238 lines
29 KiB
XML
238 lines
29 KiB
XML
<document id="A2BA46B167D724B619BF6422460C78B2" ID-DOI="10.1080/00222933.2017.1324050" ID-ISSN="1464-5262" ID-Zenodo-Dep="5181888" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628698566541" checkinUser="felipe" docAuthor="Schmitter-Soto, Juan J." docDate="2017" docId="03806F39C96EFFDCFE43FB41D795FB7E" docLanguage="en" docName="JNatHist.51.23-24.1331-1424.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2017.1324050" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Astyanax macal Schmitter-Soto 2017, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1389" masterDocId="FFB91741C957FFE7FFBEFFB5D316FFA7" masterDocTitle="A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species" masterLastPageNumber="1424" masterPageNumber="1331" pageNumber="1387" updateTime="1699050015946" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="023B212FD1258D8C1C2F05E1082C9E35">A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species</mods:title>
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<mods:namePart id="1FEBD79BAEA0A90FA25177367071789C">Schmitter-Soto, Juan J.</mods:namePart>
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<mods:title id="08B7C03214D7DDD8FCF0DAE4DD311D4B">Journal of Natural History</mods:title>
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<mods:part id="02EDD2F5C22DC31629891B00CADF66B3">
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<mods:date id="9FDA10B51AF13CD7990B9BB23476B2E8">2017</mods:date>
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<mods:title id="C44961463CB373B3B5EA713BEC9F5C62">J. Nat. Hist.</mods:title>
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<mods:detail id="8627937000052ABE45B543AA4564AEEB" type="pubDate">
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<mods:number id="96B56C6056C3CE3EDC5BA45C970D20C6">2017-07-03</mods:number>
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<mods:number id="9CA42918A3F3C78E6FB1D539B0917008">51</mods:number>
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<mods:number id="872558E733872B59CB47DFBA0710BCF3">21 - 24</mods:number>
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<mods:url id="1E2BE325FA935C7F1A6D6D03D06FCCC9">http://dx.doi.org/10.1080/00222933.2017.1324050</mods:url>
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<mods:classification id="2FC7A741C86898F2CA64FDC995E15CCF">journal article</mods:classification>
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<mods:identifier id="75B7953A1D8DC2DC5AE1A60C41565BC9" type="DOI">10.1080/00222933.2017.1324050</mods:identifier>
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<mods:identifier id="B757B0E04C625500681C182B3A0A4CFB" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03806F39C96EFFDCFE43FB41D795FB7E" LSID="urn:lsid:plazi:treatment:03806F39C96EFFDCFE43FB41D795FB7E" httpUri="http://treatment.plazi.org/id/03806F39C96EFFDCFE43FB41D795FB7E" lastPageId="59" lastPageNumber="1389" pageId="57" pageNumber="1387">
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<subSubSection id="C3338DA4C96EFFDEFE43FB41D031FAA9" box="[509,807,1268,1294]" pageId="57" pageNumber="1387" type="nomenclature">
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<paragraph id="8B96DE2FC96EFFDEFE43FB41D031FAA9" blockId="57.[509,807,1268,1328]" box="[509,807,1268,1294]" pageId="57" pageNumber="1387">
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<heading id="D0DE6943C96EFFDEFE43FB41D031FAA9" box="[509,807,1268,1294]" centered="true" fontSize="11" level="2" pageId="57" pageNumber="1387" reason="5">
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<taxonomicName id="4C29A5ACC96EFFDEFE43FB41D1D7FAA9" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[509,705,1268,1294]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="species" species="macal" status="sp. nov.">
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<emphasis id="B95D023DC96EFFDEFE43FB41D1D7FAA9" bold="true" box="[509,705,1268,1294]" italics="true" pageId="57" pageNumber="1387">Astyanax macal</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C96EFFDEFD74FB40D031FAA9" box="[714,807,1269,1294]" pageId="57" pageNumber="1387" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3338DA4C96EFFDEFDAFFAA2D142FA3B" pageId="57" pageNumber="1387" type="description">
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||
<paragraph id="8B96DE2FC96EFFDEFDAFFAA2D004FA97" blockId="57.[509,807,1268,1328]" box="[529,786,1303,1328]" pageId="57" pageNumber="1387">
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(
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<figureCitation id="1312C2AAC96EFFDEFDA7FAA2D183FA97" box="[537,661,1303,1328]" captionStart="Figure 10" captionStartId="14.[160,225,1180,1203]" captionTargetBox="[168,1153,146,1149]" captionTargetId="figure-190@14.[163,1153,142,1149]" captionTargetPageId="14" captionText="Figure 10. (a) An opercle with parallel sides (Astyanax macal sp. nov., UMMZ 178599); (b) sides divergent at mid-length (A. tehuacanensis sp. nov., UMMZ 198853). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.5181909" httpUri="https://zenodo.org/record/5181909/files/figure.png" pageId="57" pageNumber="1387">Figures 10</figureCitation>
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(a), 15, 35)
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</paragraph>
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<paragraph id="8B96DE2FC96EFFDEFF1EFAD7D16FFADD" blockId="57.[160,633,1378,1436]" box="[160,633,1378,1402]" pageId="57" pageNumber="1387">
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<taxonomicName id="4C29A5ACC96EFFDEFF1EFAD7D252FADD" box="[160,324,1378,1402]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="species" species="undefined-4">
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<emphasis id="B95D023DC96EFFDEFF1EFAD7D215FADD" box="[160,259,1378,1402]" italics="true" pageId="57" pageNumber="1387">Astyanax</emphasis>
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sp. 4
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</taxonomicName>
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.
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<bibRefCitation id="EFB8A3DEC96EFFDEFEEEFAD7D165FADD" author="Ornelas-Garcia CP & Dominguez-Dominguez O & Doadrio I" box="[336,627,1378,1402]" pageId="57" pageNumber="1387" pagination="340" refId="ref48693" refString="Ornelas-Garcia CP, Dominguez-Dominguez O, Doadrio I. 2008. Evolutionary history of the fish genus Astyanax Baird and Girard 1854 (Actinopterygii, Characidae) in Mesoamerica reveals multiple morphological homoplasies. BMC Evol Biol. 8: 340." type="journal article" year="2008">Ornelas-García et al. 2008</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8B96DE2FC96EFFDEFF1EFA31D142FA3B" blockId="57.[160,633,1378,1436]" box="[160,596,1412,1436]" pageId="57" pageNumber="1387">
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<taxonomicName id="4C29A5ACC96EFFDEFF1EFA31D215FA3B" authorityName="Baird and Girard" authorityYear="1854" box="[160,259,1412,1436]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="genus">
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<emphasis id="B95D023DC96EFFDEFF1EFA31D215FA3B" box="[160,259,1412,1436]" italics="true" pageId="57" pageNumber="1387">Astyanax</emphasis>
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</taxonomicName>
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‘Macal’,
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<bibRefCitation id="EFB8A3DEC96EFFDEFED9FA31D158FA3B" author="Schmitter-Soto JJ" box="[359,590,1412,1436]" pageId="57" pageNumber="1387" pagination="101 - 130" refId="ref49172" refString="Schmitter-Soto JJ. 2016. A phylogeny of Astyanax (Characiformes: Characidae) in Central and North America. Zootaxa. 4109: 101 - 130." type="journal article" year="2016">Schmitter-Soto 2016</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C96EFFDEFF1EFA7CD10AF915" pageId="57" pageNumber="1387" type="diagnosis">
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<paragraph id="8B96DE2FC96EFFDEFF1EFA7CD202FA45" blockId="57.[160,1156,1481,1714]" box="[160,276,1481,1506]" pageId="57" pageNumber="1387">
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<emphasis id="B95D023DC96EFFDEFF1EFA7CD202FA45" bold="true" box="[160,276,1481,1506]" italics="true" pageId="57" pageNumber="1387">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8B96DE2FC96EFFDEFF1EFA59D10AF915" blockId="57.[160,1156,1481,1714]" pageId="57" pageNumber="1387">
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Diagnosed from other
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<taxonomicName id="4C29A5ACC96EFFDEFE12FA59D17AF9A3" box="[428,620,1516,1540]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B95D023DC96EFFDEFE12FA59D119F9A3" box="[428,527,1516,1540]" italics="true" pageId="57" pageNumber="1387">Astyanax</emphasis>
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species
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</taxonomicName>
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in Yucatan, Peten and
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<collectingCountry id="F33E9EBFC96EFFDEFC3BFA59D0D0F9A3" box="[901,966,1516,1540]" name="Belize" pageId="57" pageNumber="1387">Belize</collectingCountry>
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as follows: predorsal length, 50% SL or shorter (50% or longer in
