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<mods:title id="EE6E0182ABF7058F6F77A490982CD023">Phylogeny and revision of a colorful Neotropical genus of rove beetles: Xenopygus Bernhauer (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="6C31536D9D0B044B165B5A7C0EEE115F">Caron, Edilson</mods:namePart>
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<subSubSection id="C3436526FFA3FF9929996189FDEBFF1C" box="[151,535,151,178]" pageId="18" pageNumber="77" type="nomenclature">
<paragraph id="8BE636ADFFA3FF9929996189FDEBFF1C" blockId="18.[151,535,151,178]" box="[151,535,151,178]" pageId="18" pageNumber="77">
<heading id="D0AE81C1FFA3FF9929996189FDEBFF1C" bold="true" box="[151,535,151,178]" fontSize="11" level="1" pageId="18" pageNumber="77" reason="1">
<taxonomicName id="4C594D2EFFA3FF9929996189FDEBFF1C" ID-CoL="5CBD4" authority="Solsky" authorityName="Solsky" box="[151,535,151,178]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF9929996189FDEBFF1C" bold="true" box="[151,535,151,178]" pageId="18" pageNumber="77">
<emphasis id="B92DEABFFFA3FF9929996189FE57FF1F" bold="true" box="[151,427,151,177]" italics="true" pageId="18" pageNumber="77">Xenopygus peruvianus</emphasis>
(Solsky)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3436526FFA3FF99299961C0FB47FE9C" pageId="18" pageNumber="77" type="reference_group">
<paragraph id="8BE636ADFFA3FF99299961C0FC9FFEBD" blockId="18.[151,1408,222,306]" pageId="18" pageNumber="77">
<treatmentCitationGroup id="AB491183FFA3FF99299961C0FC9FFEBD" pageId="18" pageNumber="77">
<taxonomicName id="4C594D2EFFA3FF99299961C0FDA9FF5A" ID-CoL="3S53W" authority="Solsky, 1872: 306" authorityName="Solsky" authorityPageNumber="306" authorityYear="1872" box="[151,597,222,245]" class="Insecta" family="Staphylinidae" genus="Xanthopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF99299961C0FE6DFF5B" box="[151,401,222,245]" italics="true" pageId="18" pageNumber="77">Xanthopygus peruvianus</emphasis>
:
<treatmentCitation id="0AF810BCFFA3FF99289161C0FDA9FF5A" author="Solsky" box="[415,597,222,244]" page="306" pageId="18" pageNumber="77" year="1872">
<bibRefCitation id="EFC84B5CFFA3FF99289161C0FDD9FF5A" author="Solsky" box="[415,549,222,244]" pageId="18" pageNumber="77" refString="Solsky, S. M. (1872) Enumeration et description des coleopteres de la famille des Staphylinides recueillis par Mrs. C. Jelsky et le Baron de Nolken pendant leurs voyages dans l'Amerique du Sud en 1870 et 1871. Horae Societatis Entomologicae Rossicae, 8, 289 - 314." type="journal article" year="1872">Solsky, 1872</bibRefCitation>
: 306
</treatmentCitation>
</taxonomicName>
(original description,
<typeStatus id="54E2880FFFA3FF992A3861C1FC9EFF5A" box="[822,866,223,244]" pageId="18" pageNumber="77">type</typeStatus>
locality: Monte-Rico,
<collectingCountry id="F34E763DFFA3FF992D5761C1FB76FF5A" box="[1113,1162,223,244]" name="Peru" pageId="18" pageNumber="77">Peru</collectingCountry>
, cited as
<emphasis id="B92DEABFFFA3FF992DE961C0FA8EFF5A" box="[1255,1394,222,244]" italics="true" pageId="18" pageNumber="77">Xanthopygius</emphasis>
).
