatPrecision="7" longitude="-75.7171" municipality="Cueva de los Majases" pageId="44" pageNumber="45" specimenCount="2" specimenCount-female="2" stateProvince="Santiago de Cuba">
<collectingCountry id="F34E7677AC5AFFFA0BA96744E208FCAC" box="[189,269,866,892]" name="Cuba" pageId="44" pageNumber="45">CUBA</collectingCountry>
<emphasis id="B92DEAF5AC5AFFFA0A256744E110FCAC" bold="true" box="[305,533,866,892]" pageId="44" pageNumber="45">
<collectingRegion id="499DF805AC5AFFFA0A256744E110FCAC" box="[305,533,866,892]" country="Cuba" name="Santiago de Cuba" pageId="44" pageNumber="45">Santiago de Cuba</collectingRegion>
</emphasis>
<specimenCount id="9D5FFD6EAC5AFFFA09226744E179FCAD" box="[566,636,866,893]" pageId="44" pageNumber="45" type="female">2 ♀♀</specimenCount>
, in pure ethanol;
<collectingCounty id="62874E6BAC5AFFFA08436744E0BDFCAC" box="[855,952,866,892]" pageId="44" pageNumber="45">Siboney</collectingCounty>
,
<location id="8E86603CAC5AFFFA08DC6744E7D0FCAC" LSID="urn:lsid:plazi:treatment:03F087F1AC5AFFFA096E652EE75DFC70:8E86603CAC5AFFFA08DC6744E7D0FCAC" box="[968,1237,866,892]" country="Cuba" county="Siboney" latitude="19.9623" longLatPrecision="7" longitude="-75.7171" municipality="Cueva" name="Majases" pageId="44" pageNumber="45" stateProvince="Santiago de Cuba">
<collectingMunicipality id="6B82AC9DAC5AFFFA08DC6744E716FCAC" box="[968,1043,866,892]" pageId="44" pageNumber="45">Cueva</collectingMunicipality>
de los Majases
</location>
;
<geoCoordinate id="EE6D5020AC5AFFFA0FF06744E677FCAD" box="[1252,1394,866,893]" degrees="19.9623" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="5" value="19.9623">19.9623° N</geoCoordinate>
,
<geoCoordinate id="EE6D5020AC5AFFFA0BA967A3E248FC70" box="[189,333,901,928]" degrees="75.7171" direction="west" orientation="longitude" pageId="44" pageNumber="45" precision="5" value="-75.7171">75.7171° W</geoCoordinate>
; ~
<elevation id="0074D1D4AC5AFFFA0A7C67A3E2E1FC4F" box="[360,484,901,928]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="45" unit="m" value="90.0">
<quantity id="4CA19B02AC5AFFFA0A7C67A3E2A1FC70" box="[360,420,901,928]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="45" unit="m" value="90.0">90 m</quantity>
a.s.l.
</elevation>
;
<date id="FFE71027AC5AFFFA0AE467A0E16CFC4F" box="[496,617,901,928]" pageId="44" pageNumber="45" value="2012-04">
<collectingDate id="EFA3E9CFAC5AFFFA0AE467A0E16CFC4F" box="[496,617,901,928]" pageId="44" pageNumber="45" value="2012-04">Apr. 2012</collectingDate>
</date>
;
<collectorName id="26AC5331AC5AFFFA096167A0E044FC4F" box="[629,833,901,928]" pageId="44" pageNumber="45">F. Cala Riquelme</collectorName>
leg.;
<collectionCode id="ED48AE22AC5AFFFA089767A0E0D1FC70" box="[899,980,902,928]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="44" pageNumber="45" type="Museum">ZFMK</collectionCode>
<collectorName id="26AC5331AC5AFFFA08C867A3E704FC70" box="[988,1025,901,928]" pageId="44" pageNumber="45">Cu</collectorName>
12-325
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>