268 lines
35 KiB
XML
268 lines
35 KiB
XML
<document id="B164437601DE0CD46E60781588BEE22B" ID-DOI="10.11646/zootaxa.3964.3.1" ID-GBIF-Dataset="d80458ea-4fb4-4999-92c2-b8445c9dd19a" ID-ISSN="1175-5326" ID-Zenodo-Dep="231957" ID-ZooBank="88702846-1180-400A-B7DB-4154ACD4EC49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074488649" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2015" docId="EC3687B45811FF81FF3DFE6CFA77098C" docLanguage="en" docName="zt03964p320.pdf" docOrigin="Zootaxa 3964 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kophosphaera shivapuri Wesener, 2015, new species" docType="treatment" docVersion="9" lastPageNumber="316" masterDocId="100FFFCC581DFF8EFFAAF841FFEB0C68" masterDocTitle="The Giant Pill-Millipedes of Nepal (Diplopoda, Sphaerotheriida, Zephroniidae) *" masterLastPageNumber="320" masterPageNumber="301" pageNumber="313" updateTime="1698586053603" updateUser="plazi">
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<mods:title id="414944136915F31034B6D05BC3BF243E">The Giant Pill-Millipedes of Nepal (Diplopoda, Sphaerotheriida, Zephroniidae) *</mods:title>
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<mods:name id="65E1FF8AFC03296DFB47B0B5ED36F46E" type="personal">
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<mods:namePart id="EF3080EFEDE20955EE98D94F52FDB23B">Wesener, Thomas</mods:namePart>
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<mods:title id="0A0070E1EDC0365126E0A837D9E4A4D3">Zootaxa</mods:title>
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<mods:date id="B3C31676C842F85525C713D45C166B61">2015</mods:date>
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<mods:number id="A275F949F2C7F792D7902A4832472C6E">3964</mods:number>
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<treatment id="EC3687B45811FF81FF3DFE6CFA77098C" ID-DOI="http://doi.org/10.5281/zenodo.6100373" ID-GBIF-Taxon="119617620" ID-Zenodo-Dep="6100373" LSID="urn:lsid:plazi:treatment:EC3687B45811FF81FF3DFE6CFA77098C" httpUri="http://treatment.plazi.org/id/EC3687B45811FF81FF3DFE6CFA77098C" lastPageId="15" lastPageNumber="316" pageId="12" pageNumber="313">
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<subSubSection id="2C8565295811FF82FF3DFE6CFE7B0A00" pageId="12" pageNumber="313" type="nomenclature">
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<paragraph id="642036A25811FF82FF3DFE6CFDBC0A2F" blockId="12.[151,599,1581,1640]" box="[151,599,1581,1607]" pageId="12" pageNumber="313">
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<heading id="3F6881CE5811FF82FF3DFE6CFDBC0A2F" bold="true" box="[151,599,1581,1607]" fontSize="11" level="1" pageId="12" pageNumber="313" reason="1">
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<emphasis id="56EBEAB05811FF82FF3DFE6CFDBC0A2F" bold="true" box="[151,599,1581,1607]" pageId="12" pageNumber="313">
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<taxonomicName id="A39F4D215811FF82FF3DFE6CFE2B0A2F" box="[151,448,1581,1607]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="shivapuri" status="sp. nov.">
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<emphasis id="56EBEAB05811FF82FF3DFE6CFE2B0A2F" bold="true" box="[151,448,1581,1607]" italics="true" pageId="12" pageNumber="313">Kophosphaera shivapuri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4DD857CB5811FF82FE6DFE6CFDBC0A2F" box="[455,599,1581,1607]" pageId="12" pageNumber="313" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="642036A25811FF82FF3DFE0EFE7B0A00" blockId="12.[151,599,1581,1640]" box="[151,400,1615,1640]" pageId="12" pageNumber="313">
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<figureCitation id="FCA42A275811FF82FF3DFE0EFEEA0A00" box="[151,257,1615,1640]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="12" pageNumber="313">Figures 2</figureCitation>
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D, 3C, 8A–F
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</paragraph>
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</subSubSection>
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<subSubSection id="2C8565295811FF82FF3DFED6FA870AD8" box="[151,1388,1687,1712]" pageId="12" pageNumber="313" type="etymology">
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<paragraph id="642036A25811FF82FF3DFED6FA870AD8" blockId="12.[151,1436,1687,2037]" box="[151,1388,1687,1712]" pageId="12" pageNumber="313">
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<emphasis id="56EBEAB05811FF82FF3DFED6FEFD0AD8" bold="true" box="[151,278,1687,1712]" pageId="12" pageNumber="313">Etymology</emphasis>
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.