<taxonomicName id="4C594D2EFFA3FF99299961E0FD07FEBD" ID-CoL="3S53W" authority="Bernhauer &amp; Schubert, 1914: 406" authorityName="Bernhauer &amp; Schubert" authorityPageNumber="406" authorityYear="1914" box="[151,763,253,275]" class="Insecta" family="Staphylinidae" genus="Lampropygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF99299961E0FE6BFEBD" box="[151,407,254,275]" italics="true" pageId="18" pageNumber="77">Lampropygus peruvianus</emphasis>
:
<treatmentCitation id="0AF810BCFFA3FF9928AB61E3FD07FEBD" author="Bernhauer" box="[421,763,253,275]" page="406" pageId="18" pageNumber="77" year="1914">
<bibRefCitation id="EFC84B5CFFA3FF9928AB61E3FD37FEBD" author="Bernhauer" box="[421,715,253,275]" pageId="18" pageNumber="77" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: S. Schenkling (Ed.) Coleopterorum Catalogus. 5 (57). Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer &amp; Schubert, 1914</bibRefCitation>
: 406
</treatmentCitation>
</taxonomicName>
(catalog).
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</paragraph>
<paragraph id="8BE636ADFFA3FF9929996003FB47FE9C" blockId="18.[151,1408,222,306]" box="[151,1211,284,306]" pageId="18" pageNumber="77">
<treatmentCitationGroup id="AB491183FFA3FF9929996003FB47FE9C" box="[151,1211,284,306]" pageId="18" pageNumber="77">
<taxonomicName id="4C594D2EFFA3FF9929996003FD83FE9C" ID-CoL="5CBD4" authority="Scheerpeltz, 1933: 1416" authorityName="Scheerpeltz" authorityPageNumber="1416" authorityYear="1933" box="[151,639,284,306]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF9929996003FE80FE9C" box="[151,380,285,306]" italics="true" pageId="18" pageNumber="77">Xenopygus peruvianus</emphasis>
:
<treatmentCitation id="0AF810BCFFA3FF9928846002FD83FE9C" author="Scheerpeltz" box="[394,639,284,306]" page="1416" pageId="18" pageNumber="77" year="1933">
<bibRefCitation id="EFC84B5CFFA3FF9928846002FDBEFE9C" author="Scheerpeltz" box="[394,578,284,306]" pageId="18" pageNumber="77" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. 6 (129). Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1416
</treatmentCitation>
</taxonomicName>
(catalog);
<treatmentCitation id="0AF810BCFFA3FF992BE36002FC6DFE9C" author="Irmler" box="[749,913,284,306]" page="31" pageId="18" pageNumber="77" year="1979">
<bibRefCitation id="EFC84B5CFFA3FF992BE36002FC92FE9C" author="Irmler" box="[749,878,284,306]" pageId="18" pageNumber="77" refString="Irmler, U. (1979) Taxonomie, Verbreitung und Biologie der neotropischen Staphylinidengattung Xenopygus Bernh. (Coleoptera, Staphylinidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75, 30 - 36." type="journal article" year="1979">Irmler, 1979</bibRefCitation>
: 31
</treatmentCitation>
(considered
<emphasis id="B92DEABFFFA3FF992D1A6003FB57FE9C" box="[1044,1195,285,306]" italics="true" pageId="18" pageNumber="77">species incerta</emphasis>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3436526FFA3FF9929996047FC75FC78" pageId="18" pageNumber="77" type="discussion">
<paragraph id="8BE636ADFFA3FF9929996047FE5AFD88" blockId="18.[151,1437,345,982]" pageId="18" pageNumber="77">
<emphasis id="B92DEABFFFA3FF9929996047FEF7FEDC" bold="true" box="[151,267,345,370]" pageId="18" pageNumber="77">Remarks.</emphasis>
This species was described primarily in
<taxonomicName id="4C594D2EFFA3FF992BDF6047FC9FFEDF" box="[721,867,345,369]" class="Insecta" family="Staphylinidae" genus="Xanthopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEABFFFA3FF992BDF6047FC9FFEDF" box="[721,867,345,369]" italics="true" pageId="18" pageNumber="77">Xanthopygus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC84B5CFFA3FF992A836047FBE1FEDF" author="Solsky" box="[909,1053,345,370]" pageId="18" pageNumber="77" refString="Solsky, S. M. (1872) Enumeration et description des coleopteres de la famille des Staphylinides recueillis par Mrs. C. Jelsky et le Baron de Nolken pendant leurs voyages dans l'Amerique du Sud en 1870 et 1871. Horae Societatis Entomologicae Rossicae, 8, 289 - 314." type="journal article" year="1872">Solsky, 1872</bibRefCitation>
(cited as
<emphasis id="B92DEABFFFA3FF992D866047FADEFEDF" box="[1160,1314,345,369]" italics="true" pageId="18" pageNumber="77">Xanthopygius</emphasis>
