treatments-xml/data/03/80/9F/03809F40FF9BD70DD1C2FA59FC7C2763.xml
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<document id="B6A101791903AACB9B59E06E7478C076" ID-DOI="10.11646/zootaxa.5306.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8073166" ID-ZooBank="F136C7C2-BF12-4943-BA48-4AFEF4186F1A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1687503661512" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2023" docId="03809F40FF9BD70DD1C2FA59FC7C2763" docLanguage="en" docName="zootaxa.5306.5.1.pdf" docOrigin="Zootaxa 5306 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5306.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Platymetopus rugosus" docType="treatment" docVersion="2" lastPageNumber="526" masterDocId="FFB9E738FF8CD714D155FFFFFF832116" masterDocTitle="A review of Asian species of the genus Platymetopus (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="536" masterPageNumber="501" pageNumber="524" updateTime="1687504211159" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2D61C023FFE5AFBC448482F96383C8C7">A review of Asian species of the genus Platymetopus (Coleoptera: Carabidae: Harpalini)</mods:title>
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<mods:namePart id="E5EC40F5CB71D0283EC2383A47034B55">Kataev, Boris M.</mods:namePart>
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<mods:date id="2DCE4E7506F93B4757100F9E024A093E">2023</mods:date>
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<treatment id="03809F40FF9BD70DD1C2FA59FC7C2763" LSID="urn:lsid:plazi:treatment:03809F40FF9BD70DD1C2FA59FC7C2763" httpUri="http://treatment.plazi.org/id/03809F40FF9BD70DD1C2FA59FC7C2763" lastPageId="25" lastPageNumber="526" pageId="23" pageNumber="524">
<subSubSection id="C3337DDDFF9BD703D1C2FA59FDE224D6" box="[151,609,1446,1473]" pageId="23" pageNumber="524" type="nomenclature">
<paragraph id="8B962E56FF9BD703D1C2FA59FDE224D6" blockId="23.[151,609,1446,1509]" box="[151,609,1446,1473]" pageId="23" pageNumber="524">
<heading id="D0DE993AFF9BD703D1C2FA59FDE224D6" bold="true" box="[151,609,1446,1473]" fontSize="11" level="1" pageId="23" pageNumber="524" reason="1">
<taxonomicName id="4C2955D5FF9BD703D1C2FA59FDE224D6" ID-CoL="4JSXV" authority="(Nietner, 1857)" baseAuthorityName="Nietner" baseAuthorityYear="1857" box="[151,609,1446,1473]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B95DF244FF9BD703D1C2FA59FDE224D6" bold="true" box="[151,609,1446,1473]" pageId="23" pageNumber="524">
<emphasis id="B95DF244FF9BD703D1C2FA59FE2224D7" bold="true" box="[151,417,1446,1473]" italics="true" pageId="23" pageNumber="524">Platymetopus rugosus</emphasis>
(
<bibRefCitation id="EFB853A7FF9BD703D0E4FA59FDDA24D6" author="Nietner, J." box="[433,601,1446,1472]" pageId="23" pageNumber="524" pagination="272 - 282" refId="ref27223" refString="Nietner, J. (1857) Descriptions of new Ceylon Coleoptera. The Annals and Magazine of Natural History, Series 2, 20, 272 - 282 + 368 - 374. https: // doi. org / 10.1080 / 00222935709487938" type="journal article" year="1857">Nietner, 1857</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3337DDDFF9BD703D1C2FA34FD8324F3" box="[151,512,1482,1509]" pageId="23" pageNumber="524" type="description">
<paragraph id="8B962E56FF9BD703D1C2FA34FD8324F3" blockId="23.[151,609,1446,1509]" box="[151,512,1482,1509]" pageId="23" pageNumber="524">
(
<figureCitation id="131232D3FF9BD703D1CAFA34FEAB24F3" box="[159,296,1482,1509]" captionStart="FIGURES 4245" captionStartId="22.[151,264,1128,1152]" captionTargetBox="[161,1426,278,1095]" captionTargetId="figure-72@22.[154,1433,273,1105]" captionTargetPageId="22" captionText="FIGURES 4245. Platymetopus spp., habitus (42, 43, 45) and humerus (44). 42, P. tritus Bates, 1889 (Vietnam); 43, P. rugosus rugosus (Nietner, 1857) (India); 44, 45, P. r. obscurofemoratus subsp. n. (Vietnam, holotype). Scale bars = 1.0 mm (A: 44, B: 42, 43, 45)." figureDoi="http://doi.org/10.5281/zenodo.8073200" httpUri="https://zenodo.org/record/8073200/files/figure.png" pageId="23" pageNumber="524">Figs. 4345</figureCitation>
,
<figureCitation id="131232D3FF9BD703D066FA35FEF824F3" box="[307,379,1482,1509]" captionStart="FIGURES 4656" captionStartId="24.[151,264,1416,1440]" captionTargetBox="[153,1434,183,1391]" captionTargetId="figure-27@24.[151,1436,181,1393]" captionTargetPageId="24" captionText="FIGURES 4656. Platymetopus spp., median lobe of aedeagus. 46, 47, P. tritus Bates, 1889 (Vietnam); 48, 49, P. rugosus rugosus (Nietner, 1857) (India); 50, 51, P. r. obscurofemoratus subsp. n. (Vietnam, holotype); 52, 53, P. quadrimaculatus Dejean, 1829 (Vietnam); 5456, P. figuratus pictus Andrewes, 1923 (India, Rajasthan). Dorsal (46, 48, 50, 52, 54) and lateral (47, 49, 51, 53, 55, 56) views. Scale bars = 1.0 mm (A: 4653, B: 5456)." figureDoi="http://doi.org/10.5281/zenodo.8073202" httpUri="https://zenodo.org/record/8073202/files/figure.png" pageId="23" pageNumber="524">4851</figureCitation>
,
<figureCitation id="131232D3FF9BD703D0D2FA35FE4C24F3" box="[391,463,1482,1509]" captionStart="FIGURES 5764" captionStartId="26.[151,264,972,996]" captionTargetBox="[156,1430,456,942]" captionTargetId="figure-124@26.[151,1436,453,949]" captionTargetPageId="26" captionText="FIGURES 5764. Platymetopus spp., gonocoxite, lateral view (57, 59, 61, 63), and laterotergite, gonosubcoxite, and gonocoxite, ventral view (58, 60, 62, 64). 57, 58, P. tritus Bates, 1889 (Vietnam); 59, 60, P. rugosus (Nietner, 1857) (India); 61, 62, P. quadrimaculatus Dejean, 1829 (Vietnam); 63, 64, P. figuratus pictus Andrewes, 1923 (India, Rajasthan). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8073206" httpUri="https://zenodo.org/record/8073206/files/figure.png" pageId="23" pageNumber="524">59, 60</figureCitation>
,
<figureCitation id="131232D3FF9BD703D08EFA34FE7B24F3" box="[475,504,1483,1509]" captionStart="FIGURE 66" captionStartId="29.[151,250,1166,1190]" captionTargetBox="[151,1435,345,1142]" captionTargetId="figure-87@29.[151,1435,345,1142]" captionTargetPageId="29" captionText="FIGURE 66. Platymetopus rugosus (Nietner, 1857), distribution: P. r. rugosus (Nietner, 1857) (black circles, based on examined material; white circles, based on literature data); P. r. obscurofemoratus subsp. n. (stars)." figureDoi="http://doi.org/10.5281/zenodo.8073210" httpUri="https://zenodo.org/record/8073210/files/figure.png" pageId="23" pageNumber="524">66</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3337DDDFF9BD703D1C2F9F0FBCE2731" box="[151,1101,1551,1576]" pageId="23" pageNumber="524" type="reference_group">
<paragraph id="8B962E56FF9BD703D1C2F9F0FBCE2731" blockId="23.[151,1101,1551,1576]" box="[151,1101,1551,1576]" pageId="23" pageNumber="524">
<treatmentCitationGroup id="AB390978FF9BD703D1C2F9F0FBCE2731" box="[151,1101,1551,1576]" pageId="23" pageNumber="524">
<treatmentCitation id="0A880847FF9BD703D1C2F9F0FD042731" author="Nietner, J." box="[151,647,1551,1575]" page="373" pageId="23" pageNumber="524" year="1857">
<taxonomicName id="4C2955D5FF9BD703D1C2F9F0FD042731" ID-CoL="8M9NR" authority="Nietner, 1857: 373" authorityName="Nietner" authorityPageNumber="373" authorityYear="1857" box="[151,647,1551,1575]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="rugosus" subGenus="Ophonus">
<emphasis id="B95DF244FF9BD703D1C2F9F0FF742731" box="[151,247,1551,1575]" italics="true" pageId="23" pageNumber="524">Harpalus</emphasis>
(
<emphasis id="B95DF244FF9BD703D053F9F0FEE22731" box="[262,353,1551,1575]" italics="true" pageId="23" pageNumber="524">Ophonus</emphasis>
)
<emphasis id="B95DF244FF9BD703D025F9EFFE432731" box="[368,448,1552,1575]" italics="true" pageId="23" pageNumber="524">rugosus</emphasis>
<materialsCitation id="3B41240BFF9BD703D092F9F0FD042731" box="[455,647,1551,1575]" country="Sri Lanka" location="Sri Lanka" pageId="23" pageNumber="524" specimenCount="1">
<bibRefCitation id="EFB853A7FF9BD703D092F9F0FD042731" author="Nietner, J." box="[455,647,1551,1575]" pageId="23" pageNumber="524" pagination="272 - 282" refId="ref27223" refString="Nietner, J. (1857) Descriptions of new Ceylon Coleoptera. The Annals and Magazine of Natural History, Series 2, 20, 272 - 282 + 368 - 374. https: // doi. org / 10.1080 / 00222935709487938" type="journal article" year="1857">Nietner, 1857: 373</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="549290F4FF9BD703D3C5F9EFFD41273E" box="[656,706,1552,1576]" pageId="23" pageNumber="524">Type</typeStatus>
locality: “about Colombo”,
<collectingCountry id="F33E6EC6FF9BD703D2B7F9F0FBCA2731" box="[994,1097,1551,1575]" name="Sri Lanka" pageId="23" pageNumber="524">Sri Lanka</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3337DDDFF9BD703D1C2F9B0FBE32657" pageId="23" pageNumber="524" type="materials_examined">
<paragraph id="8B962E56FF9BD703D1C2F9B0FBE32657" blockId="23.[151,1437,1614,2037]" pageId="23" pageNumber="524">
<emphasis id="B95DF244FF9BD703D1C2F9B0FE3D277E" bold="true" box="[151,446,1614,1641]" pageId="23" pageNumber="524">Type material examined.</emphasis>
<materialsCitation id="3B41240BFF9BD703D091F9B0FABD279B" collectionCode="MFNB" country="Sri Lanka" location="Nietn." municipality="Hist." pageId="23" pageNumber="524" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<typeStatus id="549290F4FF9BD703D091F9B0FDAC277F" box="[452,559,1615,1641]" pageId="23" pageNumber="524">Syntypes</typeStatus>
:
<specimenCount id="9D2FE5DFFF9BD703D36CF9B0FD1F277F" box="[569,668,1615,1641]" pageId="23" pageNumber="524" type="female">1 female</specimenCount>
, with labels “
<collectingCountry id="F33E6EC6FF9BD703D264F9B0FC04277F" box="[817,903,1615,1641]" name="Sri Lanka" pageId="23" pageNumber="524">Ceylon</collectingCountry>
,
<location id="8EF6788DFF9BD703D2C7F9B0FC5F277F" LSID="urn:lsid:plazi:treatment:03809F40FF9BD70DD1C2FA59FC7C2763:8EF6788DFF9BD703D2C7F9B0FC5F277F" box="[914,988,1615,1641]" country="Sri Lanka" municipality="Hist." name="Nietn." pageId="23" pageNumber="524">Nietn.</location>
”, “3491”, “
<taxonomicName id="4C2955D5FF9BD703D50CF9B0FA0E277F" authority="Ntn." authorityName="Ntn." box="[1113,1421,1615,1641]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="rugosus">Platymetopus rugosus Ntn.</taxonomicName>
”, and “
<collectingMunicipality id="6BF2B42CFF9BD703D187F98CFE89279B" box="[210,266,1651,1677]" pageId="23" pageNumber="524">Hist.</collectingMunicipality>
-Coll. (
<taxonomicName id="4C2955D5FF9BD703D00DF98CFE5A279B" authorityName="Linnaeus" authorityYear="1758" box="[344,473,1651,1677]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="23" pageNumber="524" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
), Nr. 3491
<taxonomicName id="4C2955D5FF9BD703D303F98CFCB2279B" box="[598,817,1651,1677]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="undetermined">Platymetopus spec.</taxonomicName>
,
<collectingCountry id="F33E6EC6FF9BD703D268F98CFC0C279B" box="[829,911,1651,1677]" name="Sri Lanka" pageId="23" pageNumber="524">Ceylan</collectingCountry>
, Nietner, Zool. Mus. Berlin” (
<collectionCode id="ED38B693FF9BD703D5B5F98CFAB5279B" box="[1248,1334,1651,1677]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="23" pageNumber="524">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B41240BFF9BD703D41FF98CFE1A27C3" collectionCode="MFNB" country="Sri Lanka" location="Hist. - Coll." pageId="23" pageNumber="524" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D2FE5DFFF9BD703D41FF98CFA1B279B" box="[1354,1432,1651,1677]" pageId="23" pageNumber="524" type="male">1 male</specimenCount>
,
<specimenCount id="9D2FE5DFFF9BD703D1C2F968FF7E27A7" box="[151,253,1687,1713]" pageId="23" pageNumber="524" type="female">1 female</specimenCount>
, with labels “3491”, and “
<location id="8EF6788DFF9BD703D37EF968FD2027A7" LSID="urn:lsid:plazi:treatment:03809F40FF9BD70DD1C2FA59FC7C2763:8EF6788DFF9BD703D37EF968FD2027A7" box="[555,675,1687,1713]" country="Sri Lanka" name="Hist. - Coll." pageId="23" pageNumber="524">Hist.-Coll.</location>
(
<taxonomicName id="4C2955D5FF9BD703D3E6F968FCB727A7" authorityName="Linnaeus" authorityYear="1758" box="[691,820,1687,1713]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="23" pageNumber="524" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
), Nr. 3491
<taxonomicName id="4C2955D5FF9BD703D2E0F968FB1127A7" box="[949,1170,1687,1713]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="undetermined">Platymetopus spec.</taxonomicName>
,
<collectingCountry id="F33E6EC6FF9BD703D5CAF968FB7227A7" box="[1183,1265,1687,1713]" name="Sri Lanka" pageId="23" pageNumber="524">Ceylan</collectingCountry>
, Nietner, Zool. Mus. Berlin” (
<collectionCode id="ED38B693FF9BD703D06EF944FE1227C3" box="[315,401,1723,1749]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="23" pageNumber="524">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B41240BFF9BD703D0F3F944FABC27EF" collectionCode="MFNB" country="Sri Lanka" location="Mus. Colombo" municipality="Nietner" pageId="23" pageNumber="524" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<specimenCount id="9D2FE5DFFF9BD703D0F3F944FD8B27C3" box="[422,520,1723,1749]" pageId="23" pageNumber="524" type="female">1 female</specimenCount>
(?), with labels “
<collectingCountry id="F33E6EC6FF9BD703D39BF944FCA727C3" box="[718,804,1723,1749]" name="Sri Lanka" pageId="23" pageNumber="524">Ceylon</collectingCountry>
,
<location id="8EF6788DFF9BD703D264F944FC5C27C3" LSID="urn:lsid:plazi:treatment:03809F40FF9BD70DD1C2FA59FC7C2763:8EF6788DFF9BD703D264F944FC5C27C3" box="[817,991,1723,1749]" country="Sri Lanka" municipality="Nietner" name="Mus. Colombo" pageId="23" pageNumber="524">Mus. Colombo</location>
”, “61932”, “
<taxonomicName id="4C2955D5FF9BD703D53BF944FB5A27C3" authorityName="MacLeay" authorityYear="1825" box="[1134,1241,1723,1749]" class="Insecta" family="Carabidae" genus="Dioryche" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="genus">Dioryche</taxonomicName>
rugosa
<collectingMunicipality id="6BF2B42CFF9BD703D460F944FA0D27C3" box="[1333,1422,1723,1749]" pageId="23" pageNumber="524">Nietner</collectingMunicipality>
”, and “Hist.-Coll. (
<taxonomicName id="4C2955D5FF9BD703D002F920FE5B27EF" authorityName="Linnaeus" authorityYear="1758" box="[343,472,1759,1785]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="23" pageNumber="524" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
), Nr. 61932,
<taxonomicName id="4C2955D5FF9BD703D33DF920FD5327EF" authorityName="MacLeay" authorityYear="1825" box="[616,720,1759,1785]" class="Insecta" family="Carabidae" genus="Dioryche" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="genus">Dioryche</taxonomicName>
rugosa Nietner, Colombo, Zool. Mus. Berlin” (
<collectionCode id="ED38B693FF9BD703D5B7F920FABB27EF" box="[1250,1336,1759,1785]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="23" pageNumber="524">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B41240BFF9BD703D41FF920FBDF2657" collectionCode="MIZW" collectorName="Inst. Zool. P. A. N. Warszawa" country="Sri Lanka" location="Nietner" municipality="Nietn." pageId="23" pageNumber="524" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D2FE5DFFF9BD703D41FF920FA1B27EF" box="[1354,1432,1759,1785]" pageId="23" pageNumber="524" type="male">1 male</specimenCount>
, with labels “
<collectingCountry id="F33E6EC6FF9BD703D077F8FCFEFB260B" box="[290,376,1795,1821]" name="Sri Lanka" pageId="23" pageNumber="524">Ceylon</collectingCountry>
,
<location id="8EF6788DFF9BD703D0D6F8FCFE54260B" LSID="urn:lsid:plazi:treatment:03809F40FF9BD70DD1C2FA59FC7C2763:8EF6788DFF9BD703D0D6F8FCFE54260B" box="[387,471,1795,1821]" country="Sri Lanka" municipality="Nietn." name="Nietner" pageId="23" pageNumber="524">Nietner</location>
s”, “
<taxonomicName id="4C2955D5FF9BD703D358F8FCFDFB260B" authorityName="MacLeay" authorityYear="1825" box="[525,632,1795,1821]" class="Insecta" family="Carabidae" genus="Dioryche" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="genus">Dioryche</taxonomicName>
rugosa
<collectingMunicipality id="6BF2B42CFF9BD703D385F8FCFC95260B" box="[720,790,1795,1821]" pageId="23" pageNumber="524">Nietn.</collectingMunicipality>
x”, “128”, “
<taxonomicName id="4C2955D5FF9BD703D2CAF8FCFA1E260B" authority="Nietn., H. E. Andrewes" authorityName="Nietn., H. E. Andrewes" box="[927,1437,1795,1821]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="rugosus">Platymetopus rugosus Nietn., H.E. Andrewes</taxonomicName>
det.”, “
<collectorName id="26DC4B80FF9BD703D1B3F8D8FDAB2657" box="[230,552,1831,1857]" pageId="23" pageNumber="524">Inst. Zool. P.A.N. Warszawa</collectorName>
, Cotypus, Nr. 1429”, and “MIZ 237162” (
<collectionCode id="ED38B693FF9BD703D555F8D8FBD72657" box="[1024,1108,1831,1857]" pageId="23" pageNumber="524">MIZW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3337DDDFF9BD70DD192F8B5FB3924D7" lastPageId="25" lastPageNumber="526" pageId="23" pageNumber="524" type="description">
<paragraph id="8B962E56FF9BD703D192F8B5FAF92673" blockId="23.[151,1437,1614,2037]" box="[199,1402,1866,1893]" pageId="23" pageNumber="524">
<emphasis id="B95DF244FF9BD703D192F8B5FEDA2672" bold="true" box="[199,345,1866,1892]" pageId="23" pageNumber="524">Description.</emphasis>
Body length: 6.27.2 (
<emphasis id="B95DF244FF9BD703D309F8B4FDF32672" box="[604,624,1867,1892]" italics="true" pageId="23" pageNumber="524">m</emphasis>
6.5) mm. Body proportions in
<tableCitation id="C6AB1BEDFF9BD703D298F8B4FBA32673" box="[973,1056,1867,1893]" captionStart="TABLE 1" captionStartId="4.[152,244,152,176]" captionText="TABLE 1. Variation of body length and proportions in species of Platymetopus Dejean, 1829." pageId="23" pageNumber="524">Table 1</tableCitation>
. Habitus as in
<figureCitation id="131232D3FF9BD703D592F8B4FAF52673" box="[1223,1398,1866,1893]" captionStart="FIGURES 4245" captionStartId="22.[151,264,1128,1152]" captionTargetBox="[161,1426,278,1095]" captionTargetId="figure-72@22.[154,1433,273,1105]" captionTargetPageId="22" captionText="FIGURES 4245. Platymetopus spp., habitus (42, 43, 45) and humerus (44). 42, P. tritus Bates, 1889 (Vietnam); 43, P. rugosus rugosus (Nietner, 1857) (India); 44, 45, P. r. obscurofemoratus subsp. n. (Vietnam, holotype). Scale bars = 1.0 mm (A: 44, B: 42, 43, 45)." figureDoi="http://doi.org/10.5281/zenodo.8073200" httpUri="https://zenodo.org/record/8073200/files/figure.png" pageId="23" pageNumber="524">Figs. 43 and 45</figureCitation>
.
</paragraph>
<paragraph id="8B962E56FF9BD703D192F890FE3426E3" blockId="23.[151,1437,1614,2037]" pageId="23" pageNumber="524">Body dark brown to black, with unicolorous elytra; dorsum mat or slightly shiny, without metallic tinge. Palpi and antennae dark, black or almost black; antennomeres 1 and 2 occasionally slightly paler partially, mostly basally, brown or dark brown. Femora yellow or more or less infuscate; tibiae yellow or brownish yellow, infuscate at apex; tarsi dark brown to black.</paragraph>
<caption id="DF567EDEFF94D70CD1C2FA77FC06271B" ID-DOI="http://doi.org/10.5281/zenodo.8073202" ID-Zenodo-Dep="8073202" httpUri="https://zenodo.org/record/8073202/files/figure.png" pageId="24" pageNumber="525" startId="24.[151,264,1416,1440]" targetBox="[153,1434,183,1391]" targetPageId="24" targetType="figure">
<paragraph id="8B962E56FF94D70CD1C2FA77FC06271B" blockId="24.[151,1437,1416,1549]" pageId="24" pageNumber="525">
<emphasis id="B95DF244FF94D70CD1C2FA77FED424B7" bold="true" box="[151,343,1416,1441]" pageId="24" pageNumber="525">FIGURES 4656.</emphasis>
<taxonomicName id="4C2955D5FF94D70CD00AFA76FD9D24B6" box="[351,542,1417,1441]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95DF244FF94D70CD00AFA76FE6A24B7" box="[351,489,1417,1441]" italics="true" pageId="24" pageNumber="525">Platymetopus</emphasis>
spp.
</taxonomicName>
, median lobe of aedeagus. 46, 47,
<taxonomicName id="4C2955D5FF94D70CD2D8FA76FBE224B7" authority="Bates, 1889" authorityName="Bates" authorityYear="1889" box="[909,1121,1417,1441]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="tritus">
<emphasis id="B95DF244FF94D70CD2D8FA76FC5D24B6" box="[909,990,1417,1440]" italics="true" pageId="24" pageNumber="525">P. tritus</emphasis>
<bibRefCitation id="EFB853A7FF94D70CD2B3FA76FBE224B7" author="Bates, H. W." box="[998,1121,1417,1441]" pageId="24" pageNumber="525" pagination="261 - 286" refId="ref25884" refString="Bates, H. W. (1889) Contributions a la faune indo-chinoise. 3 e memoire. Annales de la Societe Entomologique de France, Series 6, 9, 261 - 286." type="journal article" year="1889">Bates, 1889</bibRefCitation>
</taxonomicName>
(Vietnam); 48, 49,
<taxonomicName id="4C2955D5FF94D70CD464FA76FE0C24D3" authority="(Nietner, 1857)" baseAuthorityName="Nietner" baseAuthorityYear="1857" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="subSpecies" species="rugosus" subSpecies="rugosus">
<emphasis id="B95DF244FF94D70CD464FA76FF6424D2" italics="true" pageId="24" pageNumber="525">P. rugosus rugosus</emphasis>
(
<bibRefCitation id="EFB853A7FF94D70CD1ADFA52FE0A24D3" author="Nietner, J." box="[248,393,1453,1477]" pageId="24" pageNumber="525" pagination="272 - 282" refId="ref27223" refString="Nietner, J. (1857) Descriptions of new Ceylon Coleoptera. The Annals and Magazine of Natural History, Series 2, 20, 272 - 282 + 368 - 374. https: // doi. org / 10.1080 / 00222935709487938" type="journal article" year="1857">Nietner, 1857</bibRefCitation>
)
</taxonomicName>
(India); 50, 51,
<taxonomicName id="4C2955D5FF94D70CD317FA52FCA824D3" authority="Kataev, 2023" authorityName="Kataev" authorityYear="2023" box="[578,811,1453,1477]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="subSpecies" species="rugosus" status="subsp. n." subSpecies="obscurofemoratus">
<emphasis id="B95DF244FF94D70CD317FA52FCA824D3" box="[578,811,1453,1477]" italics="true" pageId="24" pageNumber="525">P. r. obscurofemoratus</emphasis>
</taxonomicName>
<emphasis id="B95DF244FF94D70CD260FA52FC1B24D3" bold="true" box="[821,920,1453,1477]" pageId="24" pageNumber="525">
<taxonomicNameLabel id="A26E4F3FFF94D70CD260FA52FC1B24D3" box="[821,920,1453,1477]" pageId="24" pageNumber="525" rank="subSpecies">subsp. n.</taxonomicNameLabel>
</emphasis>
(Vietnam, holotype); 52, 53,
<taxonomicName id="4C2955D5FF94D70CD581FA52FE9C24FF" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="B95DF244FF94D70CD581FA52FA1E24D3" box="[1236,1437,1453,1477]" italics="true" pageId="24" pageNumber="525">P. quadrimaculatus</emphasis>
<bibRefCitation id="EFB853A7FF94D70CD1C2FA2EFE9C24FF" author="Dejean, P. F. M. A." box="[151,287,1489,1513]" pageId="24" pageNumber="525" refId="ref26072" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829</bibRefCitation>
</taxonomicName>
(Vietnam); 5456,
<taxonomicName id="4C2955D5FF94D70CD0BDFA2EFCCD24FF" authority="Andrewes, 1923" authorityName="Andrewes" authorityYear="1923" box="[488,846,1489,1513]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="subSpecies" species="figuratus" subSpecies="pictus">
<emphasis id="B95DF244FF94D70CD0BDFA2EFD2124FE" box="[488,674,1489,1513]" italics="true" pageId="24" pageNumber="525">P. figuratus pictus</emphasis>
<bibRefCitation id="EFB853A7FF94D70CD3F2FA2EFCCD24FF" author="Andrewes, H. E." box="[679,846,1489,1513]" pageId="24" pageNumber="525" pagination="223 - 251" refId="ref25271" refString="Andrewes, H. E. (1923) Descriptions of some new Carabidae from Ceylon. (Part I.). Spolia Zeylanica, 12 (46), 223 - 251." type="journal article" year="1923">Andrewes, 1923</bibRefCitation>
</taxonomicName>
(India, Rajasthan). Dorsal (46, 48, 50, 52, 54) and lateral (47, 49, 51, 53, 55, 56) views. Scale bars = 1.0 mm (A: 4653, B: 5456).
</paragraph>
</caption>
<paragraph id="8B962E56FF94D70CD192F9A0FB08263B" blockId="24.[151,1437,1631,2017]" pageId="24" pageNumber="525">Head with large convex eyes; tempora convex; genae very narrow, eyes ventrally almost touching buccal fissure. Labral apex almost straight. Supraorbital pore very small, located very close to supraorbital suture slightly before of level of posterior margin of eye. Frons and vertex coarsely and densely rugose-punctate, without microsculpture between punctures. Mandibles short, with almost straight or slightly arcuate mesal margin apically. Antennae moderately long, slightly surpassing pronotal basal margin or (in some females) not reaching it; antennomeres 411 markedly enlarged and rather thin, antennomeres 58 about 1.51.7 times as long as wide.</paragraph>
<paragraph id="8B962E56FF94D70DD192F8C8FF7A2057" blockId="24.[151,1437,1631,2017]" lastBlockId="25.[151,1437,151,1653]" lastPageId="25" lastPageNumber="526" pageId="24" pageNumber="525">Pronotum wide, widest slightly before middle. Sides rounded in apical two-thirds, generally slightly sinuate or occasionally almost rectilinear in basal third; lateral bead complete, very narrow throughout; medial lateral seta inserted almost in widest point. Apical margin comparatively shallowly emarginate, bordered along entire length. Apical angles wide, only slightly prominent anteriorly, rounded at apex. Basal margin markedly longer than apical margin, only slightly shorter than elytral base, distinctly bordered along entire length, ciliate. Basal angles obtusangular, with a tiny denticle at apex. Disc moderately convex, more or less depressed laterobasally. Basal foveae wide and shallow, outlines indistinct, fused with laterobasal depressions, or indistinct. Median line roughly engraved, short, markedly not reaching anterior and basal margins. Anterior and posterior transverse depressions indistinct. Surface densely and coarsely rugose-punctate, with fine obliterate microsculpture recognized only laterally.</paragraph>
<paragraph id="8B962E56FF95D70DD192FEB4FD662393" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">
Elytra oval, comparatively wide, widest behind middle, convex on disc and somewhat gently sloping to apex. Humeri slightly angularly rounded, with distinct small denticle. Lateral margins slightly serrate basally (
<figureCitation id="131232D3FF95D70DD46FFE90FA13209F" box="[1338,1424,367,393]" captionStart="FIGURES 4245" captionStartId="22.[151,264,1128,1152]" captionTargetBox="[161,1426,278,1095]" captionTargetId="figure-72@22.[154,1433,273,1105]" captionTargetPageId="22" captionText="FIGURES 4245. Platymetopus spp., habitus (42, 43, 45) and humerus (44). 42, P. tritus Bates, 1889 (Vietnam); 43, P. rugosus rugosus (Nietner, 1857) (India); 44, 45, P. r. obscurofemoratus subsp. n. (Vietnam, holotype). Scale bars = 1.0 mm (A: 44, B: 42, 43, 45)." figureDoi="http://doi.org/10.5281/zenodo.8073200" httpUri="https://zenodo.org/record/8073200/files/figure.png" pageId="25" pageNumber="526">Fig. 44</figureCitation>
). Preapical sinuation deep and rather short; sutural angle acutangular, narrowly rounded at tip. Basal border slightly sinuate, forming a very obtuse angle with lateral margin. Striae slightly impressed along entire length, indistinctly punctate anteriorly, gradually becoming smooth behind middle. Intervals almost flat or weakly convex (more clearly at apex), generally equally wide. Discal setigerous pores absent. Marginal umbilicate series more or less clearly divided in basal and apical groups, each consisting of 57 setigerous pores. Punctation fine and dense, mostly with four or five punctures across each interval in its middle portion. Microsculpture between punctures present throughout, consisting of slightly transverse meshes.
</paragraph>
<paragraph id="8B962E56FF95D70DD192FD70FC4323DB" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">Prosternal process bordered. Proepisternum distinctly, more or less coarsely punctate. Metepisternum much longer than wide, strongly narrowed posteriorly. Wings fully developed.</paragraph>
<paragraph id="8B962E56FF95D70DD192FD28FB482203" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">Metatarsus slender, in both sexes longer than HWmin and shorter than HWmax, moderately densely setose ventrally. Male pro- and mesotarsomeres 14 only slightly dilated, each with adhesive vestiture.</paragraph>
<paragraph id="8B962E56FF95D70DD192FCE0FE3D224B" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">Apex of abdominal sternite slightly concave in male and rounded in female, in both sexes with two pairs of marginal setigerous pores.</paragraph>
<paragraph id="8B962E56FF95D70DD192FC98FC1422DF" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">
Female genitalia (
<figureCitation id="131232D3FF95D70DD0C6FC98FD9C2297" box="[403,543,870,897]" captionStart="FIGURES 5764" captionStartId="26.[151,264,972,996]" captionTargetBox="[156,1430,456,942]" captionTargetId="figure-124@26.[151,1436,453,949]" captionTargetPageId="26" captionText="FIGURES 5764. Platymetopus spp., gonocoxite, lateral view (57, 59, 61, 63), and laterotergite, gonosubcoxite, and gonocoxite, ventral view (58, 60, 62, 64). 57, 58, P. tritus Bates, 1889 (Vietnam); 59, 60, P. rugosus (Nietner, 1857) (India); 61, 62, P. quadrimaculatus Dejean, 1829 (Vietnam); 63, 64, P. figuratus pictus Andrewes, 1923 (India, Rajasthan). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8073206" httpUri="https://zenodo.org/record/8073206/files/figure.png" pageId="25" pageNumber="526">Figs. 59, 60</figureCitation>
): laterotergite with membranous angulate apex and with one apical seta; distal mesal angle of gonosubcoxite not prominent, with a long seta at apex; gonocoxite very narrow, about as long as gonosubcoxite, with one short thin seta on dorsal ridge of outer side.
</paragraph>
<paragraph id="8B962E56FF95D70DD192FC2CFA10254F" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">
Median lobe of aedeagus (
<figureCitation id="131232D3FF95D70DD0A4FC2CFDFB22FB" box="[497,632,978,1005]" captionStart="FIGURES 4656" captionStartId="24.[151,264,1416,1440]" captionTargetBox="[153,1434,183,1391]" captionTargetId="figure-27@24.[151,1436,181,1393]" captionTargetPageId="24" captionText="FIGURES 4656. Platymetopus spp., median lobe of aedeagus. 46, 47, P. tritus Bates, 1889 (Vietnam); 48, 49, P. rugosus rugosus (Nietner, 1857) (India); 50, 51, P. r. obscurofemoratus subsp. n. (Vietnam, holotype); 52, 53, P. quadrimaculatus Dejean, 1829 (Vietnam); 5456, P. figuratus pictus Andrewes, 1923 (India, Rajasthan). Dorsal (46, 48, 50, 52, 54) and lateral (47, 49, 51, 53, 55, 56) views. Scale bars = 1.0 mm (A: 4653, B: 5456)." figureDoi="http://doi.org/10.5281/zenodo.8073202" httpUri="https://zenodo.org/record/8073202/files/figure.png" pageId="25" pageNumber="526">Figs. 4851</figureCitation>
) in lateral view comparatively strongly bent ventrally in apical quarter; its ventral margin almost straight or slightly arcuate medially; apex almost not bent, without a ventroapical denticle; in dorsal view median lobe almost straight or scarcely curved, more or less evenly narrowed apically; terminal lamella slightly longer than wide, triangular, blunted apically; internal sac with a narrow curved spine preapically.
</paragraph>
<paragraph id="8B962E56FF95D70DD192FB9DFB3924D7" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">
<emphasis id="B95DF244FF95D70DD192FB9DFEE1256A" bold="true" box="[199,354,1122,1148]" pageId="25" pageNumber="526">Comparison.</emphasis>
This small species with dark unicolorous elytra is readily distinguished from all other Asian congeners by a combination of several features unique to the region:
<emphasis id="B95DF244FF95D70DD2DBFB79FC2025B6" bold="true" box="[910,931,1158,1184]" pageId="25" pageNumber="526">1,</emphasis>
antennae dark throughout, almost black, with basal antennomeres clearly infuscate and with antennomeres 411 markedly enlarged and rather thin;
<emphasis id="B95DF244FF95D70DD5A5FB55FA8625D2" bold="true" box="[1264,1285,1194,1220]" pageId="25" pageNumber="526">2,</emphasis>
elytral lateral margin slightly serrate basally;
<emphasis id="B95DF244FF95D70DD0A2FB31FD8F25FE" bold="true" box="[503,524,1230,1256]" pageId="25" pageNumber="526">3,</emphasis>
genae very narrow, eyes ventrally almost touching buccal fissure;
<emphasis id="B95DF244FF95D70DD5A2FB31FA8F25FE" bold="true" box="[1271,1292,1230,1256]" pageId="25" pageNumber="526">4,</emphasis>
median lobe of aedeagus with a preapical spine in internal sac. In these characters,
<taxonomicName id="4C2955D5FF95D70DD2C0FB0CFB8B241A" box="[917,1032,1267,1292]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B95DF244FF95D70DD2C0FB0CFB8B241A" box="[917,1032,1267,1292]" italics="true" pageId="25" pageNumber="526">P. rugosus</emphasis>
</taxonomicName>
is taxonomically isolated from other Asian species and apparently more related to some Afrotropical species, for example,
<taxonomicName id="4C2955D5FF95D70DD53AFAE8FF512443" authority="Basilewsky, 1948" authorityName="Basilewsky" authorityYear="1948" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="platythorax">
<emphasis id="B95DF244FF95D70DD53AFAE8FA8E2427" box="[1135,1293,1303,1329]" italics="true" pageId="25" pageNumber="526">P. platythorax</emphasis>
<bibRefCitation id="EFB853A7FF95D70DD443FAE8FF512443" author="Basilewsky, P." pageId="25" pageNumber="526" pagination="209 - 227" refId="ref25724" refString="Basilewsky, P. (1948) Coleopteres harpalides nouveaux du Museum National d'Histoire Naturelle. Revue Francaise d'Entomologie, 14, 209 - 227." type="journal article" year="1948">Basilewsky, 1948</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2955D5FF95D70DD189FAC4FDA32443" authority="Facchini, 2004" authorityName="Facchini" authorityYear="2004" box="[220,544,1339,1365]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="cribricollis">
<emphasis id="B95DF244FF95D70DD189FAC4FEF12443" box="[220,370,1339,1365]" italics="true" pageId="25" pageNumber="526">P. cribricollis</emphasis>
<bibRefCitation id="EFB853A7FF95D70DD02DFAC4FDA32443" author="Facchini, S." box="[376,544,1339,1365]" pageId="25" pageNumber="526" pagination="113 - 120" refId="ref26223" refString="Facchini, S. (2004) Three new species of Platymetopus Dejean, 1829 from Africa (Coleoptera, Carabidae, Harpalinae). Giornale italiano di Entomologia, 11, 113 - 120." type="journal article" year="2004">Facchini, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2955D5FF95D70DD37CFAC4FCB32443" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" box="[553,816,1339,1365]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="lepidus">
<emphasis id="B95DF244FF95D70DD37CFAC4FD172443" box="[553,660,1339,1365]" italics="true" pageId="25" pageNumber="526">P. lepidus</emphasis>
<bibRefCitation id="EFB853A7FF95D70DD3CCFAC4FCB32443" author="Dejean, P. F. M. A." box="[665,816,1339,1365]" pageId="25" pageNumber="526" refId="ref26072" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2955D5FF95D70DD26FFAC4FBFC2443" authority="Basilewsky, 1947" authorityName="Basilewsky" authorityYear="1947" box="[826,1151,1339,1365]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="straeleni">
<emphasis id="B95DF244FF95D70DD26FFAC4FC352443" box="[826,950,1339,1365]" italics="true" pageId="25" pageNumber="526">P. straeleni</emphasis>
Basilewsky, 1947
</taxonomicName>
and
<taxonomicName id="4C2955D5FF95D70DD5E6FAC4FAB42443" authorityName="LaFerte-Senectere" authorityYear="1853" box="[1203,1335,1339,1365]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B95DF244FF95D70DD5E6FAC4FAB42443" box="[1203,1335,1339,1365]" italics="true" pageId="25" pageNumber="526">P. alternans</emphasis>
</taxonomicName>
LaFerte- Senectere, 1853, which also have serrate elytral margin and enlarged antennomeres. Very indistinct serration of lateral elytral margin occasionally also occurs in
<taxonomicName id="4C2955D5FF95D70DD39CFA7CFCC8248B" authorityName="Boheman" authorityYear="1848" box="[713,843,1411,1437]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="figuratus">
<emphasis id="B95DF244FF95D70DD39CFA7CFCC8248B" box="[713,843,1411,1437]" italics="true" pageId="25" pageNumber="526">P. figuratus</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C2955D5FF95D70DD22EFA7CFC72248A" box="[891,1009,1411,1436]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B95DF244FF95D70DD22EFA7CFC72248A" box="[891,1009,1411,1436]" italics="true" pageId="25" pageNumber="526">P. rugosus</emphasis>
</taxonomicName>
, the serration of the elytral margin is always present and distinct, though the degree of expression of this feature varies individually.
</paragraph>
</subSubSection>
<subSubSection id="C3337DDDFF95D70DD192FA35FAFB2747" pageId="25" pageNumber="526" type="distribution">
<paragraph id="8B962E56FF95D70DD192FA35FAFB2747" blockId="25.[151,1437,151,1653]" pageId="25" pageNumber="526">
<emphasis id="B95DF244FF95D70DD192FA35FEDA24F2" bold="true" box="[199,345,1482,1508]" pageId="25" pageNumber="526">Distribution</emphasis>
(
<figureCitation id="131232D3FF95D70DD03EFA34FE4124F3" box="[363,450,1483,1509]" captionStart="FIGURE 66" captionStartId="29.[151,250,1166,1190]" captionTargetBox="[151,1435,345,1142]" captionTargetId="figure-87@29.[151,1435,345,1142]" captionTargetPageId="29" captionText="FIGURE 66. Platymetopus rugosus (Nietner, 1857), distribution: P. r. rugosus (Nietner, 1857) (black circles, based on examined material; white circles, based on literature data); P. r. obscurofemoratus subsp. n. (stars)." figureDoi="http://doi.org/10.5281/zenodo.8073210" httpUri="https://zenodo.org/record/8073210/files/figure.png" pageId="25" pageNumber="526">Fig. 66</figureCitation>
). This species was described from many specimens collected in
<collectingCountry id="F33E6EC6FF95D70DD5F9FA34FAA324F3" box="[1196,1312,1483,1509]" name="Sri Lanka" pageId="25" pageNumber="526">Sri Lanka</collectingCountry>
. Andrewes (1921b, 1928, 1930) recorded it from
<collectingCountry id="F33E6EC6FF95D70DD337FA10FD30271F" box="[610,691,1519,1545]" name="Sri Lanka" pageId="25" pageNumber="526">Ceylon</collectingCountry>
(
<collectingCountry id="F33E6EC6FF95D70DD39DFA10FCBC271F" box="[712,831,1519,1545]" name="Sri Lanka" pageId="25" pageNumber="526">Sri Lanka</collectingCountry>
) and the territories of the modern Indian states of
<collectingRegion id="49EDE0B4FF95D70DD1C2F9ECFEA7273B" box="[151,292,1555,1581]" country="India" name="Uttarakhand" pageId="25" pageNumber="526">Uttarakhand</collectingRegion>
,
<collectingRegion id="49EDE0B4FF95D70DD065F9ECFE4E273B" box="[304,461,1555,1581]" country="India" name="Uttar Pradesh" pageId="25" pageNumber="526">Uttar Pradesh</collectingRegion>
,
<collectingRegion id="49EDE0B4FF95D70DD082F9ECFDDC273B" box="[471,607,1555,1581]" country="India" name="Tamil Nadu" pageId="25" pageNumber="526">Tamil Nadu</collectingRegion>
,
<collectingRegion id="49EDE0B4FF95D70DD33EF9ECFD62273B" box="[619,737,1555,1581]" country="India" name="Karnataka" pageId="25" pageNumber="526">Karnataka</collectingRegion>
,
<collectingRegion id="49EDE0B4FF95D70DD3B9F9ECFCE1273B" box="[748,866,1555,1581]" country="India" name="Jharkhand" pageId="25" pageNumber="526">Jharkhand</collectingRegion>
,
<collectingRegion id="49EDE0B4FF95D70DD23BF9ECFC43273B" box="[878,960,1555,1581]" country="India" name="Odisha" pageId="25" pageNumber="526">Odisha</collectingRegion>
,
<collectingRegion id="49EDE0B4FF95D70DD29FF9ECFB02273B" box="[970,1153,1555,1581]" country="India" name="Andhra Pradesh" pageId="25" pageNumber="526">Andhra Pradesh</collectingRegion>
, and
<collectingRegion id="49EDE0B4FF95D70DD5EEF9ECFACE273B" box="[1211,1357,1555,1581]" country="India" name="West Bengal" pageId="25" pageNumber="526">West Bengal</collectingRegion>
. Based on the examined material,
<taxonomicName id="4C2955D5FF95D70DD094F9C8FDB62746" box="[449,565,1591,1616]" class="Insecta" family="Carabidae" genus="Platymetopus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B95DF244FF95D70DD094F9C8FDB62746" box="[449,565,1591,1616]" italics="true" pageId="25" pageNumber="526">P. rugosus</emphasis>
</taxonomicName>
is widespread in the Oriental region from
<collectingCountry id="F33E6EC6FF95D70DD544F9C8FBC82747" box="[1041,1099,1591,1617]" name="India" pageId="25" pageNumber="526">India</collectingCountry>
and
<collectingCountry id="F33E6EC6FF95D70DD5D4F9C8FB6C2747" box="[1153,1263,1591,1617]" name="Sri Lanka" pageId="25" pageNumber="526">Sri Lanka</collectingCountry>
to
<collectingCountry id="F33E6EC6FF95D70DD447F9C8FAF62747" box="[1298,1397,1591,1617]" name="Vietnam" pageId="25" pageNumber="526">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3337DDDFF95D70DD192F9A4FC7C2763" box="[199,1023,1627,1653]" pageId="25" pageNumber="526" type="discussion">
<paragraph id="8B962E56FF95D70DD192F9A4FC7C2763" blockId="25.[151,1437,151,1653]" box="[199,1023,1627,1653]" pageId="25" pageNumber="526">
<emphasis id="B95DF244FF95D70DD192F9A4FEB82763" bold="true" box="[199,315,1627,1653]" pageId="25" pageNumber="526">Remarks.</emphasis>
The species forms two subspecies differing in color of femora.
</paragraph>
</subSubSection>
</treatment>
</document>