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<taxonomicName id="A39F4D215811FF82FE8EFED6FE650AD8" box="[292,398,1687,1712]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="shivapuri">
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<emphasis id="56EBEAB05811FF82FE8EFED6FE650AD8" box="[292,398,1687,1712]" italics="true" pageId="12" pageNumber="313">shivapuri</emphasis>
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</taxonomicName>
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, noun in apposition, after the
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<typeStatus id="BB2488005811FF82FD74FED9FCE50AD8" box="[734,782,1688,1712]" pageId="12" pageNumber="313">type</typeStatus>
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locality, which is located in the
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<taxonomicName id="A39F4D215811FF82FBD2FED6FB0E0AD8" box="[1144,1253,1687,1712]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="shivapuri" status="sp. nov.">Shivapuri</taxonomicName>
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Mountains.
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</paragraph>
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</subSubSection>
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<subSubSection id="2C8565295811FF82FF6DFEFDFDA70B75" pageId="12" pageNumber="313" type="materials_examined">
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<paragraph id="642036A25811FF82FF6DFEFDFDA70B75" blockId="12.[151,1436,1687,2037]" pageId="12" pageNumber="313">
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<emphasis id="56EBEAB05811FF82FF6DFEFDFE400ABD" bold="true" box="[199,427,1724,1749]" pageId="12" pageNumber="313">Material examined</emphasis>
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. 1 M
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<typeStatus id="BB2488005811FF82FE5EFEFDFDBD0ABD" box="[500,598,1724,1749]" pageId="12" pageNumber="313" type="holotype">holotype</typeStatus>
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(deposited at the
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<collectionCode id="028EAE675811FF82FC8EFEFDFCB40ABD" LSID="urn:lsid:biocol.org:col:34838" box="[804,863,1724,1749]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="313">
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<emphasis id="56EBEAB05811FF82FC8EFEFDFCB40ABD" bold="true" box="[804,863,1724,1749]" pageId="12" pageNumber="313">SMF</emphasis>
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</collectionCode>
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), 'N2 field label 305',
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<collectingCountry id="1C8876325811FF82FBC9FEFDFB420ABD" box="[1123,1193,1724,1749]" name="Nepal" pageId="12" pageNumber="313">Nepal</collectingCountry>
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, Kathmandu District, Sheopuri Mountain [=
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<taxonomicName id="A39F4D215811FF82FE39FE9EFDED0A90" box="[403,518,1759,1784]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="319" phylum="Arthropoda" rank="species" species="shivapuri" status="sp. nov.">Shivapuri</taxonomicName>
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,
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<geoCoordinate id="01AB50655811FF82FDBEFEA1FD550A90" box="[532,702,1760,1784]" direction="north" orientation="latitude" pageId="12" pageNumber="313" precision="1" value="27.795227">27°47'42.82"N</geoCoordinate>
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,
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<geoCoordinate id="01AB50655811FF82FD66FE9EFC9E0A90" box="[716,885,1759,1784]" direction="east" orientation="longitude" pageId="12" pageNumber="313" precision="1" value="85.34676">85°20'48.33"E</geoCoordinate>
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], degraded forest, bushes,
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<quantity id="A3679B475811FF82FB02FEA1FAA30A9F" box="[1192,1352,1760,1784]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="12" pageNumber="313" unit="m" value="1900.0" valueMax="2100.0" valueMin="1700.0">1700–2100 m</quantity>
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, leg. J. Martens & W. Schawaller,
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<date id="102110625811FF82FE6CFF45FDA30B75" box="[454,584,1796,1821]" pageId="12" pageNumber="313" value="1988-06-25">25.VI.1988</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="2C8565295811FF82FF6DFF66FDA10B9C" pageId="12" pageNumber="313" type="diagnosis">
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<paragraph id="642036A25811FF82FF6DFF66FDA10B9C" blockId="12.[151,1436,1687,2037]" pageId="12" pageNumber="313">
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<emphasis id="56EBEAB05811FF82FF6DFF66FED10B28" bold="true" box="[199,314,1831,1856]" pageId="12" pageNumber="313">Diagnosis</emphasis>
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. Syncoxite of anterior telopod without any udder-like projections, distinguishing
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<taxonomicName id="A39F4D215811FF82FB79FF68FAB50B28" box="[1235,1374,1831,1856]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="shivapuri">