), based on individuals from Monte Rico,
<collectingCountry id="F34E763DFFA3FF9928FD6060FDD6FE3B" box="[499,554,382,405]" name="Peru" pageId="18" pageNumber="77">Peru</collectingCountry>
. In the original description, the author stated that this species is similar to
<taxonomicName id="4C594D2EFFA3FF992C8A6060FF1AFE17" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B92DEABFFFA3FF992C8A6060FF1AFE17" italics="true" pageId="18" pageNumber="77">X. bicolor</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC84B5CFFA3FF9929F760BFFE69FE17" author="Irmler" box="[249,405,417,442]" pageId="18" pageNumber="77" refString="Irmler, U. (1982) A new species of the Neotropical genus Xenopygus Bernhauer (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 36 (2), 206 - 210." type="journal article" year="1982">Irmler (1982)</bibRefCitation>
was looking for the species in Solskys collection, which is partially deposited in ZIN (Zoological Institute, Russian Academy of Sciences, Saint Petersburg,
<collectingCountry id="F34E763DFFA3FF992AA260DBFC04FE70" box="[940,1016,453,478]" name="Russia" pageId="18" pageNumber="77">Russia</collectingCountry>
) and ZMMU (Zoological Museum of Moscow University, Moscow,
<collectingCountry id="F34E763DFFA3FF9928E360F7FDC0FDAC" box="[493,572,489,514]" name="Russia" pageId="18" pageNumber="77">Russia</collectingCountry>
), and reported
<taxonomicName id="4C594D2EFFA3FF992BEC60F4FC81FDAF" box="[738,893,490,513]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF992BEC60F4FC81FDAF" box="[738,893,490,513]" italics="true" pageId="18" pageNumber="77">X. peruvianus</emphasis>
</taxonomicName>
as
<emphasis id="B92DEABFFFA3FF992AAD60F4FBB5FDAF" box="[931,1097,490,513]" italics="true" pageId="18" pageNumber="77">species incerta</emphasis>
, since he did not find the
<typeStatus id="54E2880FFFA3FF992C6260F4FA60FDAC" box="[1388,1436,490,514]" pageId="18" pageNumber="77">type</typeStatus>
specimen (
<bibRefCitation id="EFC84B5CFFA3FF9928016313FE66FD88" author="Irmler" box="[271,410,525,550]" pageId="18" pageNumber="77" refString="Irmler, U. (1982) A new species of the Neotropical genus Xenopygus Bernhauer (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 36 (2), 206 - 210." type="journal article" year="1982">Irmler 1982</bibRefCitation>
).
</paragraph>
<paragraph id="8BE636ADFFA3FF9929C9632CFB75FD77" blockId="18.[151,1437,345,982]" pageId="18" pageNumber="77">
In this study, new searches for the
<taxonomicName id="4C594D2EFFA3FF992B69632CFCFBFDE7" box="[615,775,562,585]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF992B69632CFCFBFDE7" box="[615,775,562,585]" italics="true" pageId="18" pageNumber="77">X. peruvianus</emphasis>
</taxonomicName>
material were performed in Solsky collection (ZIM and ZMMU) and in IRSNB collection (Institut Royal des Sciences Naturelles de
<collectingCountry id="F34E763DFFA3FF992D0E634BFB94FDC0" box="[1024,1128,597,622]" name="Belgium" pageId="18" pageNumber="77">Belgique</collectingCountry>
, Brussels,
<collectingCountry id="F34E763DFFA3FF992DEC634BFAB4FDC0" box="[1250,1352,597,622]" name="Belgium" pageId="18" pageNumber="77">Belgium</collectingCountry>
). There are known cases where Solsky remitted material to Charles Fauvel, with deposited collection in IRSNB (see
<bibRefCitation id="EFC84B5CFFA3FF992C566364FEE2FD18" author="Caron" pageId="18" pageNumber="77" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. http: // dx. doi. org / 10.1071 / IS 10016" type="journal article" year="2012">
Caron
<emphasis id="B92DEABFFFA3FF9929996380FF29FD1B" box="[151,213,669,693]" italics="true" pageId="18" pageNumber="77">et al.</emphasis>
2012
</bibRefCitation>
). Additionally, several staphylinids researchers were consulted to try to determinate the probable location of the
<typeStatus id="54E2880FFFA3FF99283163DCFE93FD74" box="[319,367,706,730]" pageId="18" pageNumber="77">type</typeStatus>
material of
<taxonomicName id="4C594D2EFFA3FF9928F663DCFD6FFD77" box="[504,659,706,729]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF9928F663DCFD6FFD77" box="[504,659,706,729]" italics="true" pageId="18" pageNumber="77">X. peruvianus</emphasis>
</taxonomicName>
. However, there were no positive responses.
</paragraph>
<paragraph id="8BE636ADFFA3FF9929C963FBFCAFFCE8" blockId="18.[151,1437,345,982]" pageId="18" pageNumber="77">
Since there was no specimen identified as
<taxonomicName id="4C594D2EFFA3FF992BA163F8FCB9FD53" box="[687,837,742,765]" class="Insecta" family="Staphylinidae" genus="Xanthopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvanus">
<emphasis id="B92DEABFFFA3FF992BA163F8FCB9FD53" box="[687,837,742,765]" italics="true" pageId="18" pageNumber="77">X. peruvanus</emphasis>
</taxonomicName>
in the examined collections and the
<typeStatus id="54E2880FFFA3FF992DE263F8FAE0FD50" box="[1260,1308,742,766]" pageId="18" pageNumber="77">type</typeStatus>
material is considered lost since 1982 (from
<bibRefCitation id="EFC84B5CFFA3FF992B016217FD64FC8C" author="Irmler" box="[527,664,777,802]" pageId="18" pageNumber="77" refString="Irmler, U. (1982) A new species of the Neotropical genus Xenopygus Bernhauer (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 36 (2), 206 - 210." type="journal article" year="1982">Irmler 1982</bibRefCitation>
),
<taxonomicName id="4C594D2EFFA3FF992BA26214FCB4FC8F" box="[684,840,778,801]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="77" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B92DEABFFFA3FF992BA26214FCB4FC8F" box="[684,840,778,801]" italics="true" pageId="18" pageNumber="77">X. peruvianus</emphasis>
</taxonomicName>
is here considered as
<emphasis id="B92DEABFFFA3FF992D4E6214FAEEFC8F" box="[1088,1298,777,801]" italics="true" pageId="18" pageNumber="77">species inquirenda</emphasis>
, meaning, a name of doubtful identity, and in need of further investigation.
</paragraph>
<paragraph id="8BE636ADFFA3FF9929C9624FFC75FC78" blockId="18.[151,1437,345,982]" pageId="18" pageNumber="77">
The term
<emphasis id="B92DEABFFFA3FF992839624CFDF6FCC7" box="[311,522,849,873]" italics="true" pageId="18" pageNumber="77">species inquirenda</emphasis>
is defined by the ICZN (1999) glossary and substitutes the term
<emphasis id="B92DEABFFFA3FF992DFD624CFA60FCC7" box="[1267,1436,850,873]" italics="true" pageId="18" pageNumber="77">species incerta</emphasis>
cited by
<bibRefCitation id="EFC84B5CFFA3FF9929F0626BFE60FC20" author="Irmler" box="[254,412,885,910]" pageId="18" pageNumber="77" refString="Irmler, U. (1982) A new species of the Neotropical genus Xenopygus Bernhauer (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 36 (2), 206 - 210." type="journal article" year="1982">Irmler (1982)</bibRefCitation>
, and this does not have definition by the code and can generate different interpretation. Besides, Solsky, in 1872 described the species analyzing a specimen from Monte Rico,
<collectingCountry id="F34E763DFFA3FF992D626284FB5EFC1F" box="[1132,1186,922,945]" name="Peru" pageId="18" pageNumber="77">Peru</collectingCountry>
, which, is temporarily lost. In this case, the term
<emphasis id="B92DEABFFFA3FF9928B362A0FD9EFC7B" box="[445,610,957,981]" italics="true" pageId="18" pageNumber="77">nomen dubium</emphasis>
should not be considered.
</paragraph>
</subSubSection>
</treatment>
</document>