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<taxonomicName id="4C29A5ACC96EFFDEFD44F9BAD044F980" authorityName="Hubbs" authorityYear="1936" box="[762,850,1551,1575]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="species" species="altior">
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<emphasis id="B95D023DC96EFFDEFD44F9BAD044F980" box="[762,850,1551,1575]" italics="true" pageId="57" pageNumber="1387">A. altior</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C29A5ACC96EFFDEFC2CF9BAD72EF980" baseAuthorityName="Regan" baseAuthorityYear="1908" box="[914,1080,1551,1575]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="species" species="angustifrons">
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<emphasis id="B95D023DC96EFFDEFC2CF9BAD72EF980" box="[914,1080,1551,1575]" italics="true" pageId="57" pageNumber="1387">A. angustifrons</emphasis>
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</taxonomicName>
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); least depth, 12% SL or less (12% or more in
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<taxonomicName id="4C29A5ACC96EFFDEFDECF987D1B1F9ED" authorityName="Hubbs" authorityYear="1936" box="[594,679,1586,1610]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="species" species="altior">
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<emphasis id="B95D023DC96EFFDEFDECF987D1B1F9ED" box="[594,679,1586,1610]" italics="true" pageId="57" pageNumber="1387">A. altior</emphasis>
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</taxonomicName>
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); anal fin origin usually without overlap to last dorsal-fin ray (usually overlapping in
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<taxonomicName id="4C29A5ACC96EFFDEFDCCF9E1D1C2F9CB" authorityName="Schmitter-Soto" authorityYear="2017" baseAuthorityName="Rosen" baseAuthorityYear="1972" box="[626,724,1620,1644]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="57" pageNumber="1387" phylum="Chordata" rank="species" species="baileyi">
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<emphasis id="B95D023DC96EFFDEFDCCF9E1D1C2F9CB" box="[626,724,1620,1644]" italics="true" pageId="57" pageNumber="1387">A. baileyi</emphasis>
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</taxonomicName>
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); scaly sheath in anal-fin base, double (vs single); nuptial tubercles in male can be bifid (vs always simple); rostral fontanel, sides convex (vs usually straight).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C96DFFDCFF1EFF3BD00DFD83" lastPageId="59" lastPageNumber="1389" pageId="58" pageNumber="1388" type="description">
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<paragraph id="8B96DE2FC96DFFDDFF1EFF3BD230FF00" blockId="58.[160,1157,142,1172]" box="[160,294,142,167]" pageId="58" pageNumber="1388">
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<emphasis id="B95D023DC96DFFDDFF1EFF3BD230FF00" bold="true" box="[160,294,142,167]" italics="true" pageId="58" pageNumber="1388">Description</emphasis>
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</paragraph>
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<paragraph id="8B96DE2FC96DFFDDFF1EFF07D793FF6D" blockId="58.[160,1157,142,1172]" box="[160,1157,178,202]" pageId="58" pageNumber="1388">
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A species of
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<taxonomicName id="4C29A5ACC96DFFDDFE92FF07D299FF6D" authorityName="Baird and Girard" authorityYear="1854" box="[300,399,178,202]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="58" pageNumber="1388" phylum="Chordata" rank="genus">
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<emphasis id="B95D023DC96DFFDDFE92FF07D299FF6D" box="[300,399,178,202]" italics="true" pageId="58" pageNumber="1388">Astyanax</emphasis>
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</taxonomicName>
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, subgenus
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<taxonomicName id="4C29A5ACC96DFFDDFDB1FF07D164FF6D" authorityName="Baird and Girard" authorityYear="1854" box="[527,626,178,202]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="58" pageNumber="1388" phylum="Chordata" rank="genus">
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<emphasis id="B95D023DC96DFFDDFDB1FF07D164FF6D" box="[527,626,178,202]" italics="true" pageId="58" pageNumber="1388">Astyanax</emphasis>
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</taxonomicName>
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(i.e. with a complete predorsal series of scales).
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</paragraph>
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<paragraph id="8B96DE2FC96DFFDDFF7EFF61D706FEF3" blockId="58.[160,1157,142,1172]" pageId="58" pageNumber="1388">Head profile straight; snout rounded. Lower and upper lips even, or upper lip a bit protruding; mouth somewhat upturned. Pectoral fins, not reaching posteriorly to pelvic fin origin in females (longer in males); anal and dorsal fins do not overlap vertically in females (they do, in males). Lobes of caudal fin, not equal: lower one longer.</paragraph>
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<paragraph id="8B96DE2FC96DFFDDFF7EFEEAD198FDCD" blockId="58.[160,1157,142,1172]" pageId="58" pageNumber="1388">
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D. 10; A. 21–24, mean 22.5; pect. 12. Procurrent unsegmented dorsal rays on caudal fin, 10 or more. Gill rakers on first arch, 17–20, modally 19; on lower limb, 9–11. Scales on lateral line, 34–37, modally 36; predorsal scales, 11–12; scale rows from lateral line to base of first dorsal-fin ray, 7–8.5, modally 8; to base of pelvic fin, 6–7; to base of pectoral fin, modally 4, up to 5; circumpeduncular scales, 14–16, modally 15. A double scale row on anal fin base, about 8 scales long. Nuptial tubercles only on anal fin rays, simple or bifid. Total vertebrae 34, 20 caudal. Detailed frequencies are given in
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<tableCitation id="C6ABEB94C96DFFDDFD8FFDE7D19EFDCD" box="[561,648,594,618]" captionStart="Table 3" captionStartId="16.[160,216,142,165]" captionText="Table 3. Meristics of Central and North American species in the genus Astyanax. Numbers are" pageId="58" pageNumber="1388">Table 3</tableCitation>
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.
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</paragraph>
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<paragraph id="8B96DE2FC96DFFDDFF7EFDC1D00EFD75" blockId="58.[160,1157,142,1172]" pageId="58" pageNumber="1388">
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Largest examined specimen,
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<quantity id="4CD173CAC96DFFDDFDA5FDC1D192FD2B" box="[539,644,628,652]" metricMagnitude="-2" metricUnit="m" metricValue="6.72" pageId="58" pageNumber="1388" unit="mm" value="67.2">67.2 mm</quantity>
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SL. Body depth, 32–39% SL. Head length 24–28% SL; orbital diameter, 31–34% HL; interorbital distance, 8.5–10.9% SL, mean 9.3% SL (further morphometric data appear in
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<tableCitation id="C6ABEB94C96DFFDDFD06FD0FD01AFD75" box="[696,780,698,722]" captionStart="Table 4" captionStartId="19.[160,216,137,160]" captionText="Table 4. Selected morphometric proportions of Central and North American species in the genus Astyanax (abbreviated by first letters of specific epithet). Specimens measured are those marked ‘dig’ in Appendix. All measurements given as percentage of standard length. Data are means; whole range in parentheses." pageId="58" pageNumber="1388">Table 4</tableCitation>
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).
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</paragraph>
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<paragraph id="8B96DE2FC96DFFDCFF7EFD69D0D7FE79" blockId="58.[160,1157,142,1172]" lastBlockId="59.[160,1156,142,548]" lastPageId="59" lastPageNumber="1389" pageId="58" pageNumber="1388">Anterior fontanel short, convex-sided, blunt-tipped. Supraoccipital process in dorsal view, short, wide-based; caudally undulate in lateral view. Vomer rostrally slightly concave. Dentigerous arm of premaxilla, longer; 0–4 teeth. Highest tooth on dentary, first or third; posterior teeth, abruptly smaller. Dorsal edge of longer articular arm, straight. Maxillary, with a convex anteroinferior edge; 1–3 teeth. Quadrate, dorsal process not expanded. Metapterygoid, rostral arm longer than ventral, 2 dorsorostral projections. Infraorbital II, a triangle with an angled base; infraorbital III, inferoposteriorly semicircular; infraorbital IV, rectangular with a projection; a wide contact between infraorbitals II and III. Urohyal rostral end turned up, pointed, its ventrorostral edge convex, its ventral apex closer to caudal end; no ceratohyal foramen; rostral vertices of urohyal angled, the ventral side straight. Epibranchial III, insertion of uncinate process angled, the distal segment of the main body straight. Upper pharyngeal bones, oval; lower pharyngeal plate not seen. Dorsal side of hyomandibular, convex. Opercular dorsal edge, markedly angled; sides of dorsal half, parallel; posterior edge, dorsally concave, ventrally straight-convex; ventral tip, sharp. Interopercle, posterior edge, straight-convex, with a spine. Preopercle, anterodorsal edge, with a median convexity; ventral rim, straight; posteroventral edge, curved; 2 divergent canals at angle. Five predorsal bony elements, distally expanded; rostral edge of first pterygiophore angled. Coracoid with 2–3 interdigitations in suture to cleithrum, a concave caudal edge, a double posteroinferior spine. Caudad process of postcleithrum, digitiform. Proximal edge of pelvic bone, convex. Dorsal tip of scaphium, truncate; caudal tip, sharp; dorsoposterior edge, slightly concave. Neural spines under dorsal fin, straight. Seventh caudal vertebra from tail, with a haemal spine displaced caudad. Rostral edge of largest hypuric plate, convex. Epuric plate on last neural spine, straight.</paragraph>
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<caption id="DF568EA7C96DFFDDFF1EF93AD01DF901" ID-DOI="http://doi.org/10.5281/zenodo.5182622" ID-Zenodo-Dep="5182622" box="[160,779,1679,1702]" httpUri="https://zenodo.org/record/5182622/files/figure.png" pageId="58" pageNumber="1388" startId="58.[160,225,1679,1702]" targetBox="[163,1153,1240,1648]" targetPageId="58">
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<paragraph id="8B96DE2FC96DFFDDFF1EF93AD01DF901" blockId="58.[160,779,1679,1702]" box="[160,779,1679,1702]" pageId="58" pageNumber="1388">
|
||
<emphasis id="B95D023DC96DFFDDFF1EF93AD21FF901" bold="true" box="[160,265,1679,1702]" pageId="58" pageNumber="1388">Figure 35.</emphasis>
|
||
<taxonomicName id="4C29A5ACC96DFFDDFEAFF93AD2BCF901" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[273,426,1679,1702]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="58" pageNumber="1388" phylum="Chordata" rank="species" species="macal" status="sp. nov.">
|
||
<emphasis id="B95D023DC96DFFDDFEAFF93AD2BCF901" box="[273,426,1679,1702]" italics="true" pageId="58" pageNumber="1388">Astyanax macal</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26EBF46C96DFFDDFE0DF93AD112F901" box="[435,516,1679,1702]" pageId="58" pageNumber="1388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, UMMZ 250291.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B96DE2FC96CFFDCFF7EFE5CD00DFD83" blockId="59.[160,1156,142,548]" pageId="59" pageNumber="1389">Humeral spot vertically oblong, but rather inconspicuous. Pigment on anal fin, denser distally. Caudal spot, both on peduncle and on fin rays.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C96CFFDCFF1EFDE5D0E1FD76" pageId="59" pageNumber="1389" type="materials_examined">
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFDE5D2E2FDCE" blockId="59.[160,1156,592,721]" box="[160,500,592,617]" pageId="59" pageNumber="1389">
|
||
<emphasis id="B95D023DC96CFFDCFF1EFDE5D2E2FDCE" bold="true" box="[160,500,592,617]" italics="true" pageId="59" pageNumber="1389">Type material and depositor</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFDC6D0E1FD76" blockId="59.[160,1156,592,721]" pageId="59" pageNumber="1389">
|
||
<materialsCitation id="3B41D472C96CFFDCFF1EFDC6D769FD09" collectionCode="UMMZ" collectorName="Lichtfield" country="Belize" location="Macal River" municipality="Maya Mountains" pageId="59" pageNumber="1389" specimenCode="UMMZ 250291" specimenCount="1" stateProvince="Cayo" typeStatus="holotype">
|
||
<typeStatus id="5492608DC96CFFDCFF1EFDC6D211FD2C" box="[160,263,627,651]" pageId="59" pageNumber="1389" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="DB8F7654C96CFFDCFEB1FDC6D2AEFD2C" box="[271,440,627,651]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="59" pageNumber="1389" type="University or college">UMMZ 250291</specimenCode>
|
||
,
|
||
<quantity id="4CD173CAC96CFFDCFE7DFDC6D130FD2C" box="[451,550,627,651]" metricMagnitude="-2" metricUnit="m" metricValue="6.72" pageId="59" pageNumber="1389" unit="mm" value="67.2">67.2 mm</quantity>
|
||
SL, ‘
|
||
<locationDeviation id="9A24C70DC96CFFDCFDE2FDC6D0CBFD2C" box="[604,989,627,651]" firstLocation="San Luis" pageId="59" pageNumber="1389" secondLocation="Millonarios">between San Luis and Millonarios</locationDeviation>
|
||
,
|
||
<collectingRegion id="49ED10CDC96CFFDCFC57FDC6D737FD2C" box="[1001,1057,627,651]" country="Belize" name="Cayo" pageId="59" pageNumber="1389">Cayo</collectingRegion>
|
||
District’,
|
||
<location id="8EF688F4C96CFFDCFF1EFD23D23FFD09" LSID="urn:lsid:plazi:treatment:03806F39C96EFFDCFE43FB41D795FB7E:8EF688F4C96CFFDCFF1EFD23D23FFD09" box="[160,297,662,686]" country="Belize" municipality="Maya Mountains" name="Macal River" pageId="59" pageNumber="1389" stateProvince="Cayo">Macal River</location>
|
||
,
|
||
<collectingMunicipality id="6BF24455C96CFFDCFE84FD23D2EBFD09" box="[314,509,662,686]" pageId="59" pageNumber="1389">Maya Mountains</collectingMunicipality>
|
||
, highlands of
|
||
<collectingCountry id="F33E9EBFC96CFFDCFD11FD23D1E5FD09" box="[687,755,662,686]" name="Belize" pageId="59" pageNumber="1389">Belize</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26DCBBF9C96CFFDCFC81FD23D0BEFD09" box="[831,936,662,686]" pageId="59" pageNumber="1389">Lichtfield</collectorName>
|
||
, 1960 (
|
||
<figureCitation id="1312C2AAC96CFFDCFBB8FD23D760FD09" box="[1030,1142,662,686]" captionStart="Figure 35" captionStartId="58.[160,225,1679,1702]" captionTargetBox="[163,1153,1240,1648]" captionTargetId="figure-421@58.[163,1153,1240,1648]" captionTargetPageId="58" captionText="Figure 35. Astyanax macal sp. nov., holotype, UMMZ 250291." figureDoi="http://doi.org/10.5281/zenodo.5182622" httpUri="https://zenodo.org/record/5182622/files/figure.png" pageId="59" pageNumber="1389">Figure 35</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="5492608DC96CFFDCFF1EFD0CD206FD76" box="[160,272,697,721]" pageId="59" pageNumber="1389" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectionCode id="ED3846EAC96CFFDCFEA5FD0CD270FD76" box="[283,358,697,721]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="59" pageNumber="1389" type="University or college">UMMZ</collectionCode>
|
||
178599,
|
||
<specimenCount id="9D2F15A6C96CFFDCFE6EFD0CD17BFD76" box="[464,621,697,721]" pageId="59" pageNumber="1389" type="generic">13 specimens</specimenCount>
|
||
, same collection data as
|
||
<typeStatus id="5492608DC96CFFDCFC32FD0CD0E5FD76" box="[908,1011,697,721]" pageId="59" pageNumber="1389" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C96CFFDCFF1EFD48D25BFCFC" pageId="59" pageNumber="1389" type="etymology">
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFD48D20BFCB1" blockId="59.[160,1156,765,859]" box="[160,285,765,790]" pageId="59" pageNumber="1389">
|
||
<emphasis id="B95D023DC96CFFDCFF1EFD48D20BFCB1" bold="true" box="[160,285,765,790]" italics="true" pageId="59" pageNumber="1389">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFC94D25BFCFC" blockId="59.[160,1156,765,859]" pageId="59" pageNumber="1389">
|
||
Named after the
|
||
<typeStatus id="5492608DC96CFFDCFED8FC94D28EFC9E" box="[358,408,801,825]" pageId="59" pageNumber="1389">type</typeStatus>
|
||
locality. A noun in apposition. Probably from the Mayan ‘makal’, meaning ‘yam’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C96CFFDCFF1EFC3DD11CFBD6" pageId="59" pageNumber="1389" type="distribution">
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFC3DD23AFC06" blockId="59.[160,1155,904,1033]" box="[160,300,904,929]" pageId="59" pageNumber="1389">
|
||
<emphasis id="B95D023DC96CFFDCFF1EFC3DD23AFC06" bold="true" box="[160,300,904,929]" italics="true" pageId="59" pageNumber="1389">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFC1ED0DDFBAE" blockId="59.[160,1155,904,1033]" pageId="59" pageNumber="1389">
|
||
Endemic to the upper Macal River, Maya Mountains,
|
||
<collectingCountry id="F33E9EBFC96CFFDCFD43FC1ED028FC64" box="[765,830,939,963]" name="Belize" pageId="59" pageNumber="1389">Belize</collectingCountry>
|
||
(
|
||
<figureCitation id="1312C2AAC96CFFDCFCF0FC1ED0AFFC64" box="[846,953,939,963]" captionStart="Figure 15" captionStartId="25.[160,225,848,871]" captionTargetBox="[163,1152,142,817]" captionTargetId="figure-294@25.[163,1153,142,817]" captionTargetPageId="25" captionText="Figure 15. Map of northern Central America and southern Mexico, showing distribution localities of Astyanax aeneus, A. altior, A. angustifrons, A. bacalarensis sp. nov., A. caballeroi, A. finitimus, A. macal sp. nov. and A. ocotal sp. nov. A larger symbol denotes the type locality (uncertain for A. angustifrons)." figureDoi="http://doi.org/10.5281/zenodo.5181919" httpUri="https://zenodo.org/record/5181919/files/figure.png" pageId="59" pageNumber="1389">Figure 15</figureCitation>
|
||
). There is at least one other species endemic to this river:
|
||
<taxonomicName id="4C29A5ACC96CFFDCFDCBFC7BD0F6FC41" authority="Greenfield 1990" authorityName="Greenfield" authorityYear="1990" box="[629,992,974,998]" class="Actinopterygii" family="Poeciliidae" genus="Poecilia" kingdom="Animalia" order="Cyprinodontiformes" pageId="59" pageNumber="1389" phylum="Chordata" rank="species" species="teresae">
|
||
<emphasis id="B95D023DC96CFFDCFDCBFC7BD009FC41" box="[629,799,974,998]" italics="true" pageId="59" pageNumber="1389">Poecilia teresae</emphasis>
|
||
Greenfield 1990
|
||
</taxonomicName>
|
||
. The river is a tributary of the eastern branch of the
|
||
<collectingCountry id="F33E9EBFC96CFFDCFDE8FC44D181FBAE" box="[598,663,1009,1033]" name="Belize" pageId="59" pageNumber="1389">Belize</collectingCountry>
|
||
River, Caribbean drainage.
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFB80D2C0FBE9" blockId="59.[160,522,1077,1137]" box="[160,470,1077,1102]" pageId="59" pageNumber="1389">
|
||
<emphasis id="B95D023DC96CFFDCFF1EFB80D2C0FBE9" bold="true" box="[160,470,1077,1102]" italics="true" pageId="59" pageNumber="1389">Proposed common names</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFBECD11CFBD6" blockId="59.[160,522,1077,1137]" box="[160,522,1113,1137]" pageId="59" pageNumber="1389">Macal tetra, sardinita del Macal.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C96CFFDCFF1EFB28D795FB7E" box="[160,1155,1181,1241]" pageId="59" pageNumber="1389" type="discussion">
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFB28D210FB11" blockId="59.[160,1155,1181,1241]" box="[160,262,1181,1206]" pageId="59" pageNumber="1389">
|
||
<emphasis id="B95D023DC96CFFDCFF1EFB28D210FB11" bold="true" box="[160,262,1181,1206]" italics="true" pageId="59" pageNumber="1389">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC96CFFDCFF1EFB74D795FB7E" blockId="59.[160,1155,1181,1241]" box="[160,1155,1217,1241]" pageId="59" pageNumber="1389">
|
||
<taxonomicName id="4C29A5ACC96CFFDCFF1EFB74D25FFB7E" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[160,329,1217,1241]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="59" pageNumber="1389" phylum="Chordata" rank="species" species="macal" status="sp. nov.">
|
||
<emphasis id="B95D023DC96CFFDCFF1EFB74D25FFB7E" box="[160,329,1217,1241]" italics="true" pageId="59" pageNumber="1389">Astyanax macal</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26EBF46C96CFFDCFEF1FB74D2B3FB7E" box="[335,421,1217,1241]" pageId="59" pageNumber="1389" rank="species">sp. nov.</taxonomicNameLabel>
|
||
corresponds to clade V or ‘species 4’ of
|
||
<bibRefCitation id="EFB8A3DEC96CFFDCFCE7FB74D769FB7E" author="Ornelas-Garcia CP & Dominguez-Dominguez O & Doadrio I" box="[857,1151,1217,1241]" pageId="59" pageNumber="1389" pagination="340" refId="ref48693" refString="Ornelas-Garcia CP, Dominguez-Dominguez O, Doadrio I. 2008. Evolutionary history of the fish genus Astyanax Baird and Girard 1854 (Actinopterygii, Characidae) in Mesoamerica reveals multiple morphological homoplasies. BMC Evol Biol. 8: 340." type="journal article" year="2008">Ornelas-García et al. (2008)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |