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<document id="935F7C2B7013CFDD3331DE0FE0B6B87D" ID-CLB-Dataset="80832" ID-DOI="10.5281/zenodo.6707142" ID-GBIF-Dataset="ab66b2b7-9544-4411-bf61-5bc3651d7bca" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6707142" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1656002532852" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="03F06D13FF2E20E70D9F18250FBAFA93" docLanguage="en" docName="hbmw_7_Cricetidae_0204.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Ichthyomys tweedii Anthony 1921" docType="treatment" docVersion="14" lastPageNumber="407" masterDocId="FFC9156BFFAE20670D37145C0837FFDB" masterDocTitle="Cricetidae" masterLastPageNumber="535" masterPageNumber="204" pageNumber="407" updateTime="1718799336328" updateUser="carolina">
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<mods:namePart id="94B49641F45FC02F1E0E2D88B32C990C">Russell A. Mittermeier</mods:namePart>
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<treatment id="03F06D13FF2E20E70D9F18250FBAFA93" ID-DOI="http://doi.org/10.5281/zenodo.6726748" ID-GBIF-Taxon="196221088" ID-Zenodo-Dep="6726748" LSID="urn:lsid:plazi:treatment:03F06D13FF2E20E70D9F18250FBAFA93" httpUri="http://treatment.plazi.org/id/03F06D13FF2E20E70D9F18250FBAFA93" lastPageNumber="407" pageId="128" pageNumber="407">
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<heading id="D0AE6B69FF2E20E70D9F182508CCF370" box="[168,251,3193,3243]" pageId="128" pageNumber="407">
<figureCitation id="1362C080FF2E20E70D9F182508CCF370" box="[168,251,3193,3243]" captionStart="Plate 19: Cricetidae" captionStartId="121.[142,172,3221,3242]" captionTargetBox="[11,2767,14,3647]" captionTargetPageId="120" captionText="321. Groove-toothed Cotton Rat (Sigmodon alstoni), 322. Ecuadorean Cotton Rat (Sigmodon inopinatus), 323. Peruvian Cotton Rat (Sigmodon peruanus), 324. White-eared Cotton Rat (Sigmodon leucotis), 325. Tawny-bellied Cotton Rat (Sigmodon fulviventer), 326. Yellow-nosed Cotton Rat (Sigmodon ochrognathus), 327. Hispid Cotton Rat (Sigmodon hispidus), 328. West Mexican Cotton Rat (Sigmodon mascotensis), 329. Arizona Cotton Rat (Sigmodon arizonae), 330. Toltec Cotton Rat (Sigmodon toltecus), 331. Allens Cotton Rat (Sigmodon alleni), 332. Burmeisters Cotton Rat (Sigmodon hirsutus), 333. Montane Cotton Rat (Sigmodon zanjonensis), 334. Miahuatlan Cotton Rat (Sigmodon planifrons), 335. Mussos Fish-eating Rat (Neusticomys mussoi), 336. Montane Fish-eating Rat (Neusticomys monticolus), 337. Vosss Fish-eating Rat (Neusticomys vossi), 338. Venezuelan Fish-eating Rat (Neusticomys venezuelae), 339. Guianan Fish-eating Rat (Neusticomys oyapocki), 340. Ferreiras Fish-eating Rat (Neusticomys ferreirai), 341. Peruvian Fish-eating Rat (Neusticomys peruviensis), 342. Northern Andean Water Mouse (Chibchanomys trichotis), 343. Orcess Andean Water Mouse (Chibchanomys orcesi), 344. Earless Water Mouse (Anotomys leander), 345. Pittiers Crab-eating Rat (Ichthyomys pittieri), 346. Common Crab-eating Rat (Ichthyomys hydrobates), 347. Tweedys Crab-eating Rat (Ichthyomys tweedii), 348. Stolzmanns Crab-eating Rat (Ichthyomys stolzmanni), 349. Goldman's Water Mouse (Rheomys raptor), 350. Thomas's Water Mouse (Rheomys thomasi), 351. Underwoods Water Mouse (Rheomys underwoodi), 352. Mexican Water Mouse (Rheomys mexicanus)" figureDoi="http://doi.org/10.5281/zenodo.6708483" httpUri="https://zenodo.org/record/6708483/files/figure.png" pageId="128" pageNumber="407">347.</figureCitation>
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<paragraph id="8BE6DC05FF2E20E70C3C18250B21F370" blockId="128.[166,1198,3193,3323]" box="[267,790,3193,3243]" pageId="128" pageNumber="407">
<heading id="D0AE6B69FF2E20E70C3C18250B21F370" box="[267,790,3193,3243]" pageId="128" pageNumber="407">
<vernacularName id="055AAC2BFF2E20E70C3C18250B21F370" ID-CoL="6N5ST" authorityName="Anthony" authorityYear="1921" box="[267,790,3193,3243]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">Tweedys Crab-eating Rat</vernacularName>
</heading>
</paragraph>
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<subSubSection id="C3438F8EFF2E20E70E6C18250CA8F370" box="[859,1183,3193,3243]" pageId="128" pageNumber="407" type="nomenclature">
<paragraph id="8BE6DC05FF2E20E70E6C18250CA8F370" blockId="128.[166,1198,3193,3323]" box="[859,1183,3193,3243]" pageId="128" pageNumber="407">
<heading id="D0AE6B69FF2E20E70E6C18250CA8F370" box="[859,1183,3193,3243]" pageId="128" pageNumber="407">
<taxonomicName id="4C59A786FF2E20E70E6C18250CA8F370" ID-CoL="6N5ST" authorityName="Anthony" authorityYear="1921" box="[859,1183,3193,3243]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">
<emphasis id="B92D0017FF2E20E70E6C18250CA8F370" box="[859,1183,3193,3243]" italics="true" pageId="128" pageNumber="407">Ichthyomys tweedii</emphasis>
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<subSubSection id="C3438F8EFF2E20E70D9F18E20ABBF321" pageId="128" pageNumber="407" type="vernacular_names">
<paragraph id="8BE6DC05FF2E20E70D9F18E20C9AF308" blockId="128.[166,1198,3193,3323]" box="[168,1197,3262,3283]" pageId="128" pageNumber="407">
<heading id="D0AE6B69FF2E20E70D9F18E20C9AF308" box="[168,1197,3262,3283]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E70D9F18E208C4F308" bold="true" box="[168,243,3262,3283]" pageId="128" pageNumber="407">French:</emphasis>
<vernacularName id="055AAC2BFF2E20E70DCA18E209E8F308" ID-CoL="6N5ST" authorityName="Anthony" authorityYear="1921" box="[253,479,3262,3283]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" language="fra" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">Ichthyomys de Tweedy</vernacularName>
/
<emphasis id="B92D0017FF2E20E70CC418E20A79F308" bold="true" box="[499,590,3262,3283]" pageId="128" pageNumber="407">German:</emphasis>
<vernacularName id="055AAC2BFF2E20E70F6418E20B11F308" ID-CoL="6N5ST" authorityName="Anthony" authorityYear="1921" box="[595,806,3262,3283]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" language="deu" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">Tweedy-Krabbenratte</vernacularName>
/
<emphasis id="B92D0017FF2E20E70E0B18E20BA0F308" bold="true" box="[828,919,3262,3283]" pageId="128" pageNumber="407">Spanish:</emphasis>
<vernacularName id="055AAC2BFF2E20E70E9618E20C9AF308" ID-CoL="6N5ST" authorityName="Anthony" authorityYear="1921" box="[929,1197,3262,3283]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" language="esp" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">Rata cangrejera de Tweedy</vernacularName>
</heading>
</paragraph>
<paragraph id="8BE6DC05FF2E20E70D9018B90ABBF321" blockId="128.[166,1198,3193,3323]" box="[167,652,3301,3322]" pageId="128" pageNumber="407">
<heading id="D0AE6B69FF2E20E70D9018B90ABBF321" box="[167,652,3301,3322]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E70D9018B909A9F321" bold="true" box="[167,414,3301,3322]" pageId="128" pageNumber="407">Other common names:</emphasis>
<vernacularName id="055AAC2BFF2E20E70C9318B90ABBF321" ID-CoL="6N5ST" authorityName="Anthony" authorityYear="1921" box="[420,652,3301,3322]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">Tweedy's Ichthyomyine</vernacularName>
</heading>
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<subSubSection id="C3438F8EFF2E20E70D9019700A3EF2AB" pageId="128" pageNumber="407" type="reference_group">
<paragraph id="8BE6DC05FF2E20E70D9019700A3EF2AB" blockId="128.[165,1372,3372,3486]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E70D9019700975F292" bold="true" box="[167,322,3372,3401]" pageId="128" pageNumber="407">Taxonomy.</emphasis>
<taxonomicName id="4C59A786FF2E20E70C6719700B26F292" ID-CoL="6N5ST" authority="Anthony, 1921" authorityName="Anthony" authorityYear="1921" box="[336,785,3372,3401]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">Ichthyomys tweedii Anthony, 1921</taxonomicName>
,
<materialsCitation id="3B31D658FF2E20E70E1319700A3EF2AB" ID-GBIF-Occurrence="3818814379" pageId="128" pageNumber="407">
“Portovelo, Prov. del Oro,
<collectingCountry id="F34E9C95FF2E20E709A819700D21F292" box="[1183,1302,3372,3401]" name="Ecuador" pageId="128" pageNumber="407">Ecuador</collectingCountry>
; alttude,
<quantity id="4CA171E0FF2E20E70DCE19130969F2AB" box="[249,350,3407,3440]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="128" pageNumber="407" unit="ft" value="2000.0">2000 ft.</quantity>
[=
<quantity id="4CA171E0FF2E20E70CA5191309DBF2AB" box="[402,492,3407,3440]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="128" pageNumber="407" unit="m" value="610.0">610 m</quantity>
].”
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</paragraph>
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<subSubSection id="C3438F8EFF2E20E70D9F19270D3CF243" box="[168,1291,3451,3480]" pageId="128" pageNumber="407" type="discussion">
<paragraph id="8BE6DC05FF2E20E70D9F19270D3CF243" blockId="128.[165,1372,3372,3486]" box="[168,1291,3451,3480]" pageId="128" pageNumber="407">
It includes
<taxonomicName id="4C59A786FF2E20E70C72192709E5F243" box="[325,466,3451,3480]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="caurinus">I. caurinus</taxonomicName>
which is treated as junior synonym of
<taxonomicName id="4C59A786FF2E20E70EDA19270C68F243" authorityName="Anthony" authorityYear="1921" box="[1005,1119,3451,3480]" class="Mammalia" family="Cricetidae" genus="Ichthyomys" kingdom="Animalia" order="Rodentia" pageId="128" pageNumber="407" phylum="Chordata" rank="species" species="tweedii">I. tweedii</taxonomicName>
. Monotypic.
</paragraph>
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<subSubSection id="C3438F8EFF2E20E705321579018AFEB6" pageId="128" pageNumber="407" type="distribution">
<caption id="DF268C8DFF2E20E705321579018AFEB6" ID-DOI="http://doi.org/10.5281/zenodo.6761198" ID-Zenodo-Dep="6761198" httpUri="https://zenodo.org/record/6761198/files/figure.png" inLine="true" pageId="128" pageNumber="407" startId="128.[2053,2225,293,326]" targetBox="[1438,2029,301,715]" targetPageId="128">
<paragraph id="8BE6DC05FF2E20E705321579018AFEB6" blockId="128.[2052,2646,293,720]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E7053215790082FE9D" bold="true" box="[2053,2229,293,326]" pageId="128" pageNumber="407">Distribution.</emphasis>
C Panama and lowlands and Andean foothills of W Ecuador.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3438F8EFF2E20E7053215280E94FC9C" pageId="128" pageNumber="407" type="description">
<paragraph id="8BE6DC05FF2E20E7053215280E94FC9C" blockId="128.[2052,2646,293,720]" lastBlockId="128.[1439,2649,726,1352]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E705321528012AFE4E" bold="true" box="[2053,2333,372,405]" pageId="128" pageNumber="407">Descriptive notes.</emphasis>
Head-body 143-197 mm, tail 132-155 mm, ear 8-12 mm, hindfoot 33-40 mm; weight 123-155 g. Large ichthyomyine, with dark unicolored tail, bilobed M,, and massive skull featuring broad rostra and flaring zygomatic arches. Dorsal pelage is dense and woollylooking. Back is dark brown; flanks are paler; and venter is white to yellowish gray, strongly contrasting flanks and back. Tail is uniformly dark brown, equal or somewhat smaller than head-body length. Outer edges of feet and fingers have fringes of rigid silver-to-white hair.
</paragraph>
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<subSubSection id="C3438F8EFF2E20E7089717110EA0FC4D" pageId="128" pageNumber="407" type="biology_ecology">
<paragraph id="8BE6DC05FF2E20E7089717110EA0FC4D" blockId="128.[1439,2649,726,1352]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E7089717110E27FCB5" bold="true" box="[1440,1552,845,878]" pageId="128" pageNumber="407">Habitat.</emphasis>
Small streams bordered by dense secondary vegetation from sea levelto elevations of ¢.1350 m.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2E20E7089717C70231FC67" box="[1440,2566,923,956]" pageId="128" pageNumber="407" type="food_feeding">
<paragraph id="8BE6DC05FF2E20E7089717C70231FC67" blockId="128.[1439,2649,726,1352]" box="[1440,2566,923,956]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E7089717C70E91FC67" bold="true" box="[1440,1702,923,956]" pageId="128" pageNumber="407">Food and Feeding.</emphasis>
Tweedys Crab-eating Rat eats fish, crabs, and aquatic insects.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2E20E70896179F0FBEFBD7" pageId="128" pageNumber="407" type="breeding">
<paragraph id="8BE6DC05FF2E20E70896179F0FBEFBD7" blockId="128.[1439,2649,726,1352]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E70896179F0E10FC3F" bold="true" box="[1441,1575,963,996]" pageId="128" pageNumber="407">Breeding.</emphasis>
A Tweedys Crab-eating Rat collected during late wet season (May) had two newly implanted embryos in utero.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2E20E708A8104E0E3BFB80" pageId="128" pageNumber="407" type="activity">
<paragraph id="8BE6DC05FF2E20E708A8104E0E3BFB80" blockId="128.[1439,2649,726,1352]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E708A8104E0EBAFBE8" bold="true" box="[1439,1677,1042,1075]" pageId="128" pageNumber="407">Activity patterns.</emphasis>
Tweedys Crab-eating Rat is both diurnal and nocturnal, and semiaquatic.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2E20E70896103C0264FB5A" box="[1441,2643,1120,1153]" pageId="128" pageNumber="407" type="biology_ecology">
<paragraph id="8BE6DC05FF2E20E70896103C0264FB5A" blockId="128.[1439,2649,726,1352]" box="[1441,2643,1120,1153]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E70896103C007FFB5A" bold="true" box="[1441,2120,1120,1153]" pageId="128" pageNumber="407">Movements, Home range and Social organization.</emphasis>
Tweedys Crab-eating Rats are solitary.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2E20E7089610D40274FB23" pageId="128" pageNumber="407" type="conservation">
<paragraph id="8BE6DC05FF2E20E7089610D40274FB23" blockId="128.[1439,2649,726,1352]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E7089610D40F37FB72" bold="true" box="[1441,1792,1160,1193]" pageId="128" pageNumber="407">Status and Conservation.</emphasis>
Classified as Data Deficient on The IUCN Red List. Extent of occurrence, status, and ecological requirements of Tweedys Crab-eating Rat are unknown. It could more widely distributed because it is known from two disjunct areas.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2E20E70895115B0FBAFA93" pageId="128" pageNumber="407" type="bibRefCitation_list">
<paragraph id="8BE6DC05FF2E20E70895115B0FBAFA93" blockId="128.[1439,2649,726,1352]" pageId="128" pageNumber="407">
<emphasis id="B92D0017FF2E20E70895115B0E0CFAFB" bold="true" box="[1442,1595,1287,1312]" pageId="128" pageNumber="407">Bibliography.</emphasis>
Anthony (1921), Santillan &amp; Segovia (2013), Thomas (1924b), Tirira, Boada, Samudio &amp; Pino (2008), Voss (1988, 2015b), Voss et al. (1982).
</paragraph>
</subSubSection>
</treatment>
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