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<emphasis id="56EBEAB05811FF82FB79FF68FAB50B28" box="[1235,1374,1831,1856]" italics="true" pageId="12" pageNumber="313">K. shivapuri</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="A39F4D215811FF82FF3DFF0CFED30B0C" box="[151,312,1868,1892]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="mammifera" status="stat. nov.">
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<emphasis id="56EBEAB05811FF82FF3DFF0CFED30B0C" box="[151,312,1868,1892]" italics="true" pageId="12" pageNumber="313">K. mammifera</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4DD857CB5811FF82FE95FF0CFE520B0C" box="[319,441,1869,1892]" pageId="12" pageNumber="313">new status</taxonomicNameLabel>
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. Male antennae with>45 apical cones, tarsi with 2 or 3 ventral spines, distinguishing
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<taxonomicName id="A39F4D215811FF82FA28FF0CFEEA0BE0" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="shivapuri">
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<emphasis id="56EBEAB05811FF82FA28FF0CFEEA0BE0" italics="true" pageId="12" pageNumber="313">K. shivapuri</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="A39F4D215811FF82FEEFFF30FE220BE0" box="[325,457,1905,1928]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="excavata">
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<emphasis id="56EBEAB05811FF82FEEFFF30FE220BE0" box="[325,457,1905,1928]" italics="true" pageId="12" pageNumber="313">K. excavata</emphasis>
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</taxonomicName>
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. Differing from
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<taxonomicName id="A39F4D215811FF82FD2EFF30FCEC0BE0" box="[644,775,1905,1928]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
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<emphasis id="56EBEAB05811FF82FD2EFF30FCEC0BE0" box="[644,775,1905,1928]" italics="true" pageId="12" pageNumber="313">K. martensi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4DD857CB5811FF82FCA4FF31FCA70BEF" box="[782,844,1904,1927]" pageId="12" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
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, among other characters, in the presence of a longer, divided locking carina and the shape of the process of the first pleurite, which is sharp-edged in
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<taxonomicName id="A39F4D215811FF82FB7EFFD4FAB30BC4" box="[1236,1368,1941,1964]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
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<emphasis id="56EBEAB05811FF82FB7EFFD4FAB30BC4" box="[1236,1368,1941,1964]" italics="true" pageId="12" pageNumber="313">K. martensi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4DD857CB5811FF82FAF5FFD4FA770BC4" box="[1375,1436,1941,1964]" pageId="12" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
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and well-rounded in
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<taxonomicName id="A39F4D215811FF82FE20FFF8FDF30BB9" box="[394,536,1976,2001]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="shivapuri">
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<emphasis id="56EBEAB05811FF82FE20FFF8FDF30BB9" box="[394,536,1976,2001]" italics="true" pageId="12" pageNumber="313">K. shivapuri</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="A39F4D215811FF82FD83FFF8FD5C0BB9" box="[553,695,1976,2001]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="shivapuri">
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<emphasis id="56EBEAB05811FF82FD83FFF8FD5C0BB9" box="[553,695,1976,2001]" italics="true" pageId="12" pageNumber="313">K. shivapuri</emphasis>
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</taxonomicName>
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with single, dense row of marginal bristles (
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<figureCitation id="FCA42A275811FF82FB6FFFF9FAE70BB8" box="[1221,1292,1976,2001]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="12" pageNumber="313">Fig. 2</figureCitation>
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D), while
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<taxonomicName id="A39F4D215811FF82FA28FFF8FF120B9C" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
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<emphasis id="56EBEAB05811FF82FA28FFF8FF120B9C" italics="true" pageId="12" pageNumber="313">K. martensi</emphasis>
|
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</taxonomicName>
|
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<taxonomicNameLabel id="4DD857CB5811FF82FEAAFF9CFED70B9C" box="[256,316,2013,2036]" pageId="12" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
|
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has two rows (
|
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<figureCitation id="FCA42A275811FF82FE4CFF9DFDC20B9C" box="[486,553,2012,2037]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="12" pageNumber="313">Fig. 2</figureCitation>
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E).
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</paragraph>
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</subSubSection>
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<caption id="30E0662A5810FF83FF3DFF34FE270B8F" httpUri="https://zenodo.org/record/231964/files/figure.png" pageId="13" pageNumber="314" targetBox="[170,1405,193,1875]" targetPageId="13">
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<paragraph id="642036A25810FF83FF3DFF34FE270B8F" blockId="13.[151,1436,1909,2023]" pageId="13" pageNumber="314">
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<emphasis id="56EBEAB05810FF83FF3DFF34FEF80BE3" bold="true" box="[151,275,1909,1931]" pageId="13" pageNumber="314">FIGURE 7.</emphasis>
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<taxonomicName id="A39F4D215810FF83FEB0FF34FD4C0BE3" authority="Butler, 1874" authorityName="Butler" authorityYear="1874" box="[282,679,1909,1931]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="excavata">
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<emphasis id="56EBEAB05810FF83FEB0FF34FDFB0BE3" box="[282,528,1909,1931]" italics="true" pageId="13" pageNumber="314">Kophosphaera excavata</emphasis>
|
||
(Butler, 1874)
|
||
</taxonomicName>
|
||
, male lectotype,
|
||
<emphasis id="56EBEAB05810FF83FCFFFF34FBCE0BE3" bold="true" box="[853,1061,1909,1931]" pageId="13" pageNumber="314">NHML BM1874.51</emphasis>
|
||
.
|
||
<emphasis id="56EBEAB05810FF83FB99FF34FBAE0BE3" bold="true" box="[1075,1093,1909,1931]" pageId="13" pageNumber="314">A</emphasis>
|
||
: left leg 9, posterior view;
|
||
<emphasis id="56EBEAB05810FF83FAF1FF34FA870BE3" bold="true" box="[1371,1388,1909,1931]" pageId="13" pageNumber="314">B</emphasis>
|
||
: left half of anterior telopod, anterior view;
|
||
<emphasis id="56EBEAB05810FF83FD8AFFD5FDD90BC2" bold="true" box="[544,562,1940,1962]" pageId="13" pageNumber="314">C</emphasis>
|
||
: left anterior telopod, podomeres 2–4, lateral view;
|
||
<emphasis id="56EBEAB05810FF83FB95FFD5FBBA0BC2" bold="true" box="[1087,1105,1940,1962]" pageId="13" pageNumber="314">D</emphasis>
|
||
: right posterior telopod, anterior view;
|
||
<emphasis id="56EBEAB05810FF83FF7FFFF3FF0D0BA0" bold="true" box="[213,230,1970,1992]" pageId="13" pageNumber="314">E</emphasis>
|
||
: right posterior telopod, posterior view.
|
||
<emphasis id="56EBEAB05810FF83FDD1FFF3FCFF0BA0" bold="true" box="[635,788,1970,1992]" pageId="13" pageNumber="314">Abbreviations</emphasis>
|
||
: IH = position of inner horns; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="30E0662A5813FF80FF3DFF1AFA770BA5" httpUri="https://zenodo.org/record/231965/files/figure.png" pageId="14" pageNumber="315" targetBox="[204,1372,193,1862]" targetPageId="14">
|
||
<paragraph id="642036A25813FF80FF3DFF1AFA770BA5" blockId="14.[151,1436,1883,1997]" pageId="14" pageNumber="315">
|
||
<emphasis id="56EBEAB05813FF80FF3DFF1AFEF80B19" bold="true" box="[151,275,1883,1905]" pageId="14" pageNumber="315">FIGURE 8.</emphasis>
|
||
<taxonomicName id="A39F4D215813FF80FEB0FF1AFDFE0B19" box="[282,533,1883,1905]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="shivapuri" status="sp. nov.">
|
||
<emphasis id="56EBEAB05813FF80FEB0FF1AFDFE0B19" box="[282,533,1883,1905]" italics="true" pageId="14" pageNumber="315">Kophosphaera shivapuri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="56EBEAB05813FF80FDB7FF1AFDBC0B19" bold="true" box="[541,599,1883,1905]" pageId="14" pageNumber="315">
|
||
<taxonomicNameLabel id="4DD857CB5813FF80FDB7FF1AFDBC0B19" box="[541,599,1883,1905]" pageId="14" pageNumber="315" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype,
|
||
<emphasis id="56EBEAB05813FF80FCA8FF1AFCDD0B19" bold="true" box="[770,822,1883,1905]" pageId="14" pageNumber="315">SMF</emphasis>
|
||
.
|
||
<emphasis id="56EBEAB05813FF80FCE9FF1AFCBE0B19" bold="true" box="[835,853,1883,1905]" pageId="14" pageNumber="315">A</emphasis>
|
||
: left leg 9, posterior view;
|
||
<emphasis id="56EBEAB05813FF80FBC7FF1AFB950B19" bold="true" box="[1133,1150,1883,1905]" pageId="14" pageNumber="315">B</emphasis>
|
||
: right coxa and prefemur 2, with gonopore;
|
||
<emphasis id="56EBEAB05813FF80FE91FF3BFEA60BF8" bold="true" box="[315,333,1914,1936]" pageId="14" pageNumber="315">C</emphasis>
|
||
: left half of anterior telopod, anterior view;
|
||
<emphasis id="56EBEAB05813FF80FCB6FF3BFCC50BF8" bold="true" box="[796,814,1914,1936]" pageId="14" pageNumber="315">D</emphasis>
|
||
: left anterior telopod, podomeres 2–4, lateral view;
|
||
<emphasis id="56EBEAB05813FF80FAE6FF3BFAB60BF8" bold="true" box="[1356,1373,1914,1936]" pageId="14" pageNumber="315">E</emphasis>
|
||
: right posterior telopod, anterior view;
|
||
<emphasis id="56EBEAB05813FF80FE40FFD9FE120BC6" bold="true" box="[490,505,1944,1966]" pageId="14" pageNumber="315">F</emphasis>
|
||
: right posterior telopod, posterior view.
|
||
<emphasis id="56EBEAB05813FF80FC36FFD9FBDE0BC6" bold="true" box="[924,1077,1944,1966]" pageId="14" pageNumber="315">Abbreviations</emphasis>
|
||
: Co = coxa; Go = gonopore; IH = position of inner horns; Pre = prefemur; St = stigmatic plate 2; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="2C8565295812FF81FF6DF8D6FC7609A8" pageId="15" pageNumber="316" type="description">
|
||
<paragraph id="642036A25812FF81FF6DF8D6FB910CD8" blockId="15.[151,1437,151,1509]" box="[199,1146,151,176]" pageId="15" pageNumber="316">
|
||
<emphasis id="56EBEAB05812FF81FF6DF8D6FEBA0CD8" bold="true" box="[199,337,151,176]" pageId="15" pageNumber="316">Description</emphasis>
|
||
. Measurements:
|
||
<quantity id="A3679B475812FF81FDBBF8D9FD9E0CC7" box="[529,629,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.27" pageId="15" pageNumber="316" unit="mm" value="22.7">22.7 mm</quantity>
|
||
long,
|
||
<quantity id="A3679B475812FF81FD16F8D9FCF90CC7" box="[700,786,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="15" pageNumber="316" unit="mm" value="9.8">9.8 mm</quantity>
|
||
(2nd) wide,
|
||
<quantity id="A3679B475812FF81FC0BF8D6FC1D0CC7" box="[929,1014,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="15" pageNumber="316" unit="mm" value="5.8">5.8 mm</quantity>
|
||
(2nd) high.
|
||
</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DF8FDFBF90CBD" blockId="15.[151,1437,151,1509]" box="[199,1042,188,213]" pageId="15" pageNumber="316">Colouration faded. Tergites brown, slightly dark greenish, legs dark brown.</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DF89EFCD70D0D" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">Head: Eyes with ca. 60 ocelli. Antennae short, protruding back to coxae 5. Antennomeres 1–5 with few longer setae, 6 more densely pubescent. Antennomere 6 only slightly wider than preceding antennomeres. Size relations of antennomeres 1=2=3=4=5<<6. Disc with 48/45 apical cones. Mouthparts not dissected. Palpi of gnathochilarium with sensory cones arranged in single field.</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DF92EFD370DEF" blockId="15.[151,1437,151,1509]" box="[199,732,367,392]" pageId="15" pageNumber="316">Collum glabrous except for few marginal setae.</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DF9D5FEFD0D9D" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">Thoracic shield surface glabrous except for longer setae in grooves. Grooves shallow, without any ledges. Tergites 3–12 smooth, at higher magnification chagrinated, leather-like, paratergite tips only weakly projecting posteriorly.</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DF9BEFE480E08" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">
|
||
Anal shield massive, strongly bell-shaped (
|
||
<figureCitation id="FCA42A275812FF81FD11F9BEFCEA0E70" box="[699,769,511,536]" captionStart="FIGURE 3" captionStartId="4.[151,250,614,636]" captionTargetBox="[150,1435,193,592]" captionTargetId="figure@4.[150,1436,193,593]" captionTargetPageId="4" captionText="FIGURE 3. Multi-layer photographs, anal shield and posterior tergites, lateral view. All specimens were males. A: Zephronia nepalensis n. sp., holotype, SMF; B: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176; C: K. shivapuri n. sp., holotype, SMF; D: K. martensi n. sp., holotype, SMF. Abbreviations: 12 T = tergite 12; AS = anal shield. Not to scale." httpUri="https://zenodo.org/record/231960/files/figure.png" pageId="15" pageNumber="316">Fig. 3</figureCitation>
|
||
C), covered with numerous small sensory hair. Underside carrying divided, long locking carina. Small gap dividing locking carina into two carinae of equal length. Carina located close to margin.
|
||
</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DFA2DFC460EA4" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">
|
||
Endotergum inner section covered with medium-length setae, each inserting in an elevated area (
|
||
<figureCitation id="FCA42A275812FF81FA98FA2DFA920EEC" box="[1330,1401,620,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="15" pageNumber="316">Fig. 2</figureCitation>
|
||
D). Between ridge and inner section, single row of circular cuticular impressions. Externally, single row of marginal bristles (
|
||
<figureCitation id="FCA42A275812FF81FF5DFAF5FED10EA4" box="[247,314,692,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="15" pageNumber="316">Fig. 2</figureCitation>
|
||
D). Bristles short, not reaching tergite margin (
|
||
<figureCitation id="FCA42A275812FF81FCECFAF5FC620EA4" box="[838,905,692,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="15" pageNumber="316">Fig. 2</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DFA96FC7E0E87" blockId="15.[151,1437,151,1509]" box="[199,917,727,752]" pageId="15" pageNumber="316">First stigma-carrying plate with a well-rounded projecting apex.</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DFABDFDF00F5F" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">Pleurite 1 laterally elongated into a slender, well-rounded process. Pleurite 2 also weakly elongated into a broader and well-rounded process.</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DFB04FAAF0FCD" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">
|
||
Leg 1 with 2, 2 with 3, 3 with 5 ventral spines. Leg pairs 4–21 with 7 or 8 ventral spines. Legs 3–21 with single apical spine, which is absent in leg pairs 1 and 2. Coxa process strongly developed and well-rounded (
|
||
<figureCitation id="FCA42A275812FF81FA99FB26FA9C0FE8" box="[1331,1399,871,896]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[204,1372,193,1862]" captionTargetId="figure@14.[204,1376,193,1862]" captionTargetPageId="14" captionText="FIGURE 8. Kophosphaera shivapuri n. sp., male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with gonopore; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Co = coxa; Go = gonopore; IH = position of inner horns; Pre = prefemur; St = stigmatic plate 2; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231965/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
|
||
A), absent from leg pair 1 and 2. Femur 1.5, tarsus 3.2 times longer than wide. Femur with short toothed ridge.
|
||
</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DFBEEFD640F84" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">
|
||
Male gonopore large, simple pit covered by partly membranous plate, located centrally at the mesal margin of coxa 2, size ca. 1/3 of coxal height (
|
||
<figureCitation id="FCA42A275812FF81FD80FB95FD860F84" box="[554,621,980,1005]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[204,1372,193,1862]" captionTargetId="figure@14.[204,1376,193,1862]" captionTargetPageId="14" captionText="FIGURE 8. Kophosphaera shivapuri n. sp., male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with gonopore; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Co = coxa; Go = gonopore; IH = position of inner horns; Pre = prefemur; St = stigmatic plate 2; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231965/files/figure.png" pageId="15" pageNumber="316">Fig. 8</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DFBB6FC0C08AD" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">
|
||
Anterior telopod (
|
||
<figureCitation id="FCA42A275812FF81FE3FFBB6FE350878" box="[405,478,1015,1040]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[204,1372,193,1862]" captionTargetId="figure@14.[204,1376,193,1862]" captionTargetPageId="14" captionText="FIGURE 8. Kophosphaera shivapuri n. sp., male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with gonopore; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Co = coxa; Go = gonopore; IH = position of inner horns; Pre = prefemur; St = stigmatic plate 2; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231965/files/figure.png" pageId="15" pageNumber="316">Figs 8</figureCitation>
|
||
C, D): syncoxite wider than podomere 1, with few longer setae. Podomere 1 larger than podomere 2 or 3, rectangular, wider than following podomeres. Podomere 2 large, posterior process not visible in anterior view, process relatively slender only weakly curved, protruding as far as podomere 4. Podomere 3 shorter and slenderer than podomere 2. Podomere 4 long, reaching almost half the size of podomere 3, towards process of podomere 2 with at least one sclerotized spine. Podomeres 1 and
|
||
<quantity id="A3679B475812FF81FBA2FCC9FBDD08C8" box="[1032,1078,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="316" unit="in" value="2.0">2 in</quantity>
|
||
anterior view sparsely covered with long setae, podomeres 3 and 4 glabrous except for some marginal hair.
|
||
</paragraph>
|
||
<paragraph id="642036A25812FF81FF6DFC8EFC7609A8" blockId="15.[151,1437,151,1509]" pageId="15" pageNumber="316">
|
||
Posterior telopod (
|
||
<figureCitation id="FCA42A275812FF81FE3FFC8EFE310880" box="[405,474,1231,1256]" captionStart="FIGURE 8" captionStartId="14.[151,250,1883,1905]" captionTargetBox="[204,1372,193,1862]" captionTargetId="figure@14.[204,1376,193,1862]" captionTargetPageId="14" captionText="FIGURE 8. Kophosphaera shivapuri n. sp., male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with gonopore; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Co = coxa; Go = gonopore; IH = position of inner horns; Pre = prefemur; St = stigmatic plate 2; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231965/files/figure.png" pageId="15" pageNumber="316">Figs 8</figureCitation>
|
||
E, F): typical for
|
||
<taxonomicName id="A39F4D215812FF81FD30FC8EFCD40880" box="[666,831,1231,1256]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56EBEAB05812FF81FD30FC8EFCD40880" box="[666,831,1231,1256]" italics="true" pageId="15" pageNumber="316">Kophosphaera</emphasis>
|
||
</taxonomicName>
|
||
, with 4 podomeres, 2 membranous ledges and lacking any lobes, or sclerotized teeth. Podomere 3 straight, 2.3 times longer than wide. Hollowed-out inner margin with membranous ledge. Podomere 4 well-developed, 0.5 times as long as podomere 3, with three large sclerotized spines next to membranous ledge. Podomeres 3 and 4 curved towards immovable finger. Immovable finger wide, apically only weakly tapering, shorter than podomeres 3 and 4. Podomeres 1 and
|
||
<quantity id="A3679B475812FF81FB9DFD21FB880910" box="[1079,1123,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="316" unit="in" value="2.0">2 in</quantity>
|
||
anterior view covered with setae, in posterior view podomere 1 setose, but podomere 2 only with marginal hair. Podomere
|
||
<quantity id="A3679B475812FF81FB11FDC5FB0D09F5" box="[1211,1254,1412,1437]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="15" pageNumber="316" unit="in" value="3.0">3 in</quantity>
|
||
posterior aspect glabrous, anteriorly with setae in its basal half, podomere 4 glabrous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2C8565295812FF81FF6DFD8DFA77098C" box="[199,1436,1484,1509]" pageId="15" pageNumber="316" type="distribution">
|
||
<paragraph id="642036A25812FF81FF6DFD8DFA77098C" blockId="15.[151,1437,151,1509]" box="[199,1436,1484,1509]" pageId="15" pageNumber="316">
|
||
<emphasis id="56EBEAB05812FF81FF6DFD8DFEB2098D" bold="true" box="[199,345,1484,1509]" pageId="15" pageNumber="316">Distribution</emphasis>
|
||
. Only known from the mountain of
|
||
<taxonomicName id="A39F4D215812FF81FD46FD8DFCB7098D" box="[748,860,1484,1509]" class="Diplopoda" family="Zephronidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="319" phylum="Arthropoda" rank="species" species="shivapuri" status="sp. nov.">Shivapuri</taxonomicName>
|
||
, north of Kathmandu in central
|
||
<collectingCountry id="1C8876325812FF81FB14FD8DFAEA098D" box="[1214,1281,1484,1509]" name="Nepal" pageId="15" pageNumber="316">Nepal</collectingCountry>
|
||
(Fig. Map 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |