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<document id="4A2E881C6147B0EDD08C011D019B32A1" ID-CLB-Dataset="80832" ID-DOI="10.5281/zenodo.6707142" ID-GBIF-Dataset="ab66b2b7-9544-4411-bf61-5bc3651d7bca" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6707142" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1656002532852" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="03F06D13FF4020890D9C1E5E0EAAFB2E" docLanguage="en" docName="hbmw_7_Cricetidae_0204.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Calomys callosus" docType="treatment" docVersion="11" lastPageNumber="517" masterDocId="FFC9156BFFAE20670D37145C0837FFDB" masterDocTitle="Cricetidae" masterLastPageNumber="535" masterPageNumber="204" pageNumber="517" updateTime="1718799336328" updateUser="carolina">
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<mods:title id="B72CB2C4659254AC1854B1A9A65B75C0">Cricetidae</mods:title>
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<mods:namePart id="37AB8F1EE8A1FAAAC3C681BB76C47EC5">Don E. Wilson</mods:namePart>
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<mods:namePart id="5998E17C60DAFEE47667E8C1077418F9">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="3490EAF21A6731C47EE792E08DE7A72C">Thomas E. Lacher, Jr</mods:namePart>
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<treatment id="03F06D13FF4020890D9C1E5E0EAAFB2E" ID-DOI="http://doi.org/10.5281/zenodo.6727654" ID-GBIF-Taxon="196220988" ID-Zenodo-Dep="6727654" LSID="urn:lsid:plazi:treatment:03F06D13FF4020890D9C1E5E0EAAFB2E" httpUri="http://treatment.plazi.org/id/03F06D13FF4020890D9C1E5E0EAAFB2E" lastPageNumber="517" pageId="238" pageNumber="517">
<subSubSection id="C3438F8EFF4020890D9C1E5E08CBF5EB" box="[171,252,2562,2608]" pageId="238" pageNumber="517" type="multiple">
<paragraph id="8BE6DC05FF4020890D9C1E5E08CBF5EB" blockId="238.[169,1097,2562,2686]" box="[171,252,2562,2608]" pageId="238" pageNumber="517">
<heading id="D0AE6B69FF4020890D9C1E5E08CBF5EB" box="[171,252,2562,2608]" pageId="238" pageNumber="517">
<figureCitation id="1362C080FF4020890D9C1E5E08CBF5EB" box="[171,252,2562,2608]" captionStart="Plate 30: Cricetidae" captionStartId="235.[108,138,3253,3278]" captionTargetBox="[12,2727,14,3647]" captionTargetPageId="234" captionText="696. Calacas White-tailed Mouse (Calassomys apicalis), 697. Peruvian Vesper Mouse (Calomys sorella), 698. Short-tailed Vesper Mouse (Calomys miurus), 699. Coastal Vesper Mouse (Calomys chinchilico), 700. Arequipa Vesper Mouse (Calomys achaku), 701. Long-tailed Vesper Mouse (Calomys frida), 702. Andean Vesper Mouse (Calomys lepidus), 703. Drylands Vesper Mouse (Calomys musculinus), 704. Hummelincks Vesper Mouse (Calomys hummelincki), 705. Delicate Vesper Mouse (Calomys tener), 706. Small Vesper Mouse (Calomys laucha), 707. Rejected Vesper Mouse (Calomys expulsus), 708. Tocantins Vesper Mouse (Calomys tocantinsi), 709. Large Vesper Mouse (Calomys callosus), 710. Crafty Vesper Mouse (Calomys callidus), 711. Bolivian Vesper Mouse (Calomys boliviae), 712. Cordoba Vesper Mouse (Calomys venustus), 713. Cerqueiras Vesper Mouse (Calomys cerqueirai), 714. Hairy-footed Gerbil Mouse (Eligmodontia hirtipes), 715. Andean Gerbil Mouse (Eligmodontia puerulus), 716. Bolson Gerbil Mouse (Eligmodontia bolsonensis), 717. Lowland Gerbil Mouse (Eligmodontia typus), 718. Dune Gerbil Mouse (Eligmodontia dunaris), 719. Monte Gerbil Mouse (Eligmodontia moreni), 720. Morgans Gerbil Mouse (Eligmodontia morgani), 721. Pale Pericote (Graomys domorum), 722. Chaco Pericote (Graomys chacoensis), 723. Otro Cerro Pericote (Graomys edithae), 724. Common Pericote (Graomys griseoflavus)" figureDoi="http://doi.org/10.5281/zenodo.6708599" httpUri="https://zenodo.org/record/6708599/files/figure.png" pageId="238" pageNumber="517">709.</figureCitation>
</heading>
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<subSubSection id="C3438F8EFF4020890C3A1E5E0A82F5EB" box="[269,693,2562,2608]" pageId="238" pageNumber="517" type="vernacular_names">
<paragraph id="8BE6DC05FF4020890C3A1E5E0A82F5EB" blockId="238.[169,1097,2562,2686]" box="[269,693,2562,2608]" pageId="238" pageNumber="517">
<heading id="D0AE6B69FF4020890C3A1E5E0A82F5EB" box="[269,693,2562,2608]" pageId="238" pageNumber="517">
<vernacularName id="055AAC2BFF4020890C3A1E5E0A82F5EB" ID-CoL="PYL2" baseAuthorityName="Rengger" baseAuthorityYear="1830" box="[269,693,2562,2608]" class="Mammalia" family="Cricetidae" genus="Calomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="238" pageNumber="517" phylum="Chordata" rank="species" species="callosus">Large Vesper Mouse</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF4020890E361E5E0C13F5EB" box="[769,1060,2562,2608]" pageId="238" pageNumber="517" type="nomenclature">
<paragraph id="8BE6DC05FF4020890E361E5E0C13F5EB" blockId="238.[169,1097,2562,2686]" box="[769,1060,2562,2608]" pageId="238" pageNumber="517">
<heading id="D0AE6B69FF4020890E361E5E0C13F5EB" box="[769,1060,2562,2608]" pageId="238" pageNumber="517">
<taxonomicName id="4C59A786FF4020890E361E5E0C13F5EB" ID-CoL="PYL2" baseAuthorityName="Rengger" baseAuthorityYear="1830" box="[769,1060,2562,2608]" class="Mammalia" family="Cricetidae" genus="Calomys" kingdom="Animalia" order="Rodentia" pageId="238" pageNumber="517" phylum="Chordata" rank="species" species="callosus">
<emphasis id="B92D0017FF4020890E361E5E0C13F5EB" box="[769,1060,2562,2608]" italics="true" pageId="238" pageNumber="517">Calomys callosus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF4020890D9E1E1E0A23F5A5" pageId="238" pageNumber="517" type="vernacular_names">
<paragraph id="8BE6DC05FF4020890D9E1E1E0C7EF58C" blockId="238.[169,1097,2562,2686]" box="[169,1097,2626,2647]" pageId="238" pageNumber="517">
<heading id="D0AE6B69FF4020890D9E1E1E0C7EF58C" box="[169,1097,2626,2647]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF4020890D9E1E1E08C2F58C" bold="true" box="[169,245,2626,2647]" pageId="238" pageNumber="517">French:</emphasis>
<vernacularName id="055AAC2BFF4020890DC91E1E09F9F58C" ID-CoL="PYL2" baseAuthorityName="Rengger" baseAuthorityYear="1830" box="[254,462,2626,2647]" class="Mammalia" family="Cricetidae" genus="Calomys" kingdom="Animalia" language="fra" order="Rodentia" pageId="238" pageNumber="517" phylum="Chordata" rank="species" species="callosus">Calomys de Rengger</vernacularName>
/
<emphasis id="B92D0017FF4020890CD41E1E0A0AF58C" bold="true" box="[483,573,2626,2647]" pageId="238" pageNumber="517">German:</emphasis>
<vernacularName id="055AAC2BFF4020890F701E1E0B32F58C" ID-CoL="PYL2" baseAuthorityName="Rengger" baseAuthorityYear="1830" box="[583,773,2626,2647]" class="Mammalia" family="Cricetidae" genus="Calomys" kingdom="Animalia" language="deu" order="Rodentia" pageId="238" pageNumber="517" phylum="Chordata" rank="species" species="callosus">GroRRe Vespermaus</vernacularName>
/
<emphasis id="B92D0017FF4020890E2D1E1E0B42F58C" bold="true" box="[794,885,2626,2647]" pageId="238" pageNumber="517">Spanish:</emphasis>
<vernacularName id="055AAC2BFF4020890E481E1E0C7EF58C" ID-CoL="PYL2" baseAuthorityName="Rengger" baseAuthorityYear="1830" box="[895,1097,2626,2647]" class="Mammalia" family="Cricetidae" genus="Calomys" kingdom="Animalia" language="esp" order="Rodentia" pageId="238" pageNumber="517" phylum="Chordata" rank="species" species="callosus">Ratén laucha grande</vernacularName>
</heading>
</paragraph>
<paragraph id="8BE6DC05FF4020890D9E1E350A23F5A5" blockId="238.[169,1097,2562,2686]" box="[169,532,2665,2686]" pageId="238" pageNumber="517">
<heading id="D0AE6B69FF4020890D9E1E350A23F5A5" box="[169,532,2665,2686]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF4020890D9E1E350997F5A5" bold="true" box="[169,416,2665,2686]" pageId="238" pageNumber="517">Other common names:</emphasis>
<vernacularName id="055AAC2BFF4020890C9D1E350A23F5A5" ID-CoL="PYL2" baseAuthorityName="Rengger" baseAuthorityYear="1830" box="[426,532,2665,2686]" class="Mammalia" family="Cricetidae" genus="Calomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="238" pageNumber="517" phylum="Chordata" rank="species" species="callosus">Big Laucha</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF4020890E391EE90C40F3EB" pageId="238" pageNumber="517" type="reference_group">
<paragraph id="8BE6DC05FF4020890E391EE90B4AF3D3" blockId="238.[781,1373,2741,3160]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF4020890E391EE90B9EF515" bold="true" box="[782,937,2741,2766]" pageId="238" pageNumber="517">Taxonomy.</emphasis>
<taxonomicName id="4C59A786FF4020890EF71EE90D61F515" authority="Rengger, 1830" authorityName="Rengger" authorityYear="1830" box="[960,1366,2741,2766]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="238" pageNumber="517" phylum="Chordata" rank="species" species="callosus">Mus callosus Rengger, 1830</taxonomicName>
,
<materialsCitation id="3B31D658FF4020890E381E880B9FF49F" ID-GBIF-Occurrence="3818814615" pageId="238" pageNumber="517">“am Ufer des Paraguaystromes, ungefahr unter dem sieben und zwanzigsten Breitengrade.”</materialsCitation>
Interpreted by P. Hershkovitz in 1962 as “opposite mouth of Rio Bermejo, Department of Villa del Pilar, Paraguay.” Restricted byJ. R. Contreras in 1992 to vicinity of Ciudad Pilar, Corrientes, Argentina.
</paragraph>
<paragraph id="8BE6DC05FF4020890E3918530C40F3EB" blockId="238.[781,1373,2741,3160]" box="[782,1143,3087,3120]" pageId="238" pageNumber="517">This species is monotypic.</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF4020890E3A18630B99F3A6" pageId="238" pageNumber="517" type="distribution">
<caption id="DF268C8DFF4020890E3A18630B99F3A6" ID-DOI="http://doi.org/10.5281/zenodo.6708092" ID-Zenodo-Dep="6708092" httpUri="https://zenodo.org/record/6708092/files/figure.png" inLine="true" pageId="238" pageNumber="517" targetBox="[167,758,2739,3154]" targetPageId="238">
<paragraph id="8BE6DC05FF4020890E3A18630B99F3A6" blockId="238.[781,1373,2741,3160]" lastBlockId="238.[167,1372,3168,3474]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF4020890E3A18630B8AF383" bold="true" box="[781,957,3135,3160]" pageId="238" pageNumber="517">Distribution.</emphasis>
N &amp; E Bolivia, extreme W Brazil (Mato Grosso State), Paraguay, and N
<taxonomicName id="4C59A786FF4020890E2D183C0B9DF3A6" box="[794,938,3168,3197]" class="Magnoliopsida" family="Rosaceae" genus="Argentina" kingdom="Plantae" order="Rosales" pageId="238" pageNumber="517" phylum="Tracheophyta" rank="genus">Argentina</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3438F8EFF4020890D9018D80EEEFE3A" pageId="238" pageNumber="517" type="description">
<paragraph id="8BE6DC05FF4020890D9018D80EEEFE3A" blockId="238.[167,1372,3168,3474]" lastBlockId="238.[1445,2656,291,1269]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF4020890D9018D8099AF37E" bold="true" box="[167,429,3204,3237]" pageId="238" pageNumber="517">Descriptive notes.</emphasis>
Head-body 99-122 mm, tail 78-110 mm, ear 16-19 mm, hindfoot 21-23 mm; weight 30-52 g. The Large
<taxonomicName id="4C59A786FF4020890E0918F70C3AF317" authority="Mouse" authorityName="Mouse" box="[830,1037,3243,3276]" class="Magnoliopsida" family="Apiaceae" genus="Vesper" kingdom="Plantae" order="Apiales" pageId="238" pageNumber="517" phylum="Tracheophyta" rank="genus">Vesper Mouse</taxonomicName>
is medium-sized, with long and moderately lax pelage or short and appressed in wetter habitats. Upperparts of body are ocherous to tawny and often with fine mix of black or dark brown; rump and sides are generally paler; weakly defined ocherous lateral band is sometimes present; and venter is whitish to gray, sometimes washed with buff, with individual hairs dark gray at their bases. Eye-ring is defined by buffy hair, sometimes evident. Tail is bicolored, grayish brown above and whitish below. Ears are small, rounded and lightly haired; pale-colored post-auricular patches are present. Forefeet and hindfeet are white above, with whitish ungual tufts; hindfeet are narrow. Mammae count is 10-14. Diploid number of 2n = 50 is often assigned to the Large
<taxonomicName id="4C59A786FF4020890B31159C0EEEFE3A" authority="Mouse." box="[1542,1753,448,481]" class="Magnoliopsida" family="Apiaceae" genus="Vesper" kingdom="Plantae" order="Apiales" pageId="238" pageNumber="517" phylum="Tracheophyta" rank="genus">Vesper Mouse.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF402089089E15B401FEFDEB" pageId="238" pageNumber="517" type="biology_ecology">
<paragraph id="8BE6DC05FF402089089E15B401FEFDEB" blockId="238.[1445,2656,291,1269]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF402089089E15B40E2FFDD2" bold="true" box="[1449,1560,488,521]" pageId="238" pageNumber="517">Habitat.</emphasis>
Various habitats from Dry Chaco and semideciduous forests to grassy enclaves, secondary growth, old fields, and agriculture-dominated landscapes.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF402089089F16670E16FDA5" pageId="238" pageNumber="517" type="food_feeding">
<paragraph id="8BE6DC05FF402089089F16670E16FDA5" blockId="238.[1445,2656,291,1269]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF402089089F16670E8CFD83" bold="true" box="[1448,1723,571,600]" pageId="238" pageNumber="517">Food and Feeding.</emphasis>
The Large
<taxonomicName id="4C59A786FF4020890A4516670009FD83" authority="Mouse" authorityName="Mouse" box="[1906,2110,571,600]" class="Magnoliopsida" family="Apiaceae" genus="Vesper" kingdom="Plantae" order="Apiales" pageId="238" pageNumber="517" phylum="Tracheophyta" rank="genus">Vesper Mouse</taxonomicName>
is granivorous, frugivore, and insectivorous.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF402089089F16DA0E32FC98" pageId="238" pageNumber="517" type="breeding">
<paragraph id="8BE6DC05FF402089089F16DA0E32FC98" blockId="238.[1445,2656,291,1269]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF402089089F16DA0E06FD7C" bold="true" box="[1448,1585,646,679]" pageId="238" pageNumber="517">Breeding.</emphasis>
Nests of the Large
<taxonomicName id="4C59A786FF4020890A7C16DA0024FD7C" authority="Mouse" authorityName="Mouse" box="[1867,2067,646,679]" class="Magnoliopsida" family="Apiaceae" genus="Vesper" kingdom="Plantae" order="Apiales" pageId="238" pageNumber="517" phylum="Tracheophyta" rank="genus">Vesper Mouse</taxonomicName>
are generally aboveground in clumps of grass or in branches of dead trees. Nests built in laboratory were spherical, made of finely shredded, interwoven plant material; those in the field were simple depressions hollowed in the ground 10-15 cm deep. Gestation is 22 days; litters average 5 young.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF4020890891171700FFFC49" pageId="238" pageNumber="517" type="activity">
<paragraph id="8BE6DC05FF4020890891171700FFFC49" blockId="238.[1445,2656,291,1269]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF402089089117170EA3FCB7" bold="true" box="[1446,1684,843,876]" pageId="238" pageNumber="517">Activity patterns.</emphasis>
The Large
<taxonomicName id="4C59A786FF4020890A741717003DFCB7" authority="Mouse" authorityName="Mouse" box="[1859,2058,843,876]" class="Magnoliopsida" family="Apiaceae" genus="Vesper" kingdom="Plantae" order="Apiales" pageId="238" pageNumber="517" phylum="Tracheophyta" rank="genus">Vesper Mouse</taxonomicName>
is nocturnal, crepuscular, and terrestrial, although some individuals were also arboreal in captivity.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF402089089F17C10094FC3A" pageId="238" pageNumber="517" type="biology_ecology">
<paragraph id="8BE6DC05FF402089089F17C10094FC3A" blockId="238.[1445,2656,291,1269]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF402089089F17C100BBFC61" bold="true" box="[1448,2188,925,954]" pageId="238" pageNumber="517">Movements, Home range and Social organization.</emphasis>
Densities of the Large
<taxonomicName id="4C59A786FF40208904CA17C10E34FC3A" authority="Mouse" authorityName="Mouse" class="Magnoliopsida" family="Apiaceae" genus="Vesper" kingdom="Plantae" order="Apiales" pageId="238" pageNumber="517" phylum="Tracheophyta" rank="genus">Vesper Mouse</taxonomicName>
are 3—4 ind/ha; home ranges are 100-1000 m?.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF402089089F17B401F5FBD2" box="[1448,2498,1000,1033]" pageId="238" pageNumber="517" type="conservation">
<paragraph id="8BE6DC05FF402089089F17B401F5FBD2" blockId="238.[1445,2656,291,1269]" box="[1448,2498,1000,1033]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF402089089F17B40F32FBD2" bold="true" box="[1448,1797,1000,1033]" pageId="238" pageNumber="517">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF402089089F10440EAAFB2E" pageId="238" pageNumber="517" type="bibRefCitation_list">
<paragraph id="8BE6DC05FF402089089F10440EAAFB2E" blockId="238.[1445,2656,291,1269]" pageId="238" pageNumber="517">
<emphasis id="B92D0017FF402089089F10440E09FBEA" bold="true" box="[1448,1598,1048,1073]" pageId="238" pageNumber="517">Bibliography.</emphasis>
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</paragraph>
</subSubSection>
</treatment>
